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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L392	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Hipposideros dyacorum	Hipposideros dyacorum	Hipposideros dyacorum	Hipposideros dyacorum	Hipposideros dyacorum	Hipposideros dyacorum	Hipposideros dyacorum	Hipposideros dyacorum	Hipposideros dyacorum	Hipposideros dyacorum	Hipposideros dyacorum	Hipposideros dyacorum	Hipposideros dyacorum	Hipposideros dyacorum	Hipposideros dyacorum		[MSW3] bicolor species group.; [HMW] Hipposiderus [sic] dyacorum Thomas, 1902 , “Mount Mulu, Baram District, Sarawak ,” Borneo, Malaysia . Hipposideros dyacorum was formerly included in the bicolor species group, but is now placed in the new tzter species group. Monotypic.; [batnames2022]  bicolor species group.; [batnames2023]  bicolor species group.; [batnames2025_1.7] bicolorspecies group.														dyacorum				dyacorum 	dyacorum 			dyacorum O. Thomas, 1902		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Dyak leaf-nosed bat	Borneo	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Hipposideros dyacorum	Malaysia, Sarawak, Baram, Mt. Mulu.	Thomas	1902	Ann. Mag. Nat. Hist., ser. 7, 9:271.	Distribution: Confined to Borneo.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Dyak leaf-nosed bat	Malaya, Borneo; ref. 4.146	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1902	Ann. Mag. Nat. Hist., ser. 7, 9:271.		Borneo, peninsular Thailand.	Malaysia, Sarawak, Baram, Mt. Mulu.		THOMAS	1902	Anterior upper premolar greatly reduced, com pletely extruded from toothrow. Anterior lower premolar only one quarter the length and height of the posterior lower premolar. Anterior leaf without a median emargination. Posterior leaf supported by a well-defined median septum and weaker lateral septa. Pterygoids long, sphenoidal bridge wide. Size relatively small (forearm length, 39-42 mm).	Distribution: Confined to Borneo.	No subspecies.		62	species	H. dyacorum	THOMAS	1902	Hipposideros	genus	Hipposideros dyacorum				Anterior upper premolar greatly reduced, com pletely extruded from toothrow. Anterior lower premolar only one quarter the length and height of the posterior lower premolar. Anterior leaf without a median emargination. Posterior leaf supported by a well-defined median septum and weaker lateral septa. Pterygoids long, sphenoidal bridge wide. Size relatively small (forearm length, 39-42 mm).	No subspecies.		15. H. dyacorum THOMAS 1902 [bicolor group].	15	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Hipposideros dyacorum	Hipposideros		dyacorum	Thomas		1902		Ann. Mag. Nat. Hist., ser. 7	9		271		Dayak Leaf-nosed Bat	Malaysia, Sarawak, Baram, Mt. Mulu.	Borneo (including Sarawak, Malaysia), Peninsular Thailand.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).		bicolor species group.	03BD87A2C66CA21FF8A3F3A3FCC240C7	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	253	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C66CA21FF8A3F3A3FCC240C7.xml	Hipposideros dyacorum	Hipposideridae	Hipposideros	dyacorum	Thomas	1902	Phyllorhine dayak @fr | Dayak-Rundblattnase @de | Hiposidérido dayak @es | Dayak Roundleaf Bat @en	Hipposiderus [sic] dyacorum Thomas, 1902 , “Mount Mulu, Baram District, Sarawak ,” Borneo, Malaysia . Hipposideros dyacorum was formerly included in the bicolor species group, but is now placed in the new tzter species group. Monotypic.		Forearm 38—43 mm . Noseleaf of the Dayak Leaf-nosed Bat is small and rounded, without supplementary leaflet. Intemarial septum is swollen and broader at base, and narrow in middle. Pelage is uniformly dark brown, with slightly paler tips.	Lowland rainforest to semi-evergreen forest.	The Dayak Leaf-nosed Bat feeds on small insects in forest understory.	In peninsular Thailand , lactating females were captured in May-June. On following pages: 71. House-dwelling Leaf-nosed Bat ( Hipposideros einnaythu ); 72. Andersen's Leaf-nosed Bat (( Hipposideros halophyllus }; 75. Kolar Leaf-nosed Bat ( Hipposideros hypophyllus }; 76. Phou Khao Khouay Leaf-nosed nosed Bat ( Hipposideros macrobullatus }; 79. Maggie Taylor's Leaf-nosed Bat ( Hipposideros maggietaylorae ); 80. Malayan 82. Philippine Forest Leaf-nosed Bat ( Hipposideros obscurus ); 83. Orbiculus Leaf-nosed Bat ( Hipposideros orbiculus 86. Philippine Pygmy Leaf-nosed Bat ( Hipposideros pygmaeus }; 87. Ridley’ s Leaf-nosed Bat ( Hipposideros ridleyi }; 88 Hipposideros gentilis ); 73. Fulvus Leaf-nosed Bat ( Hipposideros fulvus }; 74. Thailand Leaf-nosed Bat Bat ( Hipposideros khaokhouayensis }; 77. Kunz's Leaf-nosed Bat ( Hipposideros kunzi }; 78. Big-eared Leaf-Leaf-nosed Bat ( Hipposideros nequam ); 81. Nicobar Leaf-nosed Bat ( Hipposideros nicobarulae };}; 84. Biak Leaf-nosed Bat ( Hipposideros papua }; 85. Pomona Leaf-nosed Bat ( Hipposideros pomona };. Laotian Leaf-nosed Bat ( Hipposideros rotalis }.	T he Dayak Leaf-nosed Bat roosts in limestone caves, under rocks, and in hollow trees. Call frequency of the F segment is 167-170 kHz.	The Dayak Leaf-nosed Bat was caught in the same trap with Kunz’s Leaf-nosed Bats { H. kunzî ), Bicolored Leaf-nosed Bats { H. bicolor ), Bornean Leaf-nosed Bats { H. doriae ), and other forest-dwelling Murino. and Kerivoula.	Classified as Least Concern on TTze IUCNRed List. The major threat to the Dayak Leaf-nosed Bat is habitat loss.	Corbet & Hill (1992) | Francis & Kingston (2008) | Kingston et al. (2006) | Payne et al. (1985) | Phillipps & Phillipps (2016) | Simmons (2005)	https://zenodo.org/record/3749175/files/figure.png	70. Dayak Leaf-nosed Bat Hipposideros dyacorum French: Phyllorhine dayak / German: Dayak-Rundblattnase / Spanish: Hiposidérido dayak Other common names: Dayak Roundleaf Bat Taxonomy. Hipposiderus [sic] dyacorum Thomas, 1902 , “Mount Mulu, Baram District, Sarawak ,” Borneo, Malaysia . Hipposideros dyacorum was formerly included in the bicolor species group, but is now placed in the new tzter species group. Monotypic. Distribution. Malay Peninsula and Borne©:- Descriptive notes. Forearm 38—43 mm . Noseleaf of the Dayak Leaf-nosed Bat is small and rounded, without supplementary leaflet. Intemarial septum is swollen and broader at base, and narrow in middle. Pelage is uniformly dark brown, with slightly paler tips. Habitat. Lowland rainforest to semi-evergreen forest. Food and Feeding. The Dayak Leaf-nosed Bat feeds on small insects in forest understory. Breeding. In peninsular Thailand , lactating females were captured in May-June. On following pages: 71. House-dwelling Leaf-nosed Bat ( Hipposideros einnaythu ); 72. Andersen's Leaf-nosed Bat (( Hipposideros halophyllus }; 75. Kolar Leaf-nosed Bat ( Hipposideros hypophyllus }; 76. Phou Khao Khouay Leaf-nosed nosed Bat ( Hipposideros macrobullatus }; 79. Maggie Taylor's Leaf-nosed Bat ( Hipposideros maggietaylorae ); 80. Malayan 82. Philippine Forest Leaf-nosed Bat ( Hipposideros obscurus ); 83. Orbiculus Leaf-nosed Bat ( Hipposideros orbiculus 86. Philippine Pygmy Leaf-nosed Bat ( Hipposideros pygmaeus }; 87. Ridley’ s Leaf-nosed Bat ( Hipposideros ridleyi }; 88 Hipposideros gentilis ); 73. Fulvus Leaf-nosed Bat ( Hipposideros fulvus }; 74. Thailand Leaf-nosed Bat Bat ( Hipposideros khaokhouayensis }; 77. Kunz's Leaf-nosed Bat ( Hipposideros kunzi }; 78. Big-eared Leaf-Leaf-nosed Bat ( Hipposideros nequam ); 81. Nicobar Leaf-nosed Bat ( Hipposideros nicobarulae };}; 84. Biak Leaf-nosed Bat ( Hipposideros papua }; 85. Pomona Leaf-nosed Bat ( Hipposideros pomona };. Laotian Leaf-nosed Bat ( Hipposideros rotalis }. Activity patterns. T he Dayak Leaf-nosed Bat roosts in limestone caves, under rocks, and in hollow trees. Call frequency of the F segment is 167-170 kHz. Movements, Home range and Social organization. The Dayak Leaf-nosed Bat was caught in the same trap with Kunz’s Leaf-nosed Bats { H. kunzî ), Bicolored Leaf-nosed Bats { H. bicolor ), Bornean Leaf-nosed Bats { H. doriae ), and other forest-dwelling Murino. and Kerivoula. Status and Conservation. Classified as Least Concern on TTze IUCNRed List. The major threat to the Dayak Leaf-nosed Bat is habitat loss. Bibliography. Corbet & Hill (1992), Francis & Kingston (2008), Kingston et al. (2006), Payne et al. (1985), Phillipps & Phillipps (2016), Simmons (2005).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Hipposideros dyacorum	Hipposideros		dyacorum	Thomas	1902	0	Ann. Mag. Nat. Hist.	ser. 7, 9: 271	Dayak Leaf-nosed Bat	None.	Malaysia, Sarawak, Baram, Mt. Mulu.	Borneo (including Sarawak, Malaysia), Peninsular Thailand.	Not listed.	Least Concern	 bicolor species group.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hipposideros dyacorum	23	Dayak Leaf-nosed Bat	Dayak Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Hipposideros	NA	dyacorum	O. Thomas	1902	0	Hipposideros_dyacorum	Thomas, O. (1902). A new Hipposideros from Borneo. The Annals and magazine of natural history; zoology, botany, and geology, ser. 9, 9, 271.	https://www.biodiversitylibrary.org/item/63677#page/293/mode/1up	BM 1894.9.29.16		"Mount Mulu, Baram District, Sarawak," Borneo, Malaysia.			dyacorum O. Thomas, 1902	NA	NA	Thailand|Malaysia|Indonesia|Brunei	Asia	Indomalaya	LC	0	0	0	Hipposideros_dyacorum	0	sciname match	Hipposideros_dyacorum	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10132	Hipposideros dyacorum	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Hipposideros	dyacorum	Thomas, 1902		20000000	Hipposideros dyacorum	Least Concern		2020	2018-09-10 00:00:00 UTC	3.1	English	This species is listed as Least Concern because, although it is recorded in small numbers, it has a relatively wide distribution. It is sensitive to forest loss but there is insufficient data to show it is declining at the rate required to qualify for listing in a threatened category.	This species has been recorded from lowland rainforests. It roosts in limestone caves, under rocks and in hollow trees (Payne et al. 1985). It is also found in virgin forest above 600m a.s.l (Tama Abu Range, 2017 unpub data).	Deforestation as a result of logging, expanding agriculture and plantations, and fires, as well as limestone extraction represent threats to this species.	This species is found in small numbers.	Unknown	This species occurs throughout Borneo and much of Peninsular Malaysia (Simmons 2005). For Borneo, in Sarawak, this type of specimen has been collected based on different districts in Sarawak which include Miri, Mulu National Park (Isham et al. 2012), Kapit, Ulu Baleh (unpub. data 2015), Niah National Park, Lambir Hill National Park (unpub. data 2014) and Tama Abu Range (unpub. data 2017), from Bintulu, Bukit Kana Tatau, Kampun Langgir, Engkilili and Similajau National Park, from Bau Limestone Area (unpub. data 2014) and Dered Krian National Park, Lubok Antu district in Batang Ai National Park and lastly in Kuching district which include Kubah National Park, Bako National Park (2015), Tanjung Datu National Park Gunung Gading National Park and Gunung Regu (Mathew et al. 2009 unpub. data) and Santubong National Park (unpub. data 2016). Meanwhile, in Sandakan Sabah this species is recorded from Imbak Canyon and Madai Forest Reserve, Lahad Datu (unpub. data, 2017).In Peninsular Malaysia, the existence of this species is recorded from Pulau Pinang, Perak, Terenganu and few caves from Kelantan.		Terrestrial	This species occurs in protected areas. Survey and research work to improve the understanding of ecology and population biology of these bats.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Hipposideros		dyacorum	Thomas	1902	0	Ann. Mag. Nat. Hist.	ser. 7, 9: 271	Dayak Leaf-nosed Bat	None.	Malaysia, Sarawak, Baram, Mt. Mulu.	Borneo (including Sarawak, Malaysia), Peninsular Thailand.	Not listed.	Least Concern	 bicolor species group.	Hipposideros dyacorum	1004596	23	Dayak Leaf-nosed Bat	Dayak Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Hipposideros	NA	dyacorum	O. Thomas	1902	0	Hipposideros_dyacorum	Thomas, O. (1902). A new Hipposideros from Borneo. The Annals and magazine of natural history; zoology, botany, and geology, ser. 9, 9, 271.	https://www.biodiversitylibrary.org/item/63677#page/293/mode/1up	BM 1894.9.29.16		"Mount Mulu, Baram District, Sarawak," Borneo, Malaysia.			dyacorum O. Thomas, 1902	NA	NA				Thailand|Malaysia|Indonesia|Brunei	Asia	Indomalaya	LC	0	0	0	Hipposideros_dyacorum	0	sciname match	Hipposideros_dyacorum	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hipposideros_dyacorum	1004596	23	Dayak Roundleaf Bat	Dayak Roundleaf Bat|Dayak Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Hipposideros	NA	dyacorum	O. Thomas	0	Hipposiderus dyacorum	Thomas, O. 1902-04-01. A new _Hipposiderus_ from Borneo. Annals and Magazine of Natural History (7)9(52):271-272.	https://www.biodiversitylibrary.org/page/19331556	BMNH:Mamm:1894.9.29.10	holotype	https://data.nhm.ac.uk/object/3993c117-ec7e-4525-bfea-3b8a191e3657	"Mount Mulu, Baram District, Sarawak," Borneo, Malaysia.			NA	NA				Thailand|Malaysia|Indonesia|Brunei	Asia	Indomalaya	LC	0	0	0	Hipposideros_dyacorum	0	sciname match	Hipposideros_dyacorum	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Hipposideros		dyacorum	Thomas	1902	0	Ann. Mag. Nat. Hist.	ser. 7, 9: 271	Dayak Leaf-nosed Bat	None.	Malaysia, Sarawak, Baram, Mt. Mulu.	Borneo (including Sarawak, Malaysia), Peninsular Thailand.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10132/22090760/' target='_blank'>Least Concern</a>	bicolorspecies group.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hipposideros dyacorum; Hipposideros dyacorum; Hipposideros dyacorum; Hipposideros dyacorum; Hipposideros dyacorum; Hipposideros dyacorum; dyacorum; Phyllorhine dayak; Dayak-Rundblattnase; Hiposidérido dayak; Dayak Roundleaf Bat; Dayak Leaf-nosed Bat; Dayak Roundleaf Bat; Dayak Leaf-nosed Bat; Dayak Leaf-nosed Bat; H. dyacorum