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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L393	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	Hipposideros edwardshilli	Doryrhina edwardshilli	Hipposideros edwardshilli	Doryrhina edwardshilli	Hipposideros edwardshilli	Hipposideros edwardshilli	Hipposideros edwardshilli	Hipposideros edwardshilli	Hipposideros edwardshilli	Hipposideros edwardshilli		[MSW3] cyclops species group. See Flannery (1995a) and Bonaccorso (1998).; [HMW] Hipposideros edwardshilli Flannery & Colgan, 1993 , “Imonda Station ( 3°20’S 141°10’E ), Bewani Mountains, Papua New Guinea at an altitude of 240 m .” This species is monotypic.; [batnames2022] Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclops species group. See Flannery (1995a) and Bonaccorso (1998).; [MDD2022] moved from Hipposideros to Doryrhina; [batnames2023] Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclops species group. See Flannery (1995a) and Bonaccorso (1998).; [MDD2023] has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being; [MDD2025_2.0] has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being; [batnames2025_1.7] Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclopsspecies group. See Flannery (1995a) and Bonaccorso (1998).; [MDD2025_2.2] has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being														edwardshilli				edwardshilli 	edwardshilli 			edwardshilli Flannery & D. J. Colgan, 1993						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Hipposideros edwardshilli	Hipposideros		edwardshilli	Flannery and Colgan		1993		Rec. Aust. Mus.	45		45		Hill's Leaf-nosed Bat	Papua New Guinea, W Sepik, Bewani Mtns., Imonda Sta.	NW Papua New Guinea.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).		cyclops species group. See Flannery (1995a) and Bonaccorso (1998).	03BD87A2C675A200F87CEEF1F91D4A77	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	230	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C675A200F87CEEF1F91D4A77.xml	Doryrhina edwardshilli	Hipposideridae	Doryrhina	edwardshilli		1993	Phyllorhine d'Edwards Hill @fr | Hill-Rundblattnase @de | Doryrina de Edwards Hill @es | Hill's Roundleaf Bat @en	Hipposideros edwardshilli Flannery & Colgan, 1993 , “Imonda Station ( 3°20’S 141°10’E ), Bewani Mountains, Papua New Guinea at an altitude of 240 m .” This species is monotypic.	New Guinea , known only from three zones of the Bewani Mts, North Coastal Range, Sandaun Province.	Head-body 50-6 mm, tail 11-1-15 mm, ear 19-3-21-3 mm, hindfoot 8-2 mm, forearm 49-6-51 mm; weight 11 g . Hill’s Leaf-nosed Bat is similar to the Telefomin Leaf-nosed Bat (D. corynophyUus), but with darkish brown or reddish fur on dorsum, and more cinnamon color on ventral area; subadults can present grayish coloration. Fur is markedly shorter than on the Telefomin Leaf-nosed Bat: c.8-7 mm long on mid-back. Noseleaf differs in lower club being less pronounced, shorter and laterally flattened, as compared with that of the Telefomin Leaf-nosed Bat.	Hill’s Leaf-nosed Bat is a cave-roosting species reported in low hills. It can be present in forest and also in more modified and artificial environments such as gardens or rural village areas, although it always needs cave roosts nearby. It has only been reported at altitudes of c.200-300 m, which accounts for its restricted range.	Insects constitute its main food. Hill’s Leaf-nosed Bat has been reported foraging in forests, rural village areas, and gardens.	No information.	No information.	Unlike the Telefomin Leaf-nosed Bat, Hill’s Leaf-nosed Bat has not been found sharing caves with Wollaston’s Leafnosed Bat ( D. wollastoni ).	Classified as Vulnerable on The IUCN Red List (as Hipposideros edwardshiUi ). Population size and trends of Hill’s Leaf-nosed Bat remain unknown. Main threats probably include habitat degradation and transformation of native forest to agricultural lands around Bewani Mountains. In addition, the fact that the North Coastal Range is not limestone might reduce the roost availability, restricting the potential range ( 100—2000 km 2) even more. More research on this species is needed to assess its population status accurately and to discover details of its ecology.	Armstrong & Aplin (2017c) | Bonaccorso (1998) | Decher & ahr (2005) | lannery & Colgan (1993)	https://zenodo.org/record/3749215/files/figure.png	8. Hill’s Leaf-nosed Bat Doryrhina edwardshilli French: Phyllorhine d'Edwards Hill / German: Hill-Rundblattnase I Spanish: Doryrina de Edwards Hill Other common names: Hill's Roundleaf Bat Taxonomy. Hipposideros edwardshilli Flannery & Colgan, 1993 , “Imonda Station ( 3°20’S 141°10’E ), Bewani Mountains, Papua New Guinea at an altitude of 240 m .” This species is monotypic. Distribution. New Guinea , known only from three zones of the Bewani Mts, North Coastal Range, Sandaun Province. Descriptive notes. Head-body 50-6 mm, tail 11-1-15 mm, ear 19-3-21-3 mm, hindfoot 8-2 mm, forearm 49-6-51 mm; weight 11 g . Hill’s Leaf-nosed Bat is similar to the Telefomin Leaf-nosed Bat (D. corynophyUus), but with darkish brown or reddish fur on dorsum, and more cinnamon color on ventral area; subadults can present grayish coloration. Fur is markedly shorter than on the Telefomin Leaf-nosed Bat: c.8-7 mm long on mid-back. Noseleaf differs in lower club being less pronounced, shorter and laterally flattened, as compared with that of the Telefomin Leaf-nosed Bat. Habitat. Hill’s Leaf-nosed Bat is a cave-roosting species reported in low hills. It can be present in forest and also in more modified and artificial environments such as gardens or rural village areas, although it always needs cave roosts nearby. It has only been reported at altitudes of c.200-300 m, which accounts for its restricted range. Food and Feeding. Insects constitute its main food. Hill’s Leaf-nosed Bat has been reported foraging in forests, rural village areas, and gardens. Breeding. No information. Activity patterns. No information. Movements, Home range and Social organization. Unlike the Telefomin Leaf-nosed Bat, Hill’s Leaf-nosed Bat has not been found sharing caves with Wollaston’s Leafnosed Bat ( D. wollastoni ). Status and Conservation. Classified as Vulnerable on The IUCN Red List (as Hipposideros edwardshiUi ). Population size and trends of Hill’s Leaf-nosed Bat remain unknown. Main threats probably include habitat degradation and transformation of native forest to agricultural lands around Bewani Mountains. In addition, the fact that the North Coastal Range is not limestone might reduce the roost availability, restricting the potential range ( 100—2000 km 2) even more. More research on this species is needed to assess its population status accurately and to discover details of its ecology. Bibliography. Armstrong & Aplin (2017c), Bonaccorso (1998), Decher & ahr (2005), lannery & Colgan (1993).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Hipposideros edwardshilli	Hipposideros		edwardshilli	Flannery & Colgan	1993	0	Rec. Aust. Mus.	######	Hill's Leaf-nosed Bat	None.	Papua New Guinea, W Sepik, Bewani Mtns., Imonda Sta.	NW Papua New Guinea.	Not listed.	Vulnerable	Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclops species group. See Flannery (1995a) and Bonaccorso (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Doryrhina edwardshilli	23	Hill's Leaf-nosed Bat	Hill's Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Doryrhina	NA	edwardshilli	Flannery & Colgan	1993	1	Hipposideros_edwardshilli	Flannery, T. F. & Colgan, D. J. (1993). A new species and two new subspecies of Hipposideros (Chiroptera) from Western Papua New Guinea. Records of the Western Australia, 45(1), 45.	https://journals.australian.museum/flannery-and-colgan-1993-rec-aust-mus-451-4357/	AM M.21752		"Imonda Station (3°20'S 141°10'E), Bewani Mountains, Papua New Guinea at an altitude of 240 m."	-3.333	141.17	edwardshilli (Flannery & Colgan, 1993)	moved from Hipposideros to Doryrhina	Foley, N. M., Goodman, S. M., Whelan, C. V., Puechmaille, S. J., & Teeling, E. (2017). Towards navigating the Minotaur's labyrinth: cryptic diversity and taxonomic revision within the speciose genus Hipposideros (Hipposideridae). Acta Chiropterologica, 19(1), 1-18.	Papua New Guinea	Oceania	Australasia/Oceania	VU	0	0	0	Doryrhina_edwardshilli	0	sciname match	Hipposideros_edwardshilli	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10133	Hipposideros edwardshilli	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Hipposideros	edwardshilli	Flannery &; Colgan, 1993		200000000	Hipposideros edwardshilli	Vulnerable	B2ab(iii)	2021	2016-01-31 00:00:00 UTC	3.1	English	No new records of Hipposideros edwardshilli have appeared since the species was first described from the Bewani Mountains of the North Coastal Range in Sandaun (West Sepik) Province of Papua New Guinea (Flannery and Colgan 1993). A lack of information on its echolocation call frequency, as well as the presence of geographic variation in the call frequency of H. wollastoni parnabyi and a lack of information about the calls of H. w. fasensis and H. semoni , has limited the potential to detect it on acoustic surveys in areas further afield (K.N. Armstrong and K.P. Aplin unpublished data). However, given that it has been recorded only at 200-300 m asl, it does have the potential to range further than the Bewani Mountains at similar elevations throughout the North Coastal Range and Sepik catchment, and west into the Cyclops Mountains of the Indonesian Papua Province. This is a cave roosting species, but much of the North Coastal Range is not limestone, so habitat degradation combined with an anticipated limited opportunity for roosting could restrict its area of occupancy (AOO) to less than 2,000 km² around its currently known range. The absence of new records despite some recent survey work, coupled with continued clearing and the potential for forest conversion to monoculture farming in the region around the Bewani Mountains has prompted a more precautionary assessment than the previous one.	<p>This species roosts in caves in low hills and has been observed foraging for insects in rural villages and gardens (Flannery 1995; Bonaccorso 1998). It will presumably still require cave roosts within close proximity to modified habitats such as gardens to be present. ;</p>	<p></p><p> </p><p>The species is currently known from only a few sites. It is unclear whether or not it is threatened at these localities, however there is ongoing, as well as likely future, clearing and conversion of native forest habitats in and around the Bewani Mountains and the Sepik catchment. ;</p> <p></p>	The population size and trend for this species is not known.	Unknown	<p>This recently described species has been recorded from three localities in the Bewani Mountains of the North Coastal Range of Sandaun Province, Papua New Guinea (Flannery and Colgan 1993, Flannery 1995, Bonaccorso 1998). It has been found at elevations of 200 to 300 m asl. No further records of this species have emerged, and the available evidence (K.P. Aplin and K.N. Armstrong unpublished data) so far suggests that Hipposideros edwardshilli is genuinely limited to the northern coastal ranges in Papua New Guinea and possibly Papua Province of Indonesia. </p>		Terrestrial	<p>A suitable conservation measure for this poorly-known species would be the protection of the few known roosting sites. Further research on their use of modified landscapes, targeted surveys for roosts and characterisation of echolocation calls to aid wider surveys would be helpful initial efforts for this species. Surveys throughout the North Coastal Range in Papua New Guinea and the Cyclops Mountains in Papua Province of Indonesia would be helpful in clarifying its extent of occurrence. ; ;</p>	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Hipposideros		edwardshilli	Flannery & Colgan	1993	0	Rec. Aust. Mus.	45:45:00	Hill's Leaf-nosed Bat	None.	Papua New Guinea, W Sepik, Bewani Mtns., Imonda Sta.	NW Papua New Guinea.	Not listed.	Vulnerable	Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclops species group. See Flannery (1995a) and Bonaccorso (1998).	Hipposideros edwardshilli	1004569	23	Hill's Leaf-nosed Bat	Hill's Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Hipposideros	NA	edwardshilli	Flannery & Colgan	1993	0	Hipposideros_edwardshilli	Flannery, T. F. & Colgan, D. J. (1993). A new species and two new subspecies of Hipposideros (Chiroptera) from Western Papua New Guinea. Records of the Western Australia, 45(1), 45.	https://journals.australian.museum/flannery-and-colgan-1993-rec-aust-mus-451-4357/	AM M.21752		"Imonda Station (3°20'S 141°10'E), Bewani Mountains, Papua New Guinea at an altitude of 240 m."	-3.33333	141.1667	edwardshilli (Flannery & Colgan, 1993)	has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being	Foley, N. M., Goodman, S. M., Whelan, C. V., Puechmaille, S. J., & Teeling, E. (2017). Towards navigating the Minotaur's labyrinth: cryptic diversity and taxonomic revision within the speciose genus Hipposideros (Hipposideridae). Acta Chiropterologica, 19(1), 1-18.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.				Papua New Guinea	Oceania	Australasia/Oceania	VU	0	0	0	Doryrhina_edwardshilli	0	sciname match	Hipposideros_edwardshilli	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hipposideros_edwardshilli	1004569	23	Hill's Roundleaf Bat	Hill's Roundleaf Bat|Hill's Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Hipposideros	NA	edwardshilli	Flannery & D. J. Colgan	0	Hipposideros edwardshilli	Flannery, T.F. and Colgan, D.J. 1993-03-19. A new species and two new subspecies of _Hipposideros_ (Chiroptera) from Western Papua New Guinea. Records of the Australian Museum 45(1):43-57.	https://doi.org/10.3853/j.0067-1975.45.1993.129	AM M.21752	holotype		"Imonda Station (3°20'S 141°10'E), Bewani Mountains, Papua New Guinea at an altitude of 240 m."	-3.33333	141.1667	has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being	Foley, N. M., Goodman, S. M., Whelan, C. V., Puechmaille, S. J., & Teeling, E. (2017). Towards navigating the Minotaur's labyrinth: cryptic diversity and taxonomic revision within the speciose genus Hipposideros (Hipposideridae). Acta Chiropterologica, 19(1), 1-18.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.				Papua New Guinea	Oceania (Continent)	Australasia	VU	0	0	0	Doryrhina_edwardshilli	0	sciname match	Hipposideros_edwardshilli	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Hipposideros		edwardshilli	Flannery & Colgan	1993	0	Rec. Aust. Mus.	45:45:00	Hill's Leaf-nosed Bat	None.	Papua New Guinea, W Sepik, Bewani Mtns., Imonda Sta.	NW Papua New Guinea.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10133/209537105/' target='_blank'>Vulnerable</a>	Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclopsspecies group. See Flannery (1995a) and Bonaccorso (1998).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hipposideros edwardshilli; Doryrhina edwardshilli; Hipposideros edwardshilli; Doryrhina edwardshilli; Hipposideros edwardshilli; Hipposideros edwardshilli; edwardshilli; Phyllorhine d'Edwards Hill; Hill-Rundblattnase; Doryrina de Edwards Hill; Hill's Roundleaf Bat; Hill's Leaf-nosed Bat; Hill's Roundleaf Bat; Hill's Leaf-nosed Bat; Hill's Leaf-nosed Bat; H. edwardshilli