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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L388	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Hipposideros dinops	Hipposideros dinops	Hipposideros dinops	Hipposideros dinops	Hipposideros dinops	Hipposideros dinops	Hipposideros dinops	Hipposideros dinops	Hipposideros dinops	Hipposideros dinops	Hipposideros dinops	Hipposideros dinops	Hipposideros dinops	Hipposideros dinops	Hipposideros dinops		[MSW2] Includes pelingensis; see Hill (19636:113).; [MSW3] diadema species group. Does not include pelingensis; see discussion under that species. See Flannery (1995b) and Bonaccorso (1998).; [HMW] Hipposiderus [sic] dinops K Andersen, 1905 , “Rubiana [= Roviana Lagoon], a minute inlet very near western coast of New Georgia, Solomon Islands .” Hipposideros dinops is in the diadema species group. Many authors have considered this species to include pelingensis as a subspecies, but these taxa are separated by 1800 km and are diagnosably different. Here H. pelingensis is considered a good species, following N. B. Simmons in 2005. Monotypic.; [batnames2022]  diadema species group. Does not include pelingensis; see discussion under that species. See Flannery (1995b) and Bonaccorso (1998).; [IUCN] This species formerly considered conspecific with Hipposideros pelingensis from Peleng and Sulawesi Islands (Indonesia). However, the two are distinct with separate evolutionary origins (Simmons 2005, Lavery et al. 2014). The taxonomic relationships between H. dinops , H. diadema and H. demissus are complex and require further detailed study.; [batnames2023]  diadema species group. Does not include pelingensis; see discussion under that species. See Flannery (1995b) and Bonaccorso (1998).; [batnames2025_1.7] diademaspecies group. Does not include pelingensis; see discussion under that species. See Flannery (1995b) and Bonaccorso (1998).				pelingensis		pelingensis.	pelingensis, dinops							dinops	This species formerly considered conspecific with Hipposideros pelingensis from Peleng and Sulawesi Islands (Indonesia). However, the two are distinct with separate evolutionary origins (Simmons 2005, Lavery et al. 2014). The taxonomic relationships between H. dinops , H. diadema and H. demissus are complex and require further detailed study.			dinops	dinops			dinops Andersen, 1905		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Fierce leaf-nosed bat	Celebes, Peling I, Solomons	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Hipposideros dinops	Solomon Isis., New Georgia Group, Rubiana Isl.	K. Andersen	1905	Ann. Mag. Nat. Hist., ser. 7, 16:502.	Distribution: Known only from Celebes and the Solomons.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Fierce leaf-nosed bat	Sulawesi, Peling I, Solomons	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	K. Andersen	1905	Ann. Mag. Nat. Hist, ser. 7, 16:502.	Includes pelingensis; see Hill (19636:113).	Rubiana, Ysabel, Malaita and Bougainville Isis (Solomon Isis); Peleng Isl and Sulawesi (Indonesia).	Solomon Isis, New Georgia Group, Rubiana Isl.		ANDERSEN	1905	Frontal region concave with a shallow frontal depression. Interorbital region relatively narrow. Zygomatic arches markedly flared. Size large (forearm length, 93-97 mm; condylocanine length more than 32 but less than 33 mm; maxil lary toothrow length more than 14.0 mm). Inter mediate noseleaf with a prominent medial ridge.	Distribution: Known only from Celebes and the Solomons.	Two subspecies are recognized:	H. d. pelingensis (Celebes), H. d. dinops (Solomons).	66	species	H. dinops	ANDERSEN	1905	Hipposideros	genus	Hipposideros dinops				Frontal region concave with a shallow frontal depression. Interorbital region relatively narrow. Zygomatic arches markedly flared. Size large (forearm length, 93-97 mm; condylocanine length more than 32 but less than 33 mm; maxil lary toothrow length more than 14.0 mm). Inter mediate noseleaf with a prominent medial ridge.	Two subspecies are recognized:		51. H. dinops ANDERSEN 1905 [diadema group],	51	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Hipposideros dinops	Hipposideros		dinops	K. Andersen		1905		Ann. Mag. Nat. Hist., ser. 7	16		502		Fierce Leaf-nosed Bat	Solomon Isls, New Georgia Group, Rubiana Isl.	Solomon Isls; Bougainville Isl (Papua New Guinea).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).		diadema species group. Does not include pelingensis; see discussion under that species. See Flannery (1995b) and Bonaccorso (1998).	03BD87A2C678A20BF8A6F46DFA7F40E9	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	241	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C678A20BF8A6F46DFA7F40E9.xml	Hipposideros dinops	Hipposideridae	Hipposideros	dinops	K Andersen	1905	Phyllorhine féroce @fr | Salomonen-Rundblattnase @de | Hiposidérido de Andersen @es | Fierce Roundleaf Bat @en | Giant Horseshoe Bat @en	Hipposiderus [sic] dinops K Andersen, 1905 , “Rubiana [= Roviana Lagoon], a minute inlet very near western coast of New Georgia, Solomon Islands .” Hipposideros dinops is in the diadema species group. Many authors have considered this species to include pelingensis as a subspecies, but these taxa are separated by 1800 km and are diagnosably different. Here H. pelingensis is considered a good species, following N. B. Simmons in 2005. Monotypic.	Solomon Is (Bougainville, New Georgia , Santa Isabel, SanJorge, Malaita, and Guadalcanal).	Head-body 88-105 mm, tail 54-63 mm, ear 31-35 mm, forearm 86*1-98* 9 mm ; weight 78 g . Ears of the Fierce Leaf-nosed Bat are large, triangular, and slightly concave below tips. Noseleaf is well developed and has three supplementary lateral leaflets, of which the third is the smallest. Anterior leaf is large and has no central emargination. Intermediate leaf is expanded and presents a protuberance in middle. Upper border of posterior leafis semicircular with a small projection, and has medial septum and two lateral septa forming four cells. Males present a frontal sac. Fur is whitish or white with light brown spots on dorsum, while ventral fur is yellowishbrownish.	The Fierce Leaf-nosed Bat has been observed over forest habitats and tropical moist lowland. It has been reported from sea level up to 400 m .	The Fierce Leaf-nosed Bat is probably insectivorous.	No information.	The Fierce Leaf-nosed Bat occupies caves. Its CF echolocation call frequency ranges at 52-55 kHz.	The Fierce Leaf-nosed Bat has been observed roosting both alone and in small groups of up to twelve individuals.	Classified as Data Deficient on The IUCNRed List, due to the lack of information on its population status. The Fierce Leaf-nosed Bat occurs at low densities and it has only been reported from a few localities. The overall population is thought to be decreasing. Further research on ecology and threats is needed in order to establish adequate conservation measures.	Bonaccorso (1998) | Hamilton (2008c) | Hill (1963a) | Simmons (2005)	https://zenodo.org/record/3749117/files/figure.png	41. Fierce Leaf-nosed Bat Hipposideros dinops French: Phyllorhine féroce / German: Salomonen-Rundblattnase / Spanish: Hiposidérido de Andersen Other common names: Fierce Roundleaf Bat , Giant Horseshoe Bat Taxonomy. Hipposiderus [sic] dinops K Andersen, 1905 , “Rubiana [= Roviana Lagoon], a minute inlet very near western coast of New Georgia, Solomon Islands .” Hipposideros dinops is in the diadema species group. Many authors have considered this species to include pelingensis as a subspecies, but these taxa are separated by 1800 km and are diagnosably different. Here H. pelingensis is considered a good species, following N. B. Simmons in 2005. Monotypic. Distribution. Solomon Is (Bougainville, New Georgia , Santa Isabel, SanJorge, Malaita, and Guadalcanal). Descriptive notes. Head-body 88-105 mm, tail 54-63 mm, ear 31-35 mm, forearm 86*1-98* 9 mm ; weight 78 g . Ears of the Fierce Leaf-nosed Bat are large, triangular, and slightly concave below tips. Noseleaf is well developed and has three supplementary lateral leaflets, of which the third is the smallest. Anterior leaf is large and has no central emargination. Intermediate leaf is expanded and presents a protuberance in middle. Upper border of posterior leafis semicircular with a small projection, and has medial septum and two lateral septa forming four cells. Males present a frontal sac. Fur is whitish or white with light brown spots on dorsum, while ventral fur is yellowishbrownish. Habitat. The Fierce Leaf-nosed Bat has been observed over forest habitats and tropical moist lowland. It has been reported from sea level up to 400 m . Food and Feeding. The Fierce Leaf-nosed Bat is probably insectivorous. Breeding. No information. Activity patterns. The Fierce Leaf-nosed Bat occupies caves. Its CF echolocation call frequency ranges at 52-55 kHz. Movements, Home range and Social organization. The Fierce Leaf-nosed Bat has been observed roosting both alone and in small groups of up to twelve individuals. Status and Conservation. Classified as Data Deficient on The IUCNRed List, due to the lack of information on its population status. The Fierce Leaf-nosed Bat occurs at low densities and it has only been reported from a few localities. The overall population is thought to be decreasing. Further research on ecology and threats is needed in order to establish adequate conservation measures. Bibliography. Bonaccorso (1998), Hamilton (2008c), Hill (1963a), Simmons (2005).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Hipposideros dinops	Hipposideros		dinops	K. Andersen	1905	0	Ann. Mag. Nat. Hist.	ser. 7, 16: 502	Fierce Leaf-nosed Bat	None.	Solomon Isls, New Georgia Group, Rubiana Isl.	Solomon Isls; Bougainville Isl (Papua New Guinea).	Not listed.	Vulnerable	 diadema species group. Does not include pelingensis; see discussion under that species. See Flannery (1995b) and Bonaccorso (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hipposideros dinops	23	Fierce Leaf-nosed Bat	Fierce Roundleaf Bat|Giant Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Hipposideros	NA	dinops	K. Andersen	1905	0	Hipposideros_dinops	Andersen, K. (1905). On Hipposideros diadema and its closest allies. The Annals and magazine of natural history; zoology, botany, and geology, ser. 7, 16, 502.	https://www.biodiversitylibrary.org/item/63348#page/542/mode/1up	BM 1888.1.5.22		"Rubiana [= Roviana Lagoon], a minute inlet very near western coast of New Georgia, Solomon Islands."			dinops K. Andersen, 1905	NA	NA	Papua New Guinea|Solomon Islands	Oceania	Australasia/Oceania	DD	0	0	0	Hipposideros_dinops	0	sciname match	Hipposideros_dinops	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10129	Hipposideros dinops	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Hipposideros	dinops	K. Andersen, 1905	This species formerly considered conspecific with Hipposideros pelingensis from Peleng and Sulawesi Islands (Indonesia). However, the two are distinct with separate evolutionary origins (Simmons 2005, Lavery et al. 2014). The taxonomic relationships between H. dinops , H. diadema and H. demissus are complex and require further detailed study.	20000000	Hipposideros dinops	Vulnerable	B2ab(ii,iii,v); C2a(i)	2021	2021-07-30 00:00:00 UTC	3.1	English	The Fierce Leaf-nosed Bat is restricted to six island groups in the Solomon Island Archipelago; Bougainville (Papua New Guinea), Choiseul, New Georgia, Santa Isabel, Guadalcanal and Malaita (all in the Solomon Islands). There is continuing decline in the area, extent, and quality of habitat due to expanding subsistence and commercial agriculture, widespread logging, disturbance at caves and negative impacts of invasive species, such as Little Fire Ant (Wasmannia auropunctata ). It is listed as Vulnerable under criterion B2 because the species is dependent on a very small number of caves on each major island group, resulting in an area of occupancy of ca 240-400 kmÂ² over six disconnected islands. It is also listed under criterion C2 because the number of mature individuals is estimated at fewer than 10,000, the population is inferred to be continually declining and the number of mature individuals in each of the six subpopulations is estimated to be fewer than 1,000.	The Fierce Leaf-nosed Bat is a cave roosting species and is restricted to a small number of suitable caves and surrounding areas of lowland rainforest. Animals have been found roosting singly or in small groups of up to 12 individuals, often sharing caves with several other bat species, especially other species of Hipposideros (Flannery 1995, Bonaccorso 1998). It is the most robust leaf-nosed bat in Bougainville and the Solomon Islands, weighing 70-80 grams. It is believed to hunt from perches on vegetation, flying out to catch invertebrate prey (Lavery et al. 2016). Its robust dentition and relatively low frequency (52â€“55 kHz) echolocation call suggest it hunts for larger prey. Genetic sequencing has revealed genetic divergence between sympatric Hipposideros dinops and H. diadema is less than that between H. diadema samples from the Solomon Islands and mainland Papua New Guinea (Lavery 2014). However, where H. dinops and H. diadema co-occur on Guadalcanal, Santa Isabel and in the Western Province they are reproductively isolated. Hipposideros dinops is easily distinguished by its larger size and distinctive long duration, relatively low frequency, echolocation call (Lavery 2015, Pennay and Lavery 2017). It is believed that H. dinops has undergone recent, rapid speciation, diverging from H. diadema in the past 1â€“2 million years (Lavery et al. 2014).	This species is likely threatened by extensive logging, particularly in the Solomon Islands, where it is estimated almost all accessible (lowland) forests have been logged (Katovai et al. 2015). Mining may cause localised loss of habitat. Cave disturbance is also a threat, many locations are visited by hunters, and in Guadalcanal major disturbances (excavation, and fires) have been observed in caves used by this species caused by prospectors searching for rumoured treasures buried by Japanese forces in WWII. The species is not usually hunted for food, but given its large body size, it is taken opportunistically when hunting for other bats (e.g., Dobsonia inermis ) inside caves. Invasive species, particularly Little Fire Ant (Wasmannia auropunctata ), which has become widespread on some islands and is likely to be affecting both food resources and causing direct mortality for the species. It is also likely to be predated by introduced cats and rats.	The Fierce Leaf-nosed Bat is uncommon; it occupies a very limited number of caves and is absent from many apparently suitable caves (Flannery 1995, Lavery pers comm.). Where it is found, it occurs at low densities with small numbers, usually &lt;12 (Flannery 1995, Bonaccorso 1998). This bat's echolocation call has only recently been described (Pennay and Lavery 2017) and acoustic surveys may reveal it is easily detected where it occurs. Pennay and Lavery (2017) reported that its call was recorded with regularity amongst free flying bats in acoustic recordings of free flying bats on Guadalcanal and Choiseul Islands (Solomon Islands), however these recordings were undertaken in close proximity to caves known to be used by the species. Only a single Hipposideros dinops call was recorded at 223 m asl, at Kunua, during acoustic bat surveys on the west coast of Bougainville (J. Novera pers. comm.). The number of mature individuals over its range is unknown, however it is estimated to be fewer than 6,000. The population is divided into six subpopulations: Bougainville, Choiseul, Isabel, Malaita, Guadalcanal and the New Georgia (group) representing a distinct subpopulation with sea crossings between these major islands extremely rare. The number of mature individuals in each subpopulation is estimated to be fewer than 1,000 mature individuals. The taxonomic relationships between different island sub-populations are also complex, and each should probably be regarded as evolutionary significant units for the genus Hipposideros  (T. Lavery pers. comm.). The population is inferred to be declining based on observed patterns of decline in other island endemic bat species with small geographic ranges (Lavery et al. 2020) and the extensive habitat loss and degradation across the main islands in its range. More than 40% of Bougainvilleâ€™s lowland forest has been lost or degraded over the last half century (Woxvold and Novera 2021) and unsustainable logging in the Solomon Islands has occurred in almost all accessible lowland rainforest (Katovai et al. 2015). The rate of decline is likely to continue and intensify as logging rates increase on all islands.	Decreasing	The Fierce Leaf-nosed Bat is present on the islands of Bougainville (Papua New Guinea), Choiseul, New Georgia, Nggatokae Island, Santa Isabel, San Jorge, and Malaita (all in the Solomon Islands). On these islands it appears to be restricted to a few caves and surrounding lowland rainforest habitats from sea level to about 400 m asl (Flannery 1995, Bonaccorso 1998). Only 1 or 2 caves are known to be used by this species in each of the major island groups, although it is likely that unknown caves also exist. The species is dependent on a very small number of caves on each major island group, resulting in an area of occupancy of ca 240â€“400 kmÂ² over six disconnected islands.	The species is not known to be used or traded. It is not usually hunted for food, but given its large body size, it is taken opportunistically when hunting for other bats (e.g., Dobsonia inermis ) inside caves.	Terrestrial	It is not known if this species occurs in any conservation reserves. Further studies into the distribution, ecology, population numbers, and threats to this species are needed along with protection of the known roosting sites and surrounding forests. The taxonomic relationships between the different island sub-populations requires further investigation to clarify their status as evolutionary significant units for the genus Hipposideros . Conservation actions working with customary landowners to protect known roost caves from disturbance would be beneficial both to this, and other species with which it shares the caves.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Hipposideros		dinops	K. Andersen	1905	0	Ann. Mag. Nat. Hist.	ser. 7, 16: 502	Fierce Leaf-nosed Bat	None.	Solomon Isls, New Georgia Group, Rubiana Isl.	Solomon Isls; Bougainville Isl (Papua New Guinea).	Not listed.	Vulnerable	 diadema species group. Does not include pelingensis; see discussion under that species. See Flannery (1995b) and Bonaccorso (1998).	Hipposideros dinops	1004593	23	Fierce Leaf-nosed Bat	Fierce Roundleaf Bat|Giant Horseshoe Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Hipposideros	NA	dinops	K. Andersen	1905	0	Hipposideros_dinops	Andersen, K. (1905). On Hipposideros diadema and its closest allies. The Annals and magazine of natural history; zoology, botany, and geology, ser. 7, 16, 502.	https://www.biodiversitylibrary.org/item/63348#page/542/mode/1up	BM 1888.1.5.22		"Rubiana [= Roviana Lagoon], a minute inlet very near western coast of New Georgia, Solomon Islands."			dinops K. Andersen, 1905	NA	NA				Papua New Guinea|Solomon Islands	Oceania	Australasia/Oceania	DD	0	0	0	Hipposideros_dinops	0	sciname match	Hipposideros_dinops	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hipposideros_dinops	1004593	23	Fierce Roundleaf Bat	Fierce Roundleaf Bat|Giant Horseshoe Bat|Fierce Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Hipposideros	NA	dinops	Andersen	0	Hipposiderus dinops	Andersen, K.C. 1905-11-01. On _Hipposiderus diadema_ and its closest allies. Annals and Magazine of Natural History (7)16(95):497-507.	https://www.biodiversitylibrary.org/page/19215149	BMNH:Mamm:1888.1.5.22	holotype	https://data.nhm.ac.uk/object/a59ce3eb-123b-43d2-8e35-b16e320b5214	"Rubiana [= Roviana Lagoon], a minute inlet very near western coast of New Georgia, Solomon Islands."			NA	NA				Papua New Guinea|Solomon Islands	Oceania (Continent)	Australasia	VU	0	0	0	Hipposideros_dinops	0	sciname match	Hipposideros_dinops	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Hipposideros		dinops	K. Andersen	1905	0	Ann. Mag. Nat. Hist.	ser. 7, 16: 502	Fierce Leaf-nosed Bat	None.	Solomon Isls, New Georgia Group, Rubiana Isl.	Solomon Isls; Bougainville Isl (Papua New Guinea).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10129/22091011/' target='_blank'>Vulnerable</a>	diademaspecies group. Does not include pelingensis; see discussion under that species. See Flannery (1995b) and Bonaccorso (1998).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hipposideros dinops; Hipposideros dinops; Hipposideros dinops; Hipposideros dinops; Hipposideros dinops; Hipposideros dinops; dinops; Phyllorhine féroce; Salomonen-Rundblattnase; Hiposidérido de Andersen; Fierce Roundleaf Bat; Giant Horseshoe Bat; Fierce Leaf-nosed Bat; Fierce Roundleaf Bat; Giant Horseshoe Bat; Fierce Leaf-nosed Bat; Fierce Leaf-nosed Bat; H. dinops
