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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L382	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	Hipposideros corynophyllus	Hipposideros corynophyllus	Hipposideros corynophyllus	Hipposideros corynophyllus	Doryrhina corynophyllus	Hipposideros corynophyllus	Doryrhina corynophyllus	Hipposideros corynophyllus	Hipposideros corynophyllus	Hipposideros corynophyllus	Hipposideros corynophyllus	Hipposideros corynophyllus	Hipposideros corynophyllus		[MSW3] cyclops species group. Reviewed by Flannery and Colgan (1993); also see Flannery (1995a) and Bonaccorso (1998).; [HMW] Hipposideros corynophyllus Hill, 1985 , “Inum Tem (Cave), 3 km ENE of Telefomin, West Sepik Province , Papua New Guinea , 1800 m , c. 5°10’S , 141°35’E .” This species is monotypic.; [batnames2022] Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclops species group. Reviewed by Flannery and Colgan (1993); also see Flannery (1995a) and Bonaccorso (1998).; [MDD2022] moved from Hipposideros to Doryrhina; [batnames2023] Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclops species group. Reviewed by Flannery and Colgan (1993); also see Flannery (1995a) and Bonaccorso (1998).; [MDD2023] has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being; [MDD2025_2.0] has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being; [batnames2025_1.7] Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclopsspecies group. Reviewed by Flannery and Colgan (1993); also see Flannery (1995a) and Bonaccorso (1998).; [MDD2025_2.2] has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being														corynophyllus				corynophyllus 	corynophyllus 			corynophyllus J. Edwards Hill, 1986						N/A					Distribution: Known only from central New Guinea.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		E New Guinea, ref. 4.117	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Hill	1985	Mammalia, 49:527.		C New Guinea.	Papua New Guinea, W Sepik, 3 km ENE Telephomin, 1800 m.		HILL	1985	Posterior lateral leaflet extending anteriorly be neath the anterior noseleaf. Premaxillae narrow, not enclosing the junction with the maxillae. Me dian process of intermediate noseleaf well devel oped and club-shaped. Skull with a shallow frontal depression. A shallow sphenoidal depres sion present. Posterior ridge of last upper molar considerably reduced. Width of each cochlea about four times their distance apart. Rostrum relatively elongate. Interorbital region relatively wide. Size medium (forearm length, 48-49 mm).	Distribution: Known only from central New Guinea.	No subspecies.		64	species	H. corynophyllus	HILL	1985	Hipposideros	genus	Hipposideros corynophyllus				Posterior lateral leaflet extending anteriorly be neath the anterior noseleaf. Premaxillae narrow, not enclosing the junction with the maxillae. Me dian process of intermediate noseleaf well devel oped and club-shaped. Skull with a shallow frontal depression. A shallow sphenoidal depres sion present. Posterior ridge of last upper molar considerably reduced. Width of each cochlea about four times their distance apart. Rostrum relatively elongate. Interorbital region relatively wide. Size medium (forearm length, 48-49 mm).	No subspecies.		37. H. corynophyllus HILL 1985 [cyclops group].	37	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Hipposideros corynophyllus	Hipposideros		corynophyllus	Hill		1985		Mammalia	49		527		Telefomin Leaf-nosed Bat	Papua New Guinea, W Sepik, 3 km ENE Telefomin, 1,800 m.	C New Guinea.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.		cyclops species group. Reviewed by Flannery and Colgan (1993); also see Flannery (1995a) and Bonaccorso (1998).	03BD87A2C675A207FF22F275F58845A3	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	230	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C675A207FF22F275F58845A3.xml	Doryrhina corynophyllus	Hipposideridae	Doryrhina	corynophyllus		1986	Phyllorhine de Telefomin @fr | Telefomin-Rundblattnase @de | Doryrina de Telefomin @es | Telefomin Roundleaf Bat @en	Hipposideros corynophyllus Hill, 1985 , “Inum Tem (Cave), 3 km ENE of Telefomin, West Sepik Province , Papua New Guinea , 1800 m , c. 5°10’S , 141°35’E .” This species is monotypic.	Known only from a small area around Tifalmin and Telefomin in Sandaun Province , W Papua New Guinea , and also collected in Tembagapura region, Papua Province , Indonesia . Even so, its echolocation calls have been recorded in other areas and it is thought that this species has a larger distribution, probably occurring throughout the Central Range ofNew Guinea .	Head—body 54-9-63 mm, tail 9—12 mm , ear 18-9—22-6 mm, hindfoot 9-5—11-9 mm, forearm 48-54-6 mm; weight 14-5— 17 g . The Telefomin Leaf-nosed Bat has an elongated rostrum and relatively long hair, which is densely distributed even on face and muzzle. Brownish colors are present on dorsum, with paler silver on ventral area. Fur is long (c.14-2 mm). Ears are relatively long, distally narrowed, and convex on anterior or medial margin of ear, without antitragal lobe. Noseleaf covers much of muzzle, with two lateral leaflets. Anterior leaf has no median emargination, intermedian leaf is elevated laterally, and posterior leaf is high. Median processes of intermediate and posterior leaves are well developed and club-shaped. Lower club is very long and tubular.	The Telefomin Leaf-nosed Bat is found in deep limestone caves, usually located at elevations of 1600—1800 m . It also occurs in forests and in other subterranean cavities. The altitudinal range of this species is very wide, ranging from close to sea level up to 2700 m .	The Telefomin Leaf-nosed Bat probably forages in forest habitats.	The breeding biology of the Telefomin Leaf-nosed Bat is largely unknown; it is probably similar to that ofWollaston’s Leafnosed Bat (D. woUastoni).	Activity patterns of the Telefomin Leaf-nosed Bat are probably similar to those ofWollaston’s Leaf-nosed Bat.	The Telefomin Leaf-nosed Bat may be found either solitarily or in small groups. It sometimes shares caves with Wollaston’s Leaf-nosed Bat (subspecies pamabyi) in the Telefomin area.	Classified as Least Concern on The IUCNRed List (as Hipposideros corynophyllus ). Current population trends are unknown, but population is probably not strongly fragmented. Forest loss and modification might be the major threats for the Telefomin Leaf-nosed Bat, although large areas of intact forest still remain within its range. More research is needed to study and assess the ecology and population status of this species.	Armstrong & Aplin (2017b) | Bonaccorso (1998) | Decher & ahr (2005) | lannery & Colgan (1993) | lannery & Seri (1990) | Helgen (2007a) | Hill (1985)	https://zenodo.org/record/3749051/files/figure.png	6. Telefomin Leaf-nosed Bat Doryrhina corynophyllus French: Phyllorhine deTelefomin / German: Telefomin-Rundblattnase I Spanish: Doryrina deTelefomin Other common names: Telefomin Roundleaf Bat Taxonomy. Hipposideros corynophyllus Hill, 1985 , “Inum Tem (Cave), 3 km ENE of Telefomin, West Sepik Province , Papua New Guinea , 1800 m , c. 5°10’S , 141°35’E .” This species is monotypic. Distribution. Known only from a small area around Tifalmin and Telefomin in Sandaun Province , W Papua New Guinea , and also collected in Tembagapura region, Papua Province , Indonesia . Even so, its echolocation calls have been recorded in other areas and it is thought that this species has a larger distribution, probably occurring throughout the Central Range ofNew Guinea . Descriptive notes. Head—body 54-9-63 mm, tail 9—12 mm , ear 18-9—22-6 mm, hindfoot 9-5—11-9 mm, forearm 48-54-6 mm; weight 14-5— 17 g . The Telefomin Leaf-nosed Bat has an elongated rostrum and relatively long hair, which is densely distributed even on face and muzzle. Brownish colors are present on dorsum, with paler silver on ventral area. Fur is long (c.14-2 mm). Ears are relatively long, distally narrowed, and convex on anterior or medial margin of ear, without antitragal lobe. Noseleaf covers much of muzzle, with two lateral leaflets. Anterior leaf has no median emargination, intermedian leaf is elevated laterally, and posterior leaf is high. Median processes of intermediate and posterior leaves are well developed and club-shaped. Lower club is very long and tubular. Habitat. The Telefomin Leaf-nosed Bat is found in deep limestone caves, usually located at elevations of 1600—1800 m . It also occurs in forests and in other subterranean cavities. The altitudinal range of this species is very wide, ranging from close to sea level up to 2700 m . Food and Feeding. The Telefomin Leaf-nosed Bat probably forages in forest habitats. Breeding. The breeding biology of the Telefomin Leaf-nosed Bat is largely unknown; it is probably similar to that ofWollaston’s Leafnosed Bat (D. woUastoni). Activity patterns. Activity patterns of the Telefomin Leaf-nosed Bat are probably similar to those ofWollaston’s Leaf-nosed Bat. Movements, Home range and Social organization. The Telefomin Leaf-nosed Bat may be found either solitarily or in small groups. It sometimes shares caves with Wollaston’s Leaf-nosed Bat (subspecies pamabyi) in the Telefomin area. Status and Conservation. Classified as Least Concern on The IUCNRed List (as Hipposideros corynophyllus ). Current population trends are unknown, but population is probably not strongly fragmented. Forest loss and modification might be the major threats for the Telefomin Leaf-nosed Bat, although large areas of intact forest still remain within its range. More research is needed to study and assess the ecology and population status of this species. Bibliography. Armstrong & Aplin (2017b), Bonaccorso (1998), Decher & ahr (2005), lannery & Colgan (1993), lannery & Seri (1990), Helgen (2007a), Hill (1985).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Hipposideros corynophyllus	Hipposideros		corynophyllus	Hill	1985	0	Mammalia	57:47:00	Telefomin Leaf-nosed Bat	None.	Papua New Guinea, W Sepik, 3 km ENE Telefomin, 1,800 m.	C New Guinea.	Not listed.	Least Concern	Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclops species group. Reviewed by Flannery and Colgan (1993); also see Flannery (1995a) and Bonaccorso (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Doryrhina corynophyllus	23	Telefomin Leaf-nosed Bat	Telefomin Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Doryrhina	NA	corynophyllus	J. Edwards Hill	1985	1						"Inum Tem (Cave), 3 km ENE of Telefomin, West Sepik Province, Papua New Guinea, 1800 m, c. 5°10'S, 141°35'E."	-5.167	141.58	corynophyllus (J. Edwards Hill, 1985)	moved from Hipposideros to Doryrhina	Foley, N. M., Goodman, S. M., Whelan, C. V., Puechmaille, S. J., & Teeling, E. (2017). Towards navigating the Minotaur's labyrinth: cryptic diversity and taxonomic revision within the speciose genus Hipposideros (Hipposideridae). Acta Chiropterologica, 19(1), 1-18.	Indonesia|Papua New Guinea	Oceania	Australasia/Oceania	LC	0	0	0	Doryrhina_corynophyllus	0	sciname match	Hipposideros_corynophyllus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10122	Hipposideros corynophyllus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Hipposideros	corynophyllus	Hill, 1985		200000000	Hipposideros corynophyllus	Least Concern		2021	2016-07-31 00:00:00 UTC	3.1	English	<p>The few specimens of Hipposideros corynophyllus available in museums were collected as recently as 30 years ago, and are all derived from a relatively small area near Tifalmin and Telefomin in Sandaun (West Sepik) Province of Papua New Guinea. Acoustic surveys within the last five years have recorded echolocation calls attributable to this species further north and south but not significantly further afield (K.N. Armstrong and K.P. Aplin unpublished data; Armstrong and Aplin 2014). Assuming that the identification of the acoustic recordings are correct, the calculated extent of occurrence around all records is below 20,000 km². However, there is also a record 500 km west in the Tembagapura area of Papua Province, Indonesia (Helgen 2007; represented by a specimen in the Australian Museum) that suggests a wider distribution throughout the central cordillera of New Guinea. Forest removal and modification has occurred following modern developments around Telefomin, and seems likely to continue, but there are still large amounts of intact habitat within its range further afield. It is likely that future surveys will find this species across a wider area. In absence of evident threats and given its likely wide distribution, this species is listed as Least Concern (LC).</p>	This species has been recorded in deep limestone cave complexes. It may be solitary or aggregate in small groups (Bonaccorso 1998). It forages within forest habitat. Very little is known of its ecology otherwise, but is likely to be similar to Hipposideros wollastoni .	This species is currently known from very few roosting sites, and it is unclear if there are any threats to these localities. Development around Telefomin has led to forest habitat loss, but there is significant area of continuous intact forest further afield within its revised extent of occurrence.	Bonaccorso (1998) states that it is a rare species, but Flannery (1995) suggests that it is common in deep caves at 1,600 to 1,800 m a.s.l. It is likely to be locally common, though observations from acoustic surveys suggest that it occurs at relatively low density compared to some other Hipposideridae and Rhinolophidae (K.N. Armstrong and K.P. Aplin unpublished data).	Unknown	<p>Hipposideros corynophyllus is represented by almost 30 specimens in museum collections (Natural History Museum, London, Australian Museum, South Australian Museum) from around Tifalmin and Telefomin in Sandaun (West Sepik) Province, with the most recent collection almost 30 years ago. Echolocation calls attributable to this species have been recorded in the past five years from multiple localities both south and north of Telefomin, but not a significant distance east of this area (K.N. Armstrong and K.P. Aplin unpublished data; Armstrong and Aplin 2014). It has also been collected in the Tembagapura area of Papua Province, Indonesia and a wider presence throughout the central cordillera was suggested on the basis of this capture record (Helgen 2007). The altitudinal range appears to large, with collections from the Telefomin area up to 1,800 m Asl, acoustic records close to sea level and the Tembagapura record as high as 2,700 m.</p>		Terrestrial	<p>A suitable conservation measure for this species would be the protection of the known roosting sites. Further studies into the distribution, ecology, and threats to this species are needed.</p>	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Hipposideros		corynophyllus	Hill	1985	0	Mammalia	57:47:00	Telefomin Leaf-nosed Bat	None.	Papua New Guinea, W Sepik, 3 km ENE Telefomin, 1,800 m.	C New Guinea.	Not listed.	Least Concern	Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclops species group. Reviewed by Flannery and Colgan (1993); also see Flannery (1995a) and Bonaccorso (1998).	Hipposideros corynophyllus	1004567	23	Telefomin Leaf-nosed Bat	Telefomin Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Hipposideros	NA	corynophyllus	J. Edwards Hill	1985	0						"Inum Tem (Cave), 3 km ENE of Telefomin, West Sepik Province, Papua New Guinea, 1800 m, c. 5°10'S, 141°35'E."	-5.16667	141.5833	corynophyllus (J. Edwards Hill, 1985)	has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being	Foley, N. M., Goodman, S. M., Whelan, C. V., Puechmaille, S. J., & Teeling, E. (2017). Towards navigating the Minotaur's labyrinth: cryptic diversity and taxonomic revision within the speciose genus Hipposideros (Hipposideridae). Acta Chiropterologica, 19(1), 1-18.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.				Indonesia|Papua New Guinea	Oceania	Australasia/Oceania	LC	0	0	0	Doryrhina_corynophyllus	0	sciname match	Hipposideros_corynophyllus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hipposideros_corynophyllus	1004567	23	Telefomin Roundleaf Bat	Telefomin Roundleaf Bat|Telefomin Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Hipposideros	NA	corynophyllus	J. Edwards Hill	0	Hipposideros corynophyllus	Hill, J.E. 1986-03-20. Records of bats (Chiroptera) from New Guinea, with the description of a new _Hipposideros_ (Hipposideridae). Mammalia 49(4):525-536.	https://doi.org/10.1515/mamm.1985.49.4.525	BMNH:Mamm:1984.1898	holotype	https://data.nhm.ac.uk/object/c6dc90af-ee6e-4289-a928-34775a766675	"Inum Tem (Cave), 3 km ENE of Telefomin, West Sepik Province, Papua New Guinea, 1800 m, c. 5°10'S, 141°35'E."	-5.16667	141.5833	has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being	Foley, N. M., Goodman, S. M., Whelan, C. V., Puechmaille, S. J., & Teeling, E. (2017). Towards navigating the Minotaur's labyrinth: cryptic diversity and taxonomic revision within the speciose genus Hipposideros (Hipposideridae). Acta Chiropterologica, 19(1), 1-18.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.				Indonesia|Papua New Guinea	Oceania (Continent)	Australasia	LC	0	0	0	Doryrhina_corynophyllus	0	sciname match	Hipposideros_corynophyllus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Hipposideros		corynophyllus	Hill	1985	0	Mammalia	57:47:00	Telefomin Leaf-nosed Bat	None.	Papua New Guinea, W Sepik, 3 km ENE Telefomin, 1,800 m.	C New Guinea.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10122/209536979/' target='_blank'>Least Concern</a>	Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclopsspecies group. Reviewed by Flannery and Colgan (1993); also see Flannery (1995a) and Bonaccorso (1998).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hipposideros corynophyllus; Doryrhina corynophyllus; Hipposideros corynophyllus; Doryrhina corynophyllus; Hipposideros corynophyllus; Hipposideros corynophyllus; corynophyllus; Phyllorhine de Telefomin; Telefomin-Rundblattnase; Doryrina de Telefomin; Telefomin Roundleaf Bat; Telefomin Leaf-nosed Bat; Telefomin Roundleaf Bat; Telefomin Leaf-nosed Bat; Telefomin Leaf-nosed Bat; H. corynophyllus