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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L385	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Hipposideros curtus	Hipposideros curtus	Hipposideros curtus	Hipposideros curtus	Hipposideros curtus	Hipposideros curtus	Hipposideros curtus	Hipposideros curtus	Hipposideros curtus	Hipposideros curtus	Hipposideros curtus	Hipposideros curtus	Hipposideros curtus	Hipposideros curtus	Hipposideros curtus		[MSW2] Includes sandersoni; see Hill (19636:60).; [MSW3] bicolor species group. Includes sandersoni; see Hill (1963b).; [HMW] Hipposideros curtus G. M. Allen, 1921 , Sakbayeme, Cameroon . Hipposideros curtus was formerly included in the bicolor species group (4 species). Monotypic.; [batnames2022]  bicolor species group. Includes sandersoni; see Hill (1963b).; [IUCN] The taxonomic relationship between Hipposideros curtus and H. vittatus is unclear and needs further investigation.; [batnames2023]  bicolor species group. Includes sandersoni; see Hill (1963b).; [batnames2025_1.7] bicolorspecies group. Includes sandersoni; see Hill (1963b).				sandersoni		sandersoni.			sandersoni			curtus	curtus - sandersoni	curtus, sandersoni	The taxonomic relationship between Hipposideros curtus and H. vittatus is unclear and needs further investigation.	curtus	curtus - sandersoni	curtus, sandersoni 	curtus, sandersoni 	curtus	curtus  - sandersoni	curtus G. M. Allen, 1921|sandersoni Sanderson, 1937		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Short-tailed leaf-nosed bat	Nigeria, Cameroun, Fernando Poo	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Hipposideros curtus	Cameroun, Sakbayeme.	G. M. Allen	1921	Rev. Zool. Afr., 9:194.	Distribution: Confined to southern Cameroon and Fernando Poo in tropical western Africa.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Short-tailed leaf-nosed bat	Cameroun, Nigeria, Bioco	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	G. M. Allen	1921	Rev. Zool. Afr., 9:194.	Includes sandersoni; see Hill (19636:60).	Cameroon, Bioko, perhaps Nigeria.	Cameroon, Sakbayeme.		G. M. ALLEN	1921	Two lateral leaflets present, neither one extend ing anteriorly beneath anterior noseleaf to median line. Posterior noseleaf without a well-developed transverse serrated supplementary structure on its posterior face, but supported by three septa. In ternarial septum expanded to form a small disc like structure. Pterygoids short, sphenoidal bridge narrow, not concealing lateral apertures. Size me dium (forearm length, 42-47 mm).	Distribution: Confined to southern Cameroon and Fernando Poo in tropical western Africa.	No subspecies.		62	species	H. curtus	G. M. ALLEN	1921	Hipposideros	genus	Hipposideros curtus				Two lateral leaflets present, neither one extend ing anteriorly beneath anterior noseleaf to median line. Posterior noseleaf without a well-developed transverse serrated supplementary structure on its posterior face, but supported by three septa. In ternarial septum expanded to form a small disc like structure. Pterygoids short, sphenoidal bridge narrow, not concealing lateral apertures. Size me dium (forearm length, 42-47 mm).	No subspecies.		25. H. curtus G. M. ALLEN 1921 [bicolor group],	25	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Hipposideros curtus	Hipposideros		curtus	G. M. Allen		1921		Rev. Zool. Afr.	9		194		Short-tailed Leaf-nosed Bat	Cameroon, Sakbayeme.	Cameroon, Bioko (Equatorial Guinea).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).	sandersoni Sanderson, 1937.	bicolor species group. Includes sandersoni; see Hill (1963b).	03BD87A2C664A216FF55FEE8FA6E443F	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	245	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C664A216FF55FEE8FA6E443F.xml	Hipposideros curtus	Hipposideridae	Hipposideros	curtus	G. M. Allen	1921	Phyllorhine à queue courte @fr | Kurzschwanz-Rundblattnase @de | Hiposidérido de cola corta @es | Short-tailed Roundleaf Bat @en	Hipposideros curtus G. M. Allen, 1921 , Sakbayeme, Cameroon . Hipposideros curtus was formerly included in the bicolor species group (4 species). Monotypic.	Patchily in Cameroon and Equatorial Guinea , including Bioko I.	Head—body 50-55 mm, tail 18-23 mm, ear 15-22 mm, hindfoot 7—8 mm , forearm 42—47 mm . Muzzle of the Short-tailed Leaf-nosed Bat is relatively short with large but simple noseleaf that has two lateral leaflets. A frontal sac is usually present in both sexes posterior to noseleaf. Intemarial septum is swollen and partially covering the nares. Ears are separate, and relatively large. Fur is fine and short, sepiabrown dorsally, and slightly lighter ventrally.	Restricted to lowland (0-500 m) forests.	No information.	No information.	The Short-tailed Leaf-nosed Bat has been observed roosting in caves and under boulders.	Short-tailed Leaf-nosed Bats roost singly or in small groups.	Classified as Vulnerable on The IUCN Red List, because of habitat loss due to logging, and perhaps disturbance at roost sites.	Happold, M. (2013x) | Hill (1963a) | Rosevear (1965) | Sanderson (1940)	https://zenodo.org/record/3749129/files/figure.png	47. Short-tailed Leaf-nosed Bat Hipposideros curtus French: Phyllorhine à queue courte / German: Kurzschwanz-Rundblattnase I Spanish: Hiposidérido de cola corta Other common names: Short-tailed Roundleaf Bat Taxonomy. Hipposideros curtus G. M. Allen, 1921 , Sakbayeme, Cameroon . Hipposideros curtus was formerly included in the bicolor species group (4 species). Monotypic. Distribution. Patchily in Cameroon and Equatorial Guinea , including Bioko I. Descriptive notes. Head—body 50-55 mm, tail 18-23 mm, ear 15-22 mm, hindfoot 7—8 mm , forearm 42—47 mm . Muzzle of the Short-tailed Leaf-nosed Bat is relatively short with large but simple noseleaf that has two lateral leaflets. A frontal sac is usually present in both sexes posterior to noseleaf. Intemarial septum is swollen and partially covering the nares. Ears are separate, and relatively large. Fur is fine and short, sepiabrown dorsally, and slightly lighter ventrally. Habitat. Restricted to lowland (0-500 m) forests. Food and Feeding. No information. Breeding. No information. Activity patterns. The Short-tailed Leaf-nosed Bat has been observed roosting in caves and under boulders. Movements, Home range and Social organization. Short-tailed Leaf-nosed Bats roost singly or in small groups. Status and Conservation. Classified as Vulnerable on The IUCN Red List, because of habitat loss due to logging, and perhaps disturbance at roost sites. Bibliography. Happold, M. (2013x), Hill (1963a), Rosevear (1965), Sanderson (1940).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Hipposideros curtus	Hipposideros		curtus	G. M. Allen	1921	0	Rev. Zool. Afr.	0.5097	Short-tailed Leaf-nosed Bat	 sandersoni Sanderson, 1937.	Cameroon, Sakbayeme.	Cameroon, Bioko (Equatorial Guinea).	Not listed.	Endangered	 bicolor species group. Includes sandersoni; see Hill (1963b).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hipposideros curtus	23	Short-tailed Leaf-nosed Bat	Short-tailed Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Hipposideros	NA	curtus	G. M. Allen	1921	0	Hipposideros_curtus	Allen, G. M. (1921). A new horseshoe bat from West Africa. Revue zoologique africaine, 9, 194.	https://www.biodiversitylibrary.org/item/42138#page/216/mode/1up	MCZ 19305		Sakbayeme, Cameroon.			curtus G. M. Allen, 1921|sandersoni Sanderson, 1937	NA	NA	Nigeria|Cameroon|Equatorial Guinea	Africa	Afrotropic	EN	0	0	0	Hipposideros_curtus	0	sciname match	Hipposideros_curtus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10125	Hipposideros curtus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Hipposideros	curtus	G.M. Allen, 1921	The taxonomic relationship between Hipposideros curtus and H. vittatus is unclear and needs further investigation.	20000000	Hipposideros curtus	Endangered	A2c	2020	2019-05-16 00:00:00 UTC	3.1	English	This species is listed as Endangered under criterion A2(c) based on a suspected past population size reduction of more than 50% over the past 15 years (three generations; generation length = 5 years; Pacifici et al. 2013). The population decline is continuing due to loss of some previously known roosts, confirmed occupancy of only one out of 35 caves assessed in Nigeria, the limited availability of undisturbed roosts, the species being limited by quality forest, and ongoing deforestation within its range significantly reducing the area of occupancy.	This species is heavily dependent on undisturbed caves and lowland rainforest. It forages in the forest understory and is rarely caught outside forests (Perret and Allen 1956). Only a single record is known from human residence – government rest house at Mamfe (Sanderson 1940), which may have been well-lit and thus likely to have attracted insects, that in turn potentially attracted the bat. It has not been collected in cleared forest or farmland. A photo of recent captures suspected to be Hipposideros curtus captured by Cameroonian colleagues in farmlands have been re-identified as H. caffer or cf. ruber (I. Tanshi, personal observation). It has recently been captured in Cross River National Park, Nigeria (I. Tanshi, unpublished data), which has added to its known range states. Here, it is known from low to mid elevations (800 m asl), and has been captured along a river. It appears to have a disjointed distribution within this forest, where small numbers (1-8 individuals) are often captured at one particular location but none in nearby areas. Furthermore, within a sampling area of ca 1 km², the species was only encountered in very close proximity to cave-like boulders, demonstrating its dependence on appropriate roosting sites and possibly small home ranges. Also, since several individuals were frequently captured at the same site, it suggests that this species may forage in groups. The species appears to show high roost fidelity, the single known roost in CRNP has been checked annually over the last three years during which time it has been used consistently. Photographs taken in a cave at Ndian in the (iNaturalist.org) suggests that the species may also roost in Korup National Park. However, no captures have been made there, nor have there been additional visits to confirm long term use of this Ndian roost.	This species is threatened by the continued degradation and clear-cutting of its forest habitat. Although protected, forests where the species is known in Nigeria are threatened by wildfire outbreaks arising from slash and burn agriculture in surrounding farm lands (I. Tanshi, personal observation). Similarly, the species is heavily dependent on undisturbed caves and has been known to desert previously occupied caves due to disturbance or cave destruction. Some of the few previously known roosts have been lost (Happold 2013). Furthermore, while the species is known to roost with Rousettus aegyptiacus in some countries or regions (e.g. Cameroon and Bioko Island), in Nigeria, it was not found in roosts where R. aegyptiacus is hunted intensively. A search of 35 cave-like boulder roosts in Afi Mountain Wildlife Sanctuary and Cross River National Park (both in Nigeria), yielded just one roosting group of H. curtus , the lack of additional groups is thought to be due to the nature and intensity of fruit bat hunting that currently goes on at these parks.	This is a rarely encountered species that had not been captured or observed in the wild since the 1970s until recent surveys conducted in Nigeria (I. Tanshi, unpublished data). It appears to occur in small and fragmented populations with only a few (ca 10-15) individuals sharing a roost. Its population is considered to be decreasing due to deforestation, habitat degradation and cave roost disturbance and destruction (I. Tanshi, personal observation).	Decreasing	The species is endemic to Cameroon, Nigeria and Equatorial Guinea (Rio Muni and Bioko Island). In Nigeria, it is not found above 800 m asl. Previously suspected to occur in Nigeria, its presence in the country has recently been confirmed, based on captures and a single roost from two adjacent protected areas: Afi Mountain Wildlife Sanctuary and Cross River National Park (I. Tanshi, unpublished data).		Terrestrial	 Research needed Population size, distribution and trends: Intensive understory forest surveys and cave searches across previously known and other localities (for example those identified in distribution niche models) are essential for identification of additional populations and roosts. More surveys, as well as additional ecological and biogeographical studies, are needed to identify potential areas for conservation of this species.  Conservation Actions 1.) Protection of the single known roost of the species in Cross River National Park is now urgent. Cave roost protection depends on preventing all forms of disturbance of this last known roost including ending fruit bat hunting in Afi Mountain Wildlife Sanctuary and Cross River National Park. 2.) Elimination of wildfire outbreaks in both AMWS and CRNP is crucial for maintaining good foraging grounds for the species. 3.) Investigation of the human social dimension to cave roost disturbance and destruction. 4.) Extensive surveys and roost searches across the species range is essential for expanding the reach of conservation efforts and understanding its conservation ecology. 5.) Confirm the status of other previously known cave roosts.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Hipposideros		curtus	G. M. Allen	1921	0	Rev. Zool. Afr.	0.509722	Short-tailed Leaf-nosed Bat	 sandersoni Sanderson, 1937.	Cameroon, Sakbayeme.	Cameroon, Bioko (Equatorial Guinea).	Not listed.	Endangered	 bicolor species group. Includes sandersoni; see Hill (1963b).	Hipposideros curtus	1004590	23	Short-tailed Leaf-nosed Bat	Short-tailed Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Hipposideros	NA	curtus	G. M. Allen	1921	0	Hipposideros_curtus	Allen, G. M. (1921). A new horseshoe bat from West Africa. Revue zoologique africaine, 9, 194.	https://www.biodiversitylibrary.org/item/42138#page/216/mode/1up	MCZ 19305		Sakbayeme, Cameroon.			curtus G. M. Allen, 1921|sandersoni Sanderson, 1937	NA	NA				Nigeria|Cameroon|Equatorial Guinea	Africa	Afrotropic	EN	0	0	0	Hipposideros_curtus	0	sciname match	Hipposideros_curtus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hipposideros_curtus	1004590	23	Short-tailed Roundleaf Bat	Short-tailed Roundleaf Bat|Short-tailed Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Hipposideros	NA	curtus	G. M. Allen	0	Hipposideros curtus	Allen, G.M. 1921. A new horseshoe bat from West Africa. Revue zoologique africaine 9(2):193-196.	https://www.biodiversitylibrary.org/page/11299924	MCZ:Mamm:19305	holotype	https://mczbase.mcz.harvard.edu/guid/MCZ:Mamm:19305	Sakbayeme, Cameroon.	4.1	10.5	NA	NA				Nigeria|Cameroon|Equatorial Guinea	Africa	Afrotropic	EN	0	0	0	Hipposideros_curtus	0	sciname match	Hipposideros_curtus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Hipposideros		curtus	G. M. Allen	1921	0	Rev. Zool. Afr.	0.509722	Short-tailed Leaf-nosed Bat	sandersoni Sanderson, 1937.	Cameroon, Sakbayeme.	Cameroon, Bioko (Equatorial Guinea).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10125/22096364/' target='_blank'>Endangered</a>	bicolorspecies group. Includes sandersoni; see Hill (1963b).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hipposideros curtus; Hipposideros curtus; Hipposideros curtus; Hipposideros curtus; Hipposideros curtus; Hipposideros curtus; sandersoni; sandersoni; curtus; sandersoni; Phyllorhine à queue courte; Kurzschwanz-Rundblattnase; Hiposidérido de cola corta; Short-tailed Roundleaf Bat; Short-tailed Leaf-nosed Bat; Short-tailed Roundleaf Bat; Short-tailed Leaf-nosed Bat; Short-tailed Leaf-nosed Bat; H. curtus