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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L402	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	Hipposideros halophyllus	Hipposideros halophyllus	Hipposideros halophyllus	Hipposideros halophyllus	Hipposideros halophyllus	Hipposideros halophyllus	Hipposideros halophyllus	Hipposideros halophyllus	Hipposideros halophyllus	Hipposideros halophyllus	Hipposideros halophyllus	Hipposideros halophyllus	Hipposideros halophyllus		[MSW3] bicolor species group.; [HMW] Hipposideros halophyllus HM & Yenbutra, 1984 , “Khao Sa Moa Khon [= Khao Samor Khon], Tha Woong, Lop Buri , Thailand .” Hipposideros halophyllus was formerly included in the bicolor species group, but is now placed in the new ater species group. Monotypic.; [batnames2022]  bicolor species group.; [batnames2023]  bicolor species group.; [batnames2025_1.7] bicolorspecies group.														halophyllus				halophyllus 	halophyllus 			halophyllus J. Edwards Hill & Yenbutra, 1984						N/A					Distribution: Known only from central Thai land.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Thailand; ref. 4.36	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Hill and Yenbutra	1984	Bull. Brit. Mus. (Nat. Hist.) Zool., ^7:77.		Thailand.	Thailand, Lop Buri, Tha Woong, Khao Sa Moa Khan.		HILL & YENBUTRA	1984	Internarial septum expanded into a small disc-like structure. Tympanic bullae elon gate and at right angles to transversely elongate periotics. Size small (forearm length, 35-39 mm).	Distribution: Known only from central Thai land.	No subspecies.		60	species	H. halophyllus	HILL & YENBUTRA	1984	Hipposideros	genus	Hipposideros halophyllus				Internarial septum expanded into a small disc-like structure. Tympanic bullae elon gate and at right angles to transversely elongate periotics. Size small (forearm length, 35-39 mm).	No subspecies.		7. H. halophyllus HILL & YENBUTRA 1984 [bicolor group].	7	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Hipposideros halophyllus	Hipposideros		halophyllus	Hill and Yenbutra		1984		Bull. Brit. Mus. (Nat. Hist.) Zool.	47		77		Thailand Leaf-nosed Bat	Thailand, Lop Buri, Tha Woong, Khao Sa Moa Khan.	Thailand.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).		bicolor species group.	03BD87A2C66DA218F870F0C0FE694488	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	254	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C66DA218F870F0C0FE694488.xml	Hipposideros halophyllus	Hipposideridae	Hipposideros	halophyllus	HM & Yenbutra	1984	Phyllorhine de Thaïlande @fr | Thailand-Rundblattnase @de | Hiposidérido de Tailandia @es | Thailand Roundleaf Bat @en	Hipposideros halophyllus HM & Yenbutra, 1984 , “Khao Sa Moa Khon [= Khao Samor Khon], Tha Woong, Lop Buri , Thailand .” Hipposideros halophyllus was formerly included in the bicolor species group, but is now placed in the new ater species group. Monotypic.	Scattered records in N, C & S Thailand ( Chiang Mai , Phitsanulok , Uthai Thani , Lop Buri , Kanchanaburi , Ratchaburi , Petchaburi, Sa Kaeo , Phang Nga , and Nakhon Si Thammarat provinces) and N Peninsular Malaysia ( Perlis State ).	Forearm 36-39 mm. Noseleaf of the Thailand Leaf-nosed Bat lacks supplementary leaflet. Intemarial septum is kidney-shaped. Pelage is grayish brown. Rostrum is poorly inflated; anterior part is low and gradually slopes upward to pos-tenor part. P2 is small, situated the tooth row or slightly extruded. Baculum is very small, with straight shaft, slightly expanded base, and rounded tip.	Thailand Leaf-nosed Bats forage in a variety of habitats, from shrubs near ricefields and plantations, to mangroves, bamboo and primary evergreen forests. Found from sea level up to 480 m .	The Thailand Leafnosed Bat feeds on small insects over ricefields or by the edge of vegetation.	Females were found pregnant in March-April. Lactating females were captured in May-June, which is the same period that groups of young were observed roosting in caves during the night.	The Thailand Leaf-nosed Bat roosts in caves or rock crevices. Echolocation call frequency of 156-187 kHz has been reported.	The Thailand Leaf-nosed Bat roosts in small colonies of up to c.200 individuals in a single cave (e.g. Pha Daeng and Khao Singto caves); however, in Khao Samor Khon caves, a population size of c. 1000-1400 individuals was estimated. The species is commonly found coexisting with Least Leafnosed Bats { H. cineraceus ), Bicolored Leaf-nosed Bats ( H. bicolor ), Andersen’s Leafnosed Bats (H. gentilis ), Cantor’s Leaf-nosed Bats { H. galeritus ), Great Himalayan Leafnosed Bats { H. armiger ), and Horsfield’s Leaf-nosed Bats ( H. larvatus ).	Classified as Vulnerable on The IUCN Red List. The major threats to the Thailand Leaf-nosed Bat appear to be habitat loss due to limestone mining, habitat degradation by livestock and deforestation, and human disturbance. The species was reported as being hunted at Khao Samor Khon caves, and being disturbed by tourist activities in Ton Chan and Khao Yoi caves.	Douangboubpha & Soisook (2016) | Douangboubpha, Bumrungsri, Soisook, Murray et al. (2010) | Francis (2008a)	https://zenodo.org/record/3749183/files/figure.png	74. Thailand Leaf-nosed Bat Hipposideros halophyllus French: Phyllorhine deThaïlande / German: Thailand-Rundblattnase I Spanish: Hiposidérido deTailandia Other common names: Thailand Roundleaf Bat Taxonomy. Hipposideros halophyllus HM & Yenbutra, 1984 , “Khao Sa Moa Khon [= Khao Samor Khon], Tha Woong, Lop Buri , Thailand .” Hipposideros halophyllus was formerly included in the bicolor species group, but is now placed in the new ater species group. Monotypic. Distribution. Scattered records in N, C & S Thailand ( Chiang Mai , Phitsanulok , Uthai Thani , Lop Buri , Kanchanaburi , Ratchaburi , Petchaburi, Sa Kaeo , Phang Nga , and Nakhon Si Thammarat provinces) and N Peninsular Malaysia ( Perlis State ). Descriptive notes. Forearm 36-39 mm. Noseleaf of the Thailand Leaf-nosed Bat lacks supplementary leaflet. Intemarial septum is kidney-shaped. Pelage is grayish brown. Rostrum is poorly inflated; anterior part is low and gradually slopes upward to pos-tenor part. P2 is small, situated the tooth row or slightly extruded. Baculum is very small, with straight shaft, slightly expanded base, and rounded tip. Habitat. Thailand Leaf-nosed Bats forage in a variety of habitats, from shrubs near ricefields and plantations, to mangroves, bamboo and primary evergreen forests. Found from sea level up to 480 m . Food and Feeding. The Thailand Leafnosed Bat feeds on small insects over ricefields or by the edge of vegetation. Breeding. Females were found pregnant in March-April. Lactating females were captured in May-June, which is the same period that groups of young were observed roosting in caves during the night. Activity patterns. The Thailand Leaf-nosed Bat roosts in caves or rock crevices. Echolocation call frequency of 156-187 kHz has been reported. Movements, Home range and Social organization. The Thailand Leaf-nosed Bat roosts in small colonies of up to c.200 individuals in a single cave (e.g. Pha Daeng and Khao Singto caves); however, in Khao Samor Khon caves, a population size of c. 1000-1400 individuals was estimated. The species is commonly found coexisting with Least Leafnosed Bats { H. cineraceus ), Bicolored Leaf-nosed Bats ( H. bicolor ), Andersen’s Leafnosed Bats (H. gentilis ), Cantor’s Leaf-nosed Bats { H. galeritus ), Great Himalayan Leafnosed Bats { H. armiger ), and Horsfield’s Leaf-nosed Bats ( H. larvatus ). Status and Conservation. Classified as Vulnerable on The IUCN Red List. The major threats to the Thailand Leaf-nosed Bat appear to be habitat loss due to limestone mining, habitat degradation by livestock and deforestation, and human disturbance. The species was reported as being hunted at Khao Samor Khon caves, and being disturbed by tourist activities in Ton Chan and Khao Yoi caves. Bibliography. Douangboubpha & Soisook (2016), Douangboubpha, Bumrungsri, Soisook, Murray et al. (2010), Francis (2008a).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Hipposideros halophyllus	Hipposideros		halophyllus	Hill & Yenbutra	1984	0	Bull. Brit. Mus. (Nat. Hist.) Zool.	48:17:00	Thailand Leaf-nosed Bat	None.	Thailand, Lop Buri, Tha Woong, Khao Sa Moa Khan.	Thailand.	Not listed.	Vulnerable	 bicolor species group.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hipposideros halophyllus	23	Thailand Leaf-nosed Bat	Thailand Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Hipposideros	NA	halophyllus	J. Edwards Hill & Yenbutra	1984	0	Hipposideros_halophyllus	Hill, J. E., & Yenbutra, S. (1984). A new species of the Hipposideros bicolor group (Chiroptera: Hipposideridae) from Thailand. Bulletin of the British Museum (Natural History) Zoology, 47, 76.	https://www.biodiversitylibrary.org/item/19472#page/89/mode/1up	CTNRC 54-3694		"Khao Sa Moa Khon [= Khao Samor Khon], Tha Woong, Lop Buri, Thailand."			halophyllus J. Edwards Hill & Yenbutra, 1984	NA	NA	Thailand|Malaysia	Asia	Indomalaya	VU	0	0	0	Hipposideros_halophyllus	0	sciname match	Hipposideros_halophyllus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10137	Hipposideros halophyllus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Hipposideros	halophyllus	Hill &; Yenbutra, 1984		20000000	Hipposideros halophyllus	Vulnerable	C1	2016	2015-12-07 00:00:00 UTC	3.1	English	<p><span lang="EN-US">This species is assessed as Vulnerable because the population is estimated to be less than 10,000 individuals, and it is expected to decline by 15% in the next 15 years (three generations).</p>	<p><span lang="EN-US">The species requires a very specific roosting habitat of limestone caves in lowland area with an elevation range of a sea level to 480 m, and with small entrances which, in most cases, are underground (Douangboubpha et al. 2010). <span lang="EN-US">The surrounding vegetation included a range of forest types; hill evergreen, mixed deciduous, dipterocarp, bamboo and mangrove (Douangboubpha et al. 2010).<span lang="EN-US"> The species does not travel more than 2 km from the cave to forage. It is unlikely that there is any mix between the known populations as the areas between them are rice fields which are unsuitable for the species (Waengsothorn et al. 2006).</span></p>	<p><span lang="EN-US">There has been a loss of limestone habitat in the area due to limestone mining (S. Bumrungsri pers. comm). In addition most known populations are outside protected areas and the forest habitat is highly disturbed due to livestock and deforestation (S. Bumrungsri pers. comm). Known localities are severely fragmented and surrounded by urban areas or rice fields where pesticide use is high. The bats were reported being hunted by local hunters at Khao Samor Khon (Lop Buri) and disturbed by tourist activities in Ton Chan Cave (Sara Buri) and Khao Yoi Cave (Petcha Buri) (Douangboubpha et al. 2010).</p>	<p><span lang="EN-US">The largest population of this species at Khao Samor Khon, Lop Buri, Thailand is estimated as 1,000-1,400 individuals (Waengsothorn et al. 2006). Other populations, however, are much less abundant. The second largest populations are in Chiang Mai (Pha Daeng Cave) and Sa Kaeo (Khao Singto) but estimated population size at both sites are less than 200 individuals (P. Soisook pers. comm). ; ;</p>	Decreasing	<p><span lang="EN-US">This species currently is known from northern Thailand (Chiang Mai) to northern peninsular Malaysia (Perlis) (Douangboubpha et al. 2010).</p>		Terrestrial	<p><span lang="EN-US">Only five known localities of this species are inside protected area (Douangboubpha et al. 2010). The areas of suitable habitat, particularly the largest population site – Khao Samor Khon, require protection (S Bumrungsri pers. comm).</p>	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Hipposideros		halophyllus	Hill & Yenbutra	1984	0	Bull. Brit. Mus. (Nat. Hist.) Zool.	48:17:00	Thailand Leaf-nosed Bat	None.	Thailand, Lop Buri, Tha Woong, Khao Sa Moa Khan.	Thailand.	Not listed.	Vulnerable	 bicolor species group.	Hipposideros halophyllus	1004604	23	Thailand Leaf-nosed Bat	Thailand Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Hipposideros	NA	halophyllus	J. Edwards Hill & Yenbutra	1984	0	Hipposideros_halophyllus	Hill, J. E., & Yenbutra, S. (1984). A new species of the Hipposideros bicolor group (Chiroptera: Hipposideridae) from Thailand. Bulletin of the British Museum (Natural History) Zoology, 47, 76.	https://www.biodiversitylibrary.org/item/19472#page/89/mode/1up	CTNRC 54-3694		"Khao Sa Moa Khon [= Khao Samor Khon], Tha Woong, Lop Buri, Thailand."			halophyllus J. Edwards Hill & Yenbutra, 1984	NA	NA				Thailand|Malaysia	Asia	Indomalaya	VU	0	0	0	Hipposideros_halophyllus	0	sciname match	Hipposideros_halophyllus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hipposideros_halophyllus	1004604	23	Thailand Roundleaf Bat	Thailand Roundleaf Bat|Thailand Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Hipposideros	NA	halophyllus	J. Edwards Hill & Yenbutra	0	Hipposideros halophyllus	Hill, J.E. and Yenbutra, S. 1984. A new species of the _Hipposideros bicolor_ group (Chiroptera: Hipposideridae) from Thailand. Bulletin of the British Museum (Natural History) (Zoology)47(1):77-82.	https://www.biodiversitylibrary.org/page/2281561	TNRC 54-3694	holotype		"Khao Sa Moa Khon [= Khao Samor Khon], Tha Woong, Lop Buri, Thailand."			NA	NA				Thailand|Malaysia	Asia	Indomalaya	VU	0	0	0	Hipposideros_halophyllus	0	sciname match	Hipposideros_halophyllus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Hipposideros		halophyllus	Hill & Yenbutra	1984	0	Bull. Brit. Mus. (Nat. Hist.) Zool.	48:17:00	Thailand Leaf-nosed Bat	None.	Thailand, Lop Buri, Tha Woong, Khao Sa Moa Khan.	Thailand.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10137/22092544/' target='_blank'>Vulnerable</a>	bicolorspecies group.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hipposideros halophyllus; Hipposideros halophyllus; Hipposideros halophyllus; Hipposideros halophyllus; Hipposideros halophyllus; Hipposideros halophyllus; halophyllus; Phyllorhine de Thaïlande; Thailand-Rundblattnase; Hiposidérido de Tailandia; Thailand Roundleaf Bat; Thailand Leaf-nosed Bat; Thailand Roundleaf Bat; Thailand Leaf-nosed Bat; Thailand Leaf-nosed Bat; H. halophyllus