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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L383	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Hipposideros coxi	Hipposideros coxi	Hipposideros coxi	Hipposideros coxi	Hipposideros coxi	Hipposideros coxi	Hipposideros coxi	Hipposideros coxi	Hipposideros coxi	Hipposideros coxi	Hipposideros coxi	Hipposideros coxi	Hipposideros coxi	Hipposideros coxi	Hipposideros coxi		[MSW3] bicolor species group.; [HMW] Hipposiderus [sic] coxi Shelford, 1901 , “Mount Penrisen, Sarawak , Borneo [ Malaysia ], 4200 feet [= 1280 m ].” Hipposideros coxi was formerly included in the bicolor species group, but is now placed in the calcaratus species group. Monotypic.; [batnames2022]  bicolor species group.; [batnames2023]  bicolor species group.; [batnames2025_1.7] bicolorspecies group.														coxi S				coxi 	coxi 			coxi Shelford, 1901		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Cox's leaf-nosed bat	Sarawak, Borneo	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Hipposideros coxi	Malaysia, Sarawak, Mt. Penrisen, 4200 ft. (1280 m).	Shelford	1901	Ann. Mag. Nat. Hist., ser. 7, 8:113.	Distribution: Confined to Borneo.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Cox's leaf-nosed bat	Sarawak, Borneo	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Shelford	1901	Ann. Mag. Nat. Hist, ser. 7, 8:113.		Sarawak (Borneo, Malaysian part).	Malaysia, Sarawak, Mt. Penrisen, 4,200 ft. (1,280 m).		SHELFORD	1901	Two lateral leaflets present, the anterior one extending anteriorly beneath anterior noseleaf to the median line. Intermediate noseleaf with a prominent me dian eminence. Posterior noseleaf supported by a narrow median septum and two broad lateral septa, deeply pocketed. Ear pointed. Size fairly large (forearm length, 51-55 mm).	Distribution: Confined to Borneo.	No subspecies.		64	species	H. coxi	SHELFORD	1901	Hipposideros	genus	Hipposideros coxi				Two lateral leaflets present, the anterior one extending anteriorly beneath anterior noseleaf to the median line. Intermediate noseleaf with a prominent me dian eminence. Posterior noseleaf supported by a narrow median septum and two broad lateral septa, deeply pocketed. Ear pointed. Size fairly large (forearm length, 51-55 mm).	No subspecies.		31. H. coxi SHELFORD 1901 [bicolor group],	31	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Hipposideros coxi	Hipposideros		coxi	Shelford		1901		Ann. Mag. Nat. Hist., ser. 7	8		113		Cox's Leaf-nosed Bat	Malaysia, Borneo, Sarawak, Mt. Penrisen, 4,200 ft. (1,280 m).	Sarawak (Borneo, Malaysian part).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.		bicolor species group.	03BD87A2C665A217F87DF82FF6FE5327	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	246	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C665A217F87DF82FF6FE5327.xml	Hipposideros coxi	Hipposideridae	Hipposideros	coxi	Shelford	1901	Phyllorhine de Cox @fr | Cox-Rundblattnase @de | Hiposidérido de Cox @es | Cox's Roundleaf Bat @en	Hipposiderus [sic] coxi Shelford, 1901 , “Mount Penrisen, Sarawak , Borneo [ Malaysia ], 4200 feet [= 1280 m ].” Hipposideros coxi was formerly included in the bicolor species group, but is now placed in the calcaratus species group. Monotypic.	Known only from a few localities in W Borneo (N & SW Sarawak ), including Gunung Mulu, Bako, and Tanjung Datu national parks, Mt Penrissen, and Bau Limestone area; it has been reported in Central Kalimantan based on acoustic surveys, but this requires confirmation.	Forearm 53—55 mm . Ears of Cox’s Leaf-nosed Bat are covered with hairs. Noseleaf is enlarged with two lateral leaflets. Anterior leaf fully covers muzzle. Intermediate leaf is bulbous and thickened. Posterior is large with swollen thick bump in middle. Wing membrane is dark. Pelage color is dark brown with ventral side paler.	Cox’s Leaf-nosed Bat forages in forest gaps understory, e.g. trails. It has been netted in mangrove forest near rock crevices. It occurs in lowlands, from sea level up to a hill cave at 1200 m .	Cox’s Leaf-nosed Bat feeds on small beedes, moths, and other insects over shrubs, creeks and water bodies by using aerial and gleaning tactics. It can fly slowly and hover during gleaning.	No information.	Cox’s Leaf-nosed Bat is a cave-dwelling species. Echolocation is a typical CF/FM type, with a call frequency of the F segment of46-52 kHz.	Cox’s Leaf-nosed Bat roosts in caves, typically in small groups of 2-5 individuals. At Wind Cave, Sarawak , it was recorded roosting on the cave ceiling in a group of less than 20 individuals near an opening.	Classified as Endangered on The IUCNRed List. Cox’s Leafnosed Bat is endemic to Borneo and is rare; its population is very small and decreasing (area of occupancy is less than 500 km 2). Besides habitat loss, the main threats include mining and bird-nest harvesting. Recorded in Bako, Tanjung Datu, and Gunung Mulu national parks and Wind and Fairy cave reserves.	Corbet & Hill (1992) | Khan et al. (2008) | MacArthur (2016a) | Phillipps & Phillipps (2016) | Rosli et al. (2018) | Simmons (2005)	https://zenodo.org/record/3749141/files/figure.png	53. Cox’s Leaf-nosed Bat Hipposideros coxi French: Phyllorhine de Cox / German: Cox-Rundblattnase / Spanish: Hiposidérido de Cox Other common names: Cox's Roundleaf Bat Taxonomy. Hipposiderus [sic] coxi Shelford, 1901 , “Mount Penrisen, Sarawak , Borneo [ Malaysia ], 4200 feet [= 1280 m ].” Hipposideros coxi was formerly included in the bicolor species group, but is now placed in the calcaratus species group. Monotypic. Distribution. Known only from a few localities in W Borneo (N & SW Sarawak ), including Gunung Mulu, Bako, and Tanjung Datu national parks, Mt Penrissen, and Bau Limestone area; it has been reported in Central Kalimantan based on acoustic surveys, but this requires confirmation. Descriptive notes. Forearm 53—55 mm . Ears of Cox’s Leaf-nosed Bat are covered with hairs. Noseleaf is enlarged with two lateral leaflets. Anterior leaf fully covers muzzle. Intermediate leaf is bulbous and thickened. Posterior is large with swollen thick bump in middle. Wing membrane is dark. Pelage color is dark brown with ventral side paler. Habitat. Cox’s Leaf-nosed Bat forages in forest gaps understory, e.g. trails. It has been netted in mangrove forest near rock crevices. It occurs in lowlands, from sea level up to a hill cave at 1200 m . Food and Feeding. Cox’s Leaf-nosed Bat feeds on small beedes, moths, and other insects over shrubs, creeks and water bodies by using aerial and gleaning tactics. It can fly slowly and hover during gleaning. Breeding. No information. Activity patterns. Cox’s Leaf-nosed Bat is a cave-dwelling species. Echolocation is a typical CF/FM type, with a call frequency of the F segment of46-52 kHz. Movements, Home range and Social organization. Cox’s Leaf-nosed Bat roosts in caves, typically in small groups of 2-5 individuals. At Wind Cave, Sarawak , it was recorded roosting on the cave ceiling in a group of less than 20 individuals near an opening. Status and Conservation. Classified as Endangered on The IUCNRed List. Cox’s Leafnosed Bat is endemic to Borneo and is rare; its population is very small and decreasing (area of occupancy is less than 500 km 2). Besides habitat loss, the main threats include mining and bird-nest harvesting. Recorded in Bako, Tanjung Datu, and Gunung Mulu national parks and Wind and Fairy cave reserves. Bibliography. Corbet & Hill (1992), Khan eta/. (2008), MacArthur (2016a), Phillipps & Phillipps (2016), Rosli et al. (2018), Simmons (2005).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Hipposideros coxi	Hipposideros		coxi	Shelford	1901	0	Ann. Mag. Nat. Hist.	ser. 7, 8: 113	Cox's Leaf-nosed Bat	None.	Malaysia, Borneo, Sarawak, Mt. Penrisen, 4,200 ft. (1,280 m).	Sarawak (Borneo, Malaysian part).	Not listed.	Endangered	 bicolor species group.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hipposideros coxi	23	Cox's Leaf-nosed Bat	Cox's Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Hipposideros	NA	coxi	Shelford	1901	0	Hipposideros_Coxi	Shelford, R. (1901). On a new bat from Borneo. The Annals and magazine of natural history; zoology, botany, and geology, ser. 7, 8, 113.	https://www.biodiversitylibrary.org/item/94923#page/137/mode/1up	BM 1901.6.23.1		"Mount Penrisen, Sarawak, Borneo [Malaysia], 4200 feet [= 1280 m]."			coxi Shelford, 1901	NA	NA	Malaysia|Indonesia?	Asia	Indomalaya	EN	0	0	0	Hipposideros_coxi	0	sciname match	Hipposideros_coxi	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10123	Hipposideros coxi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Hipposideros	coxi	Shelford, 1901		20000000	Hipposideros coxi	Endangered	B2ab(iii)	2016	2015-12-10 00:00:00 UTC	3.1	English	<p><span lang="EN-MY">This species is assessed as Endangered because its area of occupancy (AOO) is less than<span lang="EN-MY"><span lang="EN-MY"><span lang="EN-MY"> ;500 km<span lang="EN-MY">² (based on the extent of suitable habitat and records of the species' presence). I<span lang="EN-MY">t is also severely fragmented and continuing decline is inferred in the area, extent and quality of habitat outside the protected area coverage of 41 km<span lang="EN-MY">²<span lang="EN-MY"> in southwestern Sarawak.</span></span></span></span></p>	<p>The species has been captured at cave entrances (Abdullah et al . 2010, Rahman et al . 2011, B. Lim unpub. data) and recorded roosting in caves in small groups of two to five individuals (S. Shazali 2014, E. McArthur 2015 unpub. data). At Bako and Tanjung Datu National Parks it was captured along narrow trails in the forest understorey (S. Habeebur, pers. comm). </p>	<p><span lang="EN-MY">Outside of protected areas, forest conversion, fragmentation of habitat and economic activities such as mining, quarrying, and birds’ nest harvesting (Yen 2004, Struebig 2010, Rahman 2011) are likely to be major threats to this rare species.</p>	<p>Little is known about the abundance, population size and trends for this species. Only a few individual have been captured at each locality. There is only one known roost site with two to five individuals in Wind Cave, Bau Limestone Area (S. Shazali 2014, unpub. data). It is only known from a few individuals in one cave from Gunung Mulu National Park (B. Lim 2013; E. McArthur 2015, unpub. data).<span lang="EN-MY"> </p>	Decreasing	<p>The type locality for H. coxi is at 1,280 m asl on Mount Penrisen, south west Sarawak, Borneo, Malaysia (Simmons 2005). It has been recorded in Bako National Park (Payne et al . 1985, Khan et al . 2008), Bau Limestone Area (Khan et al . 2008, Abdullah et al . 2010, Rahman et al . 2011) and Tanjung Datu National Park in southwestern Sarawak (Khan et al . 2014) and Gunung Mulu National Park (B. Lim 2013, unpub. data) in northern Sarawak. It has been reported in Central Kalimantan (Struebig et al . 2010). However, reports of the species from acoustic surveys, in this area, have yet to be substantiated with capture data and are likely to represent H. cervinus (M. Struebig, pers. comm). </p>		Terrestrial	It occurs in a few protected areas within its range: Bako, Tanjung Datu and Mulu National Parks, Wind and Fairy Cave Nature reserves. Protected area coverage in southwestern Sarawak is small (41 km²) and highly fragmented. There is an urgent need for survey and research work to improve the understanding of ecology and population biology of these bats.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Hipposideros		coxi	Shelford	1901	0	Ann. Mag. Nat. Hist.	ser. 7, 8: 113	Cox's Leaf-nosed Bat	None.	Malaysia, Borneo, Sarawak, Mt. Penrisen, 4,200 ft. (1,280 m).	Sarawak (Borneo, Malaysian part).	Not listed.	Endangered	 bicolor species group.	Hipposideros coxi	1004588	23	Cox's Leaf-nosed Bat	Cox's Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Hipposideros	NA	coxi	Shelford	1901	0	Hipposideros_Coxi	Shelford, R. (1901). On a new bat from Borneo. The Annals and magazine of natural history; zoology, botany, and geology, ser. 7, 8, 113.	https://www.biodiversitylibrary.org/item/94923#page/137/mode/1up	BM 1901.6.23.1		"Mount Penrisen, Sarawak, Borneo [Malaysia], 4200 feet [= 1280 m]."			coxi Shelford, 1901	NA	NA				Malaysia|Indonesia?	Asia	Indomalaya	EN	0	0	0	Hipposideros_coxi	0	sciname match	Hipposideros_coxi	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hipposideros_coxi	1004588	23	Cox's Roundleaf Bat	Cox's Roundleaf Bat|Cox's Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Hipposideros	NA	coxi	Shelford	0	Hipposiderus Coxi	Shelford, R. 1901-08-01. On a new bat from Borneo. Annals and Magazine of Natural History (7)8(44):113-114.	https://www.biodiversitylibrary.org/page/29980157	BMNH:Mamm:1901.6.23.1	holotype	https://data.nhm.ac.uk/object/b1f53a07-c6b8-42dc-9181-1d78e6395b50	"Mount Penrisen, Sarawak, Borneo [Malaysia], 4200 feet [= 1280 m]."			NA	NA				Malaysia|Indonesia?	Asia	Indomalaya	EN	0	0	0	Hipposideros_coxi	0	sciname match	Hipposideros_coxi	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Hipposideros		coxi	Shelford	1901	0	Ann. Mag. Nat. Hist.	ser. 7, 8: 113	Cox's Leaf-nosed Bat	None.	Malaysia, Borneo, Sarawak, Mt. Penrisen, 4,200 ft. (1,280 m).	Sarawak (Borneo, Malaysian part).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10123/22096963/' target='_blank'>Endangered</a>	bicolorspecies group.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hipposideros coxi; Hipposideros coxi; Hipposideros coxi; Hipposideros coxi; Hipposideros coxi; Hipposideros coxi; coxi S; Phyllorhine de Cox; Cox-Rundblattnase; Hiposidérido de Cox; Cox's Roundleaf Bat; Cox's Leaf-nosed Bat; Cox's Roundleaf Bat; Cox's Leaf-nosed Bat; Cox's Leaf-nosed Bat; H. coxi