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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L401	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Hipposideros griffini	Hipposideros griffini	Hipposideros griffini	Hipposideros griffini	Hipposideros griffini	Hipposideros griffini	Hipposideros griffini	Hipposideros griffini	Hipposideros griffini		[HMW] Hipposideros griffini Thong et aL, 2012 , “Cat Ba National Park, Cat Ba Island, Ha Long Bay, Vietnam , 20°48’N , 107°01'E , 248 m above sea level (m a.s.l.).” Hipposideros griffini is in the armiger species group. Monotypic.; [MDD2022] recently described; [MDD2023] recently described; [MDD2025_2.0] recently described; [MDD2025_2.2] recently described														griffini				griffini	griffini			griffini Vu Dinh Thong, Puechmaille, Denzinger, Dietz, Csorba, P. J. J. Bates, E. C. Teeling, & Schnitzler, 2012						N/A																																								NA																											03BD87A2C67DA20FFF31F64EFB1C5151	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	238	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C67DA20FFF31F64EFB1C5151.xml	Hipposideros griffini	Hipposideridae	Hipposideros	griffini	Thong, Puechmaille, Denzinger, Dietz, Csorba, Bates, Teeling & Schnitzler	2012	Phyllorhine de Griffin @fr | Griffin-Rundblattnase @de | Hiposidérido de Griffin @es | Griffin's Roundleaf Bat @en	Hipposideros griffini Thong et aL, 2012 , “Cat Ba National Park, Cat Ba Island, Ha Long Bay, Vietnam , 20°48’N , 107°01'E , 248 m above sea level (m a.s.l.).” Hipposideros griffini is in the armiger species group. Monotypic.	NE & C Vietnam , known only from Cat Ba I, Ha Long Bay ( Haiphong Province ) and Chu Mom Ray National Park (Kon Turn Province) .	Ear 27-5-30 mm, hindfoot 14-1-15-8 mm, forearm 83-3-90 mm; weight 44 g . Pelage of Griffin’s Leaf-nosed Bat is brownish or grayish, being darker on dorsum than on ventral area; on each hair, distal part is also darker than basal portion. This species presents four pairs of supplementary lateral leaflets. Second pair of leaflets is longest and widest. Fourth pair of leaflets is shortest and least pronounced, whereas posterior part of third leaflet is merged with an excrescence. Upper edge of posterior leaf is cusp-shaped. Ears are triangular and tail is long. Males present a noticeable excrescence and a sexual sac, which become more pronounced during breeding season.	Both primary and degraded forest. Griffin’s Leaf-nosed Bat has been reported in karstic mountain areas, as well as in lowlands.	Foraging behavior and diet are still unknown for Griffin’s Leafnosed Bat, but it probably forages in primary as well as disturbed forests; its diet may be based on insects.	One young was captured in June. Males have been reported breeding in August at Chu Mom Ray National Park.	Griffin’s Leaf-nosed Bat has been reported roosting in rock piles, crevices, caves, old mines, and hollow trees. Echolocation call frequency is c.76-6— 79-2 kHz.	Griffin’s Leaf-nosed Bat roosts alone or in pairs.	Not assessed on 7%e IUCNRed List due to its recent recognition as a species. More research on the ecology and population size and trends of Griffin’s Leaf-nosed Bat is needed to permit the establishment of adequate conservation and management measures.	Thong Vu Dinh, Puechmaille, Denzinger, Dietz et al. (2012)		28. Griffin’s Leaf-nosed Bat Hipposideros griffini French: Phyllorhine de Griffin / German: Griffin-Rundblattnase / Spanish: Hiposidérido de Griffin Other common names: Griffin's Roundleaf Bat Taxonomy. Hipposideros griffini Thong et aL, 2012 , “Cat Ba National Park, Cat Ba Island, Ha Long Bay, Vietnam , 20°48’N , 107°01'E , 248 m above sea level (m a.s.l.).” Hipposideros griffini is in the armiger species group. Monotypic. Distribution. NE & C Vietnam , known only from Cat Ba I, Ha Long Bay ( Haiphong Province ) and Chu Mom Ray National Park (Kon Turn Province) . Descriptive notes. Ear 27-5-30 mm, hindfoot 14-1-15-8 mm, forearm 83-3-90 mm; weight 44 g . Pelage of Griffin’s Leaf-nosed Bat is brownish or grayish, being darker on dorsum than on ventral area; on each hair, distal part is also darker than basal portion. This species presents four pairs of supplementary lateral leaflets. Second pair of leaflets is longest and widest. Fourth pair of leaflets is shortest and least pronounced, whereas posterior part of third leaflet is merged with an excrescence. Upper edge of posterior leaf is cusp-shaped. Ears are triangular and tail is long. Males present a noticeable excrescence and a sexual sac, which become more pronounced during breeding season. Habitat. Both primary and degraded forest. Griffin’s Leaf-nosed Bat has been reported in karstic mountain areas, as well as in lowlands. Food and Feeding. Foraging behavior and diet are still unknown for Griffin’s Leafnosed Bat, but it probably forages in primary as well as disturbed forests; its diet may be based on insects. Breeding. One young was captured in June. Males have been reported breeding in August at Chu Mom Ray National Park. Activity patterns. Griffin’s Leaf-nosed Bat has been reported roosting in rock piles, crevices, caves, old mines, and hollow trees. Echolocation call frequency is c.76-6— 79-2 kHz. Movements, Home range and Social organization. Griffin’s Leaf-nosed Bat roosts alone or in pairs. Status and Conservation. Not assessed on 7%e IUCNRed List due to its recent recognition as a species. More research on the ecology and population size and trends of Griffin’s Leaf-nosed Bat is needed to permit the establishment of adequate conservation and management measures. Bibliography. Thong Vu Dinh, Puechmaille, Denzinger, Dietz et al. (2012)	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Hipposideros griffini	Hipposideros		griffini	Thong, Puechmaille, Denzinger, Dietz, Csorba, Bates, Teeling & Schnitzler	2012	0	J. Mammal.	93(1): 3	Griffin&apos;s Leaf-nosed bat	None.	Vietnam, Ha Long Bay, Cat Ba Island, Cat Ba National Park	Vietnam	Not listed.	Near Threatened		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hipposideros griffini	23	Griffin's Leaf-nosed Bat	Griffin's Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Hipposideros	NA	griffini	Vu Dinh Thong, Puechmaille, Denzinger, Dietz, Csorba, P. J. J. Bates, Teeling, & Schnitzler	2012	0	Hipposideros_griffini	Thong, V. D., Puechmaille, S. J., Denzinger, A., Dietz, C., Csorba, G., Bates, P. J., ... & Schnitzler, H. U. (2012). A new species of Hipposideros (Chiroptera: Hipposideridae) from Vietnam. Journal of Mammalogy, 93(1), 3.	https://academic.oup.com/jmammal/article/93/1/1/894060	IEBR T.200809.12		"Cat Ba National Park, Cat Ba Island, Ha Long Bay, Vietnam, 20Â°48'N, 107Â°01'E, 248 m above sea level (m a.s.l.)."	20.8	107.02	griffini Vu Dinh Thong, Puechmaille, Denzinger, Dietz, Csorba, P. J. J. Bates, Teeling, & Schnitzler, 2012	recently described	Thong, V. D., Puechmaille, S. J., Denzinger, A., Dietz, C., Csorba, G., Bates, P. J., ... & Schnitzler, H. U. (2012). A new species of Hipposideros (Chiroptera: Hipposideridae) from Vietnam. Journal of Mammalogy, 93(1), 1-11.	Vietnam	Asia	Indomalaya	NT	0	0	0	Hipposideros_griffini	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Hipposideros griffini	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Hipposideros	griffini	Vu Dinh Trong, Puechmaille, Denzinger, Dietz, Csorba, Bates, Teeling &; Schnitzler, 2012		100000000	Hipposideros griffini	Near Threatened	A2c	2019	2018-08-27 00:00:00 UTC	3.1	English	The species is listed as Near Threatened because its population is likely declining at a rate of 20% to 30% over the last 15 years (proposed generation length of 5 years based on similar species, Pacifici et al. 2013) due to past and continuing disturbance of human activities to the roosting caves and degradation and loss of foraging habitat in forests.	The distribution of this species ranges from northern to southern Vietnam in three remarkably disjunct areas, including: the undisturbed forests in the limestone karst of Cat Ba National Park, secondary forests in a mountainous area in Chu Mom Ray in central Vietnam (Thong et al. 2012), and lowland forests in Cat Tien in southern Vietnam (Thong 2012). It is likely that to occur in areas that lie between these three isolated sites. There is no information about roosting sites of the species in northern and south-central regions.  A cave roost in southern Vietnam is the only known roosting site of a group up to 30 individuals. Recordings of the species in Cat Ba and Chu Mom Ray suggests that the species forages along trails under the forest canopy.	Threats to the species in northern and south-central Viet Nam are unclear but the roosting site in southern Viet Nam is likely disturbed by cave visiting and ecotourism. Foraging habitats in forests also face continued degradation and loss due to harvest.	Little is known about the population size of this species, but it is likely that the population is declining due to disturbance of tourism and habitat changes. Four individuals were captured in Cat Ba National Park, two individuals from Chu Mom Ray National Park (Thong et al. 2012) and several individuals from Cat Tien National Park (Thong 2012). It was observed in a group of less than 30 individuals inhabiting a cave in Cat Tien National Park (Thong, unpublished data). It is thought that there are at least two subpopulations, one in Cat Ba National Park in northern Viet Nam and at least one in south central and southern Vietnam. A series of surveys between 2014 and 2018 in both Cat Ba and Cat Tien National Park provided no additional information about the species.	Decreasing	This species has only been recorded from three disjunct national parks, namely Cat Ba, Chu Mom Ray (Thong et al. 2012) and Cat Tien (Thong 2012). It is probable that the species range is includes at least some areas that lie between these isolated sites.		Terrestrial	<span id="docs-internal-guid-858f78e4-7fff-e3b6-f1f6-861e1c31d909">All known localities of the species are situated within national parks, where forests are legally protected. Conservation campaigns have been implemented at Cat Ba National Park but outcomes are uncertain. The unscientific development of ecotourism in all three national parks, particularly in Cat Ba, appears as a critical threat to foraging habitats and roosting sites of the species.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Hipposideros		griffini	Thong, Puechmaille, Denzinger, Dietz, Csorba, Bates, Teeling & Schnitzler	2012	0	J. Mammal.	93(1): 3	Griffin&apos;s Leaf-nosed bat	None.	Vietnam, Ha Long Bay, Cat Ba Island, Cat Ba National Park	Vietnam	Not listed.	Near Threatened		Hipposideros griffini	1004603	23	Griffin's Leaf-nosed Bat	Griffin's Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Hipposideros	NA	griffini	Vu Dinh Thong, Puechmaille, Denzinger, Dietz, Csorba, P. J. J. Bates, Teeling, & Schnitzler	2012	0	Hipposideros_griffini	Thong, V. D., Puechmaille, S. J., Denzinger, A., Dietz, C., Csorba, G., Bates, P. J., ... & Schnitzler, H. U. (2012). A new species of Hipposideros (Chiroptera: Hipposideridae) from Vietnam. Journal of Mammalogy, 93(1), 3.	https://academic.oup.com/jmammal/article/93/1/1/894060	IEBR T.200809.12		"Cat Ba National Park, Cat Ba Island, Ha Long Bay, Vietnam, 20Â°48'N, 107Â°01'E, 248 m above sea level (m a.s.l.)."	20.8	107.0167	griffini Vu Dinh Thong, Puechmaille, Denzinger, Dietz, Csorba, P. J. J. Bates, Teeling, & Schnitzler, 2012	recently described	Thong, V. D., Puechmaille, S. J., Denzinger, A., Dietz, C., Csorba, G., Bates, P. J., ... & Schnitzler, H. U. (2012). A new species of Hipposideros (Chiroptera: Hipposideridae) from Vietnam. Journal of Mammalogy, 93(1), 1-11.				Vietnam	Asia	Indomalaya	NT	0	0	0	Hipposideros_griffini	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hipposideros_griffini	1004603	23	Griffin's Roundleaf Bat	Griffin's Roundleaf Bat|Griffin's Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Hipposideros	NA	griffini	Vu Dinh Thong, Puechmaille, Denzinger, Dietz, Csorba, P. J. J. Bates, E. C. Teeling, & Schnitzler	0	Hipposideros griffini	Thong, V.D., Puechmaille, S.J., Denzinger, A., Dietz, C., Csorba, G., Bates, P.J.J., Teeling, E.C. and Schnitzler, H.-U. 2012-02-16. A new species of _Hipposideros_ (Chiroptera: Hipposideridae) from Vietnam. Journal of Mammalogy 93(1):1-11.	https://doi.org/10.1644/11-MAMM-A-073.1	IEBR-T.200809.12	holotype		"Cat Ba National Park, Cat Ba Island, Ha Long Bay, Vietnam, 20Â°48'N, 107Â°01'E, 248 m above sea level (m a.s.l.)."	20.8	107.0167	recently described	Thong, V. D., Puechmaille, S. J., Denzinger, A., Dietz, C., Csorba, G., Bates, P. J., ... & Schnitzler, H. U. (2012). A new species of Hipposideros (Chiroptera: Hipposideridae) from Vietnam. Journal of Mammalogy, 93(1), 1-11.				Vietnam	Asia	Indomalaya	NT	0	0	0	Hipposideros_griffini	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Hipposideros		griffini	Thong, Puechmaille, Denzinger, Dietz, Csorba, Bates, Teeling & Schnitzler	2012	0	J. Mammal.	93(1): 3	Griffin&apos;s Leaf-nosed bat	None.	Vietnam, Ha Long Bay, Cat Ba Island, Cat Ba National Park	Vietnam	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/80222915/95642190/' target='_blank'>Near Threatened</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hipposideros griffini; Hipposideros griffini; Hipposideros griffini; Hipposideros griffini; Hipposideros griffini; griffini; Phyllorhine de Griffin; Griffin-Rundblattnase; Hiposidérido de Griffin; Griffin's Roundleaf Bat; Griffin's Leaf-nosed Bat; Griffin's Roundleaf Bat; Griffin&apos;s Leaf-nosed bat; H. griffini
