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authorityParentheses originalNameCombination authoritySpeciesCitation authoritySpeciesLink typeVoucher typeKind typeVoucherURIs typeLocality typeLocalityLatitude typeLocalityLongitude taxonomyNotes taxonomyNotesCitation distributionNotes distributionNotesCitation subregionDistribution countryDistribution continentDistribution biogeographicRealm iucnStatus extinct domestic flagged CMW_sciName diffSinceCMW MSW3_matchtype MSW3_sciName diffSinceMSW3 docOrigin_batnames2025 Family Genus Subgenus Species Author Date Parentheses (1=author & date in parentheses) Citation Pages Common Name Synonyms Type Locality Distribution CITES IUCN Comments column3781 column3791 subtribe CONCAT_ALTNAMES line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L407 application/vnd.openxmlformats-officedocument.spreadsheetml.sheet N/A N/A N/A N/A N/A N/A Hipposideros khaokhouayensis Hipposideros khaokhouayensis Hipposideros khaokhouayensis Hipposideros khaokhouayensis Hipposideros khaokhouayensis Hipposideros khaokhouayensis Hipposideros khaokhouayensis Hipposideros khaokhouayensis Hipposideros khaokhouayensis [HMW] Hipposideros khaokhouayensis Guillén-Servent & Francis, 2006 , “ Tak Leukwaterfall, along Nam (river) Leuk, in Phou Khao Khouay National Biodiversity Conservation Area, Saisombun Special Zone, Central Lao PDR, 18°23’N , 103°04’E , 180 m elevation.” Hipposideros khaokhouayensis was formerly included in the bicolor species group, but is now placed in the new ater species group. Monotypic.; [MDD2022] recently described; [IUCN] This new species is a member of the Hipposideros bicolor group.; [MDD2023] recently described; [MDD2025_2.0] recently described; [MDD2025_2.2] recently described khaokhouayensis This new species is a member of the Hipposideros bicolor group. khaokhouayensis khaokhouayensis khaokhouayensis GuillÃ©n-Servent & C. M. Francis, 2006 N/A NA 03BD87A2C66AA218FF5DEF31F3984475 Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions 978-84-16728-19-0 hbmw_9_Hipposideridae_210.pdf.imf hash://md5/ff84ffdac676a204fff8ff9affef4346 255 zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C66AA218FF5DEF31F3984475.xml Hipposideros khaokhouayensis Hipposideridae Hipposideros khaokhouayensis Guillen-Servent & Francis 2006 Phyllorhine du Khao Khouay @fr | Phou-Khao-Khouay-Rundblattnase @de | Hiposidérido de Khao Khouay @es | Phou Khao Khouay Roundleaf Bat @en Hipposideros khaokhouayensis Guillén-Servent & Francis, 2006 , “ Tak Leukwaterfall, along Nam (river) Leuk, in Phou Khao Khouay National Biodiversity Conservation Area, Saisombun Special Zone, Central Lao PDR, 18°23’N , 103°04’E , 180 m elevation.” Hipposideros khaokhouayensis was formerly included in the bicolor species group, but is now placed in the new ater species group. Monotypic. Endemic to Laos , known only from type locality and Ban Nampe, C.I 1 km NE of Vang Vieng, Vang Vieng district, Vientiane Province . Ear 24—26 mm , forearm 46-49 mm. Noseleaf of the Phou Khao Khouay Leaf-nosed Bat is narrow and without lateral supplementary leaflet Intemarial septum forms an intermediate-sized disk shape. Anterior leaf has narrow emargination. Dorsal pelage is generally mid-brown, with individual hairs white at base, brown in the middle, and pale at tip; ventral side is paler. Rostrum is inflated with large posterolateral chamber. Sagittal crest is moderately developed. The Phou Khao Khouay Leaf-nosed Bat was captured in patches of intact and disturbed evergreen forests both near and far from caves it is known to occupy. It can be found at elevations of 180—400 m . No information. Lactating female was captured in May-June. The Phou Khao Khouay Leaf-nosed Bat roosts in limestone caves. Echolocation call frequency is 87-91 kHz. No information. Classified as Vulnerable on IUCN Red List. The major threat to the Phou Khao Khouay Leaf-nosed Bat is habitat loss; this is aggravated by lack of law enforcement. Bates & Francis (2008a) | Francis (2008a) | Guillén-Servent & Francis (2006) 76. Phou Khao Khouay Leaf-nosed Bat Hipposideros khaokhouayensis French: Phyllorhine du Khao Khouay I German: Phou-Khao-Khouay-Rundblattnase / Spanish: Hiposidérido de Khao Khouay Other common names: Phou Khao Khouay Roundleaf Bat Taxonomy. Hipposideros khaokhouayensis Guillén-Servent & Francis, 2006 , “ Tak Leukwaterfall, along Nam (river) Leuk, in Phou Khao Khouay National Biodiversity Conservation Area, Saisombun Special Zone, Central Lao PDR, 18°23’N , 103°04’E , 180 m elevation.” Hipposideros khaokhouayensis was formerly included in the bicolor species group, but is now placed in the new ater species group. Monotypic. Distribution. Endemic to Laos , known only from type locality and Ban Nampe, C.I 1 km NE of Vang Vieng, Vang Vieng district, Vientiane Province . Descriptive notes. Ear 24—26 mm , forearm 46-49 mm. Noseleaf of the Phou Khao Khouay Leaf-nosed Bat is narrow and without lateral supplementary leaflet Intemarial septum forms an intermediate-sized disk shape. Anterior leaf has narrow emargination. Dorsal pelage is generally mid-brown, with individual hairs white at base, brown in the middle, and pale at tip; ventral side is paler. Rostrum is inflated with large posterolateral chamber. Sagittal crest is moderately developed. Habitat. The Phou Khao Khouay Leaf-nosed Bat was captured in patches of intact and disturbed evergreen forests both near and far from caves it is known to occupy. It can be found at elevations of 180—400 m . Food and Feeding. No information. Breeding. Lactating female was captured in May-June. Activity patterns. The Phou Khao Khouay Leaf-nosed Bat roosts in limestone caves. Echolocation call frequency is 87-91 kHz. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Vulnerable on IUCN Red List. The major threat to the Phou Khao Khouay Leaf-nosed Bat is habitat loss; this is aggravated by lack of law enforcement. Bibliography. Bates & Francis (2008a), Francis (2008a), Guillén-Servent & Francis (2006). Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022. Hipposideridae Hipposideros khaokhouayensis Hipposideros khaokhouayensis Guillén-Servent & Francis 2006 0 Acta Chiropterol. 8(1): 44 Phou Khao Khouay Leaf-nosed Bat None. Laos, Saisombun Special Zone, Phou Khou Khouay National Biodiversity Conservation Area, Tak Luk waterfall C Laos Not listed. Vulnerable Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023 Hipposideros khaokhouayensis 23 Phou Khao Khouay Leaf-nosed Bat Phou Khao Khouay Roundleaf Bat Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA PTEROPODIFORMES NA NA RHINOLOPHOIDEA HIPPOSIDERIDAE NA NA Hipposideros NA khaokhouayensis GuillÃ©n-Servent & Francis 2006 0 Hipposideros_khaokhouayensis GuillÃ©n-Servent, A., & Francis, C. M. (2006). A new species of bat of the Hipposideros bicolor group (Chiroptera: Hipposideridae) from Central Laos, with evidence of convergent evolution with Sundaic taxa. Acta Chiropterologica, 8(1), 44. https://bioone.org/journals/acta-chiropterologica/volume-8/issue-1/1733-5329(2006)8%5B39%3AANSOBO%5D2.0.CO%3B2/A-new-species-of-bat-of-the-Hipposideros-bicolor-group/10.3161/1733-5329(2006)8[39:ANSOBO]2.0.CO;2.full ROM 116790 "Tak Leuk waterfall, along Nam (river) Leuk, in Phou Khao Khouay National Biodiversity Conservation Area, Saisombun Special Zone, Central Lao PDR, 18Â°23'N, 103Â°04'E, 180 m elevation." 18.383 103.07 khaokhouayensis GuillÃ©n-Servent & Francis, 2006 recently described GuillÃ©n-Servent, A., & Francis, C. M. (2006). A new species of bat of the Hipposideros bicolor group (Chiroptera: Hipposideridae) from Central Laos, with evidence of convergent evolution with Sundaic taxa. Acta Chiropterologica, 8(1), 39-61. Laos Asia Indomalaya VU 0 0 0 Hipposideros_khaokhouayensis 0 unmatched NA 1 IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022]. 136819 Hipposideros khaokhouayensis ANIMALIA CHORDATA MAMMALIA CHIROPTERA HIPPOSIDERIDAE Hipposideros khaokhouayensis GuillÃ©n-Servent &; Francis, 2006 This new species is a member of the Hipposideros bicolor group. 200000000 Hipposideros khaokhouayensis Vulnerable C1 2020 2018-08-28 00:00:00 UTC 3.1 English This species is listed as Vulnerable as it is only known from three areas in Lao and Viet Nam and the global population is estimated to be between 8,000 to 10,000 individuals. Over the next 15 years (three generations; Pacifici et al. 2013), the species is projected to continue to decline by at least 10% due to ongoing degradation and loss of the species habitat. Surveys to the northwest, northeast and south of the two known localities have failed to find additional colonies, and it is not likely to have been overlooked in collections as it is a very distinctive species. In Lao PDR, these bats have been captured in patches of intact and disturbed evergreen forest. In VangVieng there were large limestone outcrops with caves in the vicinity. Capture sites in Phou Khao Khouay were not near limestone outcrops or large caves, but the roost sites are not known (GuillÃ©n-Servent and Francis 2006). In Viet Nam, it was captured around a building which is surrounded by primary forest covering on limestone areas (Thong et al. 2008). Habitat loss is a threat to this species as Phou Khao Khouay National Biodiversity Conservation Area is not well managed. One of the locations is affected by deforestation, road work and building associated with the development of the large Tad Leuk dam project (GuillÃ©n-Servent and Francis 2006). The type locality has been flooded for the dam (C. Francis pers. comm). It is known from only two sites in Lao PDR (GuillÃ©n-Servent and Francis 2006) and one site in Viet Nam (Thong et al. 2008). Surveys in surrounding areas have failed to locate the species (C. Francis pers. comm). The species is projected to be declining due to continuing habitat loss. Decreasing The species has been collected in a small area at the center of Phou Khao Khouay National Biodiversity Conservation Area, Saisombun Special Area, where it appeared to be relatively common, and near Ban Nampe, about 11 km northeast of VangVieng, VangVieng District, Vientiane Province, all in central Lao PDR (GuillÃ©n-Servent and Francis 2006). The species has also been recorded from Cat Ba Biosphere Reserve in north-eastern Vietnam (Thong et al. 2008). The species has been caught between 180 m and 400 m asl (GuillÃ©n-Servent and Francis 2006). Terrestrial This species occurs in the Phou Khao Khouay National Biodiversity Conservation Area. Further research is required to inform conservation of forest habitats. Indomalayan FALSE FALSE Global Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 Hipposideridae Hipposideros khaokhouayensis Guillén-Servent & Francis 2006 0 Acta Chiropterol. 8(1): 44 Phou Khao Khouay Leaf-nosed Bat None. Laos, Saisombun Special Zone, Phou Khou Khouay National Biodiversity Conservation Area, Tak Luk waterfall C Laos Not listed. Vulnerable Hipposideros khaokhouayensis 1004609 23 Phou Khao Khouay Leaf-nosed Bat Phou Khao Khouay Roundleaf Bat Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA PTEROPODIFORMES NA NA RHINOLOPHOIDEA Hipposideridae NA NA Hipposideros NA khaokhouayensis GuillÃ©n-Servent & Francis 2006 0 Hipposideros_khaokhouayensis GuillÃ©n-Servent, A., & Francis, C. M. (2006). A new species of bat of the Hipposideros bicolor group (Chiroptera: Hipposideridae) from Central Laos, with evidence of convergent evolution with Sundaic taxa. Acta Chiropterologica, 8(1), 44. https://bioone.org/journals/acta-chiropterologica/volume-8/issue-1/1733-5329(2006)8%5B39%3AANSOBO%5D2.0.CO%3B2/A-new-species-of-bat-of-the-Hipposideros-bicolor-group/10.3161/1733-5329(2006)8[39:ANSOBO]2.0.CO;2.full ROM 116790 "Tak Leuk waterfall, along Nam (river) Leuk, in Phou Khao Khouay National Biodiversity Conservation Area, Saisombun Special Zone, Central Lao PDR, 18Â°23'N, 103Â°04'E, 180 m elevation." 18.38333 103.0667 khaokhouayensis GuillÃ©n-Servent & Francis, 2006 recently described GuillÃ©n-Servent, A., & Francis, C. M. (2006). A new species of bat of the Hipposideros bicolor group (Chiroptera: Hipposideridae) from Central Laos, with evidence of convergent evolution with Sundaic taxa. Acta Chiropterologica, 8(1), 39-61. Laos Asia Indomalaya VU 0 0 0 Hipposideros_khaokhouayensis 0 unmatched NA 1 Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393 Hipposideros_khaokhouayensis 1004609 23 Phou Khao Khouay Roundleaf Bat Phou Khao Khouay Roundleaf Bat|Phou Khao Khouay Leaf-nosed Bat Theria Placentalia Boreoeutheria Laurasiatheria Chiroptera Yinpterochiroptera NA NA Rhinolophoidea Hipposideridae NA NA Hipposideros NA khaokhouayensis GuillÃ©n-Servent & C. M. Francis 0 Hipposideros khaokhouayensis GuillÃ©n-Servent, A. and Francis, C.M. 2006-04. A new species of bat of the _Hipposideros bicolor_ group (Chiroptera: Hipposideridae) from Central Laos, with evidence of convergent evolution with Sundaic taxa. Acta Chiropterologica 8(1):39-61. https://doi.org/10.3161/1733-5329(2006)8[39:ANSOBO]2.0.CO;2 ROM:MAM:116790 holotype http://portal.vertnet.org/o/rom/mammals?id=uri-catalog-rom-mammals-116790 "Tak Leuk waterfall, along Nam (river) Leuk, in Phou Khao Khouay National Biodiversity Conservation Area, Saisombun Special Zone, Central Lao PDR, 18Â°23'N, 103Â°04'E, 180 m elevation." 18.38333 103.0667 recently described GuillÃ©n-Servent, A., & Francis, C. M. (2006). A new species of bat of the Hipposideros bicolor group (Chiroptera: Hipposideridae) from Central Laos, with evidence of convergent evolution with Sundaic taxa. Acta Chiropterologica, 8(1), 39-61. Laos Asia Indomalaya VU 0 0 0 Hipposideros_khaokhouayensis 0 unmatched NA 1 Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586 Hipposideridae Hipposideros khaokhouayensis Guillén-Servent & Francis 2006 0 Acta Chiropterol. 8(1): 44 Phou Khao Khouay Leaf-nosed Bat None. Laos, Saisombun Special Zone, Phou Khou Khouay National Biodiversity Conservation Area, Tak Luk waterfall C Laos Not Listed Vulnerable Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505 NA Hipposideros khaokhouayensis; Hipposideros khaokhouayensis; Hipposideros khaokhouayensis; Hipposideros khaokhouayensis; Hipposideros khaokhouayensis; khaokhouayensis; Phyllorhine du Khao Khouay; Phou-Khao-Khouay-Rundblattnase; Hiposidérido de Khao Khouay; Phou Khao Khouay Roundleaf Bat; Phou Khao Khouay Leaf-nosed Bat; Phou Khao Khouay Roundleaf Bat; Phou Khao Khouay Leaf-nosed Bat; H. khaokhouayensis