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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L376	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Hipposideros caffer	Hipposideros caffer	Hipposideros caffer	Hipposideros caffer	Hipposideros caffer	Hipposideros caffer	Hipposideros caffer	Hipposideros caffer	Hipposideros caffer	Hipposideros caffer	Hipposideros caffer	Hipposideros caffer	Hipposideros caffer	Hipposideros caffer	Hipposideros caffer		[MSW3] bicolor species group. Includes tephrus; see Hayman and Hill (1971). Reviewed in part by Harrison and Bates (1991) and Horácek et al. (2000). See Taylor (2000a) for distribution map. Subspecies limits are somewhat unclear, and it is possible that this complex includes more than one species.; [HMW] Rhinolophus caffer Sundevall, 1846 , near Durban, KwaZulu-Natal Province , South Africa . Hipposideros caffer was formerly included in the bicolor species group, but is now placed in the ruber species group. There has been much confusion in the literature regarding the status of this species, particularly with respect to the rainforest H. ruber species complex. Recent molecular work suggests that H. caffer is present in southern Africa, but may also extend into East Africa. The validity of the proposed subspecies angolensis from Angola is uncertain, and the internal taxonomy of this species requires reassessment. Monotypic.; [batnames2022]  bicolor species group. Includes tephrus; see Hayman and Hill (1971). Reviewed in part by Harrison and Bates (1991) and Horácek et al. (2000).See Taylor (2000a) for distribution map. Subspecies limits are somewhat unclear, and it is possible that this complex includes more than onespecies.; [MDD2022] previously included H. tephrus; [IUCN] This species was previously part of the Hipposideros ruber complex. However, Vallo et al.  (2008) have somewhat clarified the taxonomy of this group and show that H. cafer s.s. is restricted to southern African. The northern limits of this species are currently unknown and may include Tanzania and Kenya (Vallo et al.  2008, Monadjem et al.  2013).; [batnames2023]  bicolor species group. Includes tephrus; see Hayman and Hill (1971). Reviewed in part by Harrison and Bates (1991) and Horácek et al. (2000).See Taylor (2000a) for distribution map. Subspecies limits are somewhat unclear, and it is possible that this complex includes more than onespecies.; [MDD2023] previously included H. tephrus; [MDD2025_2.0] previously included H. tephrus; [batnames2025_1.7] bicolorspecies group. Does not include tephrus; but see Hayman and Hill (1971). Reviewed in part by Harrison and Bates (1991) and Horácek et al. (2000).See Taylor (2000a) for distribution map. Subspecies limits are somewhat unclear, and it is possible that this complex includes more than onespecies.; [MDD2025_2.2] previously included H. tephrus				braima, nanus, auriantiaca		angolensis, aurantiaca, bicornis, braima, gracilis, nanus, tephrus (see Hayman and Hill, 1971:29).	caffer, tephrus, nanus, angolensis	caffer, angolensis, nanus, tephrus	aurantiaca, bicornis, gracilis; tephrus - braima			caffer, angolensis, nanus, tephrus	caffer - aurantiaca, bicornis, gracilis; tephrus - braima	caffercaffra, gracilis, patellifera, bicornis, angolensis, nanus, aurantiaca, braima	This species was previously part of the Hipposideros ruber complex. However, Vallo et al.  (2008) have somewhat clarified the taxonomy of this group and show that H. cafer s.s. is restricted to southern African. The northern limits of this species are currently unknown and may include Tanzania and Kenya (Vallo et al.  2008, Monadjem et al.  2013).	caffer, angolensis, nanus	caffer - aurantiaca, bicornis, gracilis, braima	caffer, caffra, gracilis, patellifera, bicornis, angolensis, nanus, aurantiaca, braima	caffer, gracilis, patellifera, bicornis, angolensis, nanus, aurantiacus	angolensis, caffer, nanus	caffer - aurantiaca, bicornis, braima, gracilis	caffer (Sundevall, 1847)|gracilis (W. C. H. Peters, 1851)|patellifera (W. C. H. Peters, 1852) [not used as valid]|bicornis (von Heuglin, 1861)|angolensis (de Seabra, 1898)|nanus J. A. Allen in J. A. Allen, Lang, & Chapin, 1917|aurantiacus De Beaux, 1924		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Sundevall's leaf-nosed bat Africa, Arabia	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Hipposideros caffer	South Africa, Natal, near Durban.	Sundevall	1846	Ofv. Kongl. Svenska Vet.-Akad. Forhandl. Stockholm, 3(4):118.	Distribution: Widely distributed in sub-Saharan Africa, south to the Cape Province (including Pemba and Zanzibar islands), but largely avoiding forested areas; also Morocco and southwestern Arabia).		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Sundevall's leaf-nosed bat	Africa, Arabia	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Sundevall	1846	Ofv. Kongl. Svenska Vet.-Akad. Forhandl. Stockholm, 3(4):118.		Most of subsaharan Africa except the central forested region; Morocco; Yemen; Zanzibar and Pemba.	South Africa, Natal, near Durban.		SUNDEVALL	1846	Two lateral leaflets present, neither one extend ing anteriorly beneath anterior noseleaf to median line. Posterior noseleaf with a well-developed transverse serrated supplementary structure on its posterior face, but no well-defined septa. Anterior upper premolar small but not extruded from toothrow. Median posterior narial compartment of rostral swelling relatively narrow, with wide lateral inflations. Size medium (forearm length 42-51 mm).	Distribution: Widely distributed in sub-Saharan Africa, south to the Cape Province (including Pemba and Zanzibar islands), but largely avoiding forested areas; also Morocco and southwestern Arabia).	Four subspecies are cur rently recognized (HAYMAN & HILL 1971):	H. c. caffer (entire range of species except for Morocco, the dry savannas immediately south of the Sahara, north eastern Zaire, and southwestern Africa), H. c. tephrus (Mo rocco, also dry sub-Saharan belt from Mauretania and Guinea to Sudan), H. c. nanus (northeastern Zaire), H. c. angolensis (Gabon south to Namibia).	63	species	H. caffer	SUNDEVALL	1846	Hipposideros	genus	Hipposideros caffer				Two lateral leaflets present, neither one extend ing anteriorly beneath anterior noseleaf to median line. Posterior noseleaf with a well-developed transverse serrated supplementary structure on its posterior face, but no well-defined septa. Anterior upper premolar small but not extruded from toothrow. Median posterior narial compartment of rostral swelling relatively narrow, with wide lateral inflations. Size medium (forearm length 42-51 mm).	Four subspecies are cur rently recognized (HAYMAN & HILL 1971):		27. H. caffer (SUNDEVALL 1846) [bicolor group].	27	_H. c. angolensis_ (Seabra, 1898); _H. c. caffer_ (Sundevall, 1847) (synonyms: _aurantiacus_ De Beaux, 1924, _gracilis_ (Peters, 1851), _patellifera_ (Peters, 1852)); _H. c. nanus_ Allen, 1917			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Hipposideros caffer	Hipposideros		caffer	Sundevall	y	1846		Öfv. Kongl. Svenska Vet.-Akad. Forhandl. Stockholm	3	4	118		Sundevall's Leaf-nosed Bat	South Africa, KwaZulu-Natal Prov., near Durban.	SW Arabian Peninsula including Yemen; most of subsaharan Africa except the central forested region; Morocco; Zanzibar and Pemba.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	aurantiaca De Beaux, 1924; bicornis Heuglin, 1861; gracilis Peters, 1825; angolensis Seabra, 1898; nanus J. A. Allen, 1917; tephrus Cabrera, 1906; braima Monard, 1939.	bicolor species group. Includes tephrus; see Hayman and Hill (1971). Reviewed in part by Harrison and Bates (1991) and Horácek et al. (2000). See Taylor (2000a) for distribution map. Subspecies limits are somewhat unclear, and it is possible that this complex includes more than one species.	03BD87A2C662A211F8A4F4B7FA814A03	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	247	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C662A211F8A4F4B7FA814A03.xml	Hipposideros caffer	Hipposideridae	Hipposideros	caffer		1847	Phyllorhine de Cafrerie @fr | Gewöhnliche Rundblattnase @de | Hiposidérido de Cafreria @es | Cape Leaf-nosed Bat @en | Common African Leaf-nosed Bat @en | Lesser Leaf-nosed bat @en | Sundevall’s Roundleaf Bat @en	Rhinolophus caffer Sundevall, 1846 , near Durban, KwaZulu-Natal Province , South Africa . Hipposideros caffer was formerly included in the bicolor species group, but is now placed in the ruber species group. There has been much confusion in the literature regarding the status of this species, particularly with respect to the rainforest H. ruber species complex. Recent molecular work suggests that H. caffer is present in southern Africa, but may also extend into East Africa. The validity of the proposed subspecies angolensis from Angola is uncertain, and the internal taxonomy of this species requires reassessment. Monotypic.	Widely in S Africa, but N & W boundaries are not yet known; known to occur in S DR Congo , W Angola , Zambia , Malawi , Mozambique , Namibia , N Botswana , Zimbabwe , South Africa , and Swaziland . A species morphologically identical to Ä caffer occurs widely in East Africa but whether this refers to H. caffer or H. tephrus has not yet been established.	Head-body 45-60 mm, tail 25-38 mm, ear 10-17 mm, hindfoot 6-10 mm, forearm 42-52 mm; weight 5-11 g. No obvious sexual differences in pelage or size although males tend to average slightly larger than females, but with considerable overlap. Sundevall’s Leaf-nosed Bat has short, rounded wings. Muzzle is relatively short with large but simple noseleaf that has two lateral leaflets. A frontal sac is present in both sexes, but greatly reduced in females. Intemarial septum is not swollen and does not partially cover the nares. Ears are separate, and relatively small. Fur is fine and fluffy, gray or brownish gray dorsally, paler ventrally; an orange morph exists, which is bright orange or rusty brown dorsally and paler ventrally. Chromosomal complement has 2n = 32 and FN = 60.	A wide range of wooded savanna habitats, including riparian forest, but avoids rainforest. In more arid regions, its distribution appears to be limited to riverine habitats.	Sundevall’s Leaf-nosed Bat is insectivorous, feeding predominandy on moths but possibly also caddisflies, beedes, and other arthropods. Moths are typically taken in proportion to availability, but clicking moths of the families Geometridae and Arctiidae are taken less frequently than would be expected.	Breeding in South Africa , Uganda , and Kenya (the last two populations may or may not refer to this species) is seasonal with births of single young occurring at the beginning of the rains. In South Africa , copulation and fertilization occur in April followed by a period of retarded development during the cool dry season, with births in December. Hence, gestation is 220 days at these latitudes. In Uganda , females are pregnant between December and March with births in the latter month. Young are fed milk for 60 days.	Sundevall’s Leaf-nosed Bat roosts during the day in a variety of cavities and structures including caves, hollow trees, holes in the ground, old mines, culverts under roads, and disused houses and outhouses. It remains active throughout the year, even at high latitudes in South Africa . This species, like its congeners, has a low wing loading and low aspect ratio, allowing it to fly in cluttered environments, such as thickets. Echolocation call includes a F component usually at (or near) 145 kHz, but which is highly variable by up to 15 kHz above and below.	Sundevall’s Leaf-nosed Bat typically roosts in colonies of up to several hundred individuals. However, it also roosts in small family groups, and occasionally may roost singly. In one study in Zimbabwe , a group of one adult male and seven lactating females roosted together in the same hollow tree, suggestive ofa harem mating system where a dominant male monopolizes several females. It occasionally shares its roosts with other bat species including Egyptian Slitfaced Bats (Nycteris thebaica), Rhinolophus spp. , Temminck’s Myotis (Myotis tricolor), and Natal Long-fingered Bats (Miniopterus natalensis).	Classified as Least Concern on The IUCNRed List. Sundevall’s Leaf-nosed Bat has a wide distribution, utilizes many different structures as day roosts, and usually occurs in good numbers.	Aldridge & Rautenbach (1987) | Bell (1987) | Bell & Fenton (1984) | Bernard & Cumming (1997) | Bernard & Happold (2013b) | Bernard & Meester (1982) | Bowie et al. (1999) | Cotterill (2001a) | Fenton (1985a) | Happold & Happold (1990) | Hill (1963a) | Monadjem & Reside (2008) | Monadjem, Reside & Lumsden (2007) | Monadjem, Schoeman et al. (2010) | Monadjem, Taylor et al. (2010) | Mutere (1970) | O'Shea & Vaughan (1980) | Rautenbach (1982) | Rautenbach et a/. (1993) | Schoeman & Jacobs (2008, 2011) | Thorn & Kerbis Peterhans (2009) | Vallo et al. (2008) | Van Cakenberghe et al. (2017) | Whitaker & Black (1976)	https://zenodo.org/record/3749149/files/figure.png	57. Sundevall’s Leaf-nosed Bat Hipposideros caffer French: Phyllorhine de Cafrerie I German: Gewöhnliche Rundblattnase / Spanish: Hiposidérido de Cafreria Other common names: Cape Leaf-nosed Bat , Common African Leaf-nosed Bat , Lesser Leaf-nosed bat , Sundevall’s Roundleaf Bat Taxonomy. Rhinolophus caffer Sundevall, 1846 , near Durban, KwaZulu-Natal Province , South Africa . Hipposideros caffer was formerly included in the bicolor species group, but is now placed in the ruber species group. There has been much confusion in the literature regarding the status of this species, particularly with respect to the rainforest H. ruber species complex. Recent molecular work suggests that H. caffer is present in southern Africa, but may also extend into East Africa. The validity of the proposed subspecies angolensis from Angola is uncertain, and the internal taxonomy of this species requires reassessment. Monotypic. Distribution. Widely in S Africa, but N & W boundaries are not yet known; known to occur in S DR Congo , W Angola , Zambia , Malawi , Mozambique , Namibia , N Botswana , Zimbabwe , South Africa , and Swaziland . A species morphologically identical to Ä caffer occurs widely in East Africa but whether this refers to H. caffer or H. tephrus has not yet been established. Descriptive notes. Head-body 45-60 mm, tail 25-38 mm, ear 10-17 mm, hindfoot 6-10 mm, forearm 42-52 mm; weight 5-11 g. No obvious sexual differences in pelage or size although males tend to average slightly larger than females, but with considerable overlap. Sundevall’s Leaf-nosed Bat has short, rounded wings. Muzzle is relatively short with large but simple noseleaf that has two lateral leaflets. A frontal sac is present in both sexes, but greatly reduced in females. Intemarial septum is not swollen and does not partially cover the nares. Ears are separate, and relatively small. Fur is fine and fluffy, gray or brownish gray dorsally, paler ventrally; an orange morph exists, which is bright orange or rusty brown dorsally and paler ventrally. Chromosomal complement has 2n = 32 and FN = 60. Habitat. A wide range of wooded savanna habitats, including riparian forest, but avoids rainforest. In more arid regions, its distribution appears to be limited to riverine habitats. Food and Feeding. Sundevall’s Leaf-nosed Bat is insectivorous, feeding predominandy on moths but possibly also caddisflies, beedes, and other arthropods. Moths are typically taken in proportion to availability, but clicking moths of the families Geometridae and Arctiidae are taken less frequently than would be expected. Breeding. Breeding in South Africa , Uganda , and Kenya (the last two populations may or may not refer to this species) is seasonal with births of single young occurring at the beginning of the rains. In South Africa , copulation and fertilization occur in April followed by a period of retarded development during the cool dry season, with births in December. Hence, gestation is 220 days at these latitudes. In Uganda , females are pregnant between December and March with births in the latter month. Young are fed milk for 60 days. Activity patterns. Sundevall’s Leaf-nosed Bat roosts during the day in a variety of cavities and structures including caves, hollow trees, holes in the ground, old mines, culverts under roads, and disused houses and outhouses. It remains active throughout the year, even at high latitudes in South Africa . This species, like its congeners, has a low wing loading and low aspect ratio, allowing it to fly in cluttered environments, such as thickets. Echolocation call includes a F component usually at (or near) 145 kHz, but which is highly variable by up to 15 kHz above and below. Movements, Home range and Social organization. Sundevall’s Leaf-nosed Bat typically roosts in colonies of up to several hundred individuals. However, it also roosts in small family groups, and occasionally may roost singly. In one study in Zimbabwe , a group of one adult male and seven lactating females roosted together in the same hollow tree, suggestive ofa harem mating system where a dominant male monopolizes several females. It occasionally shares its roosts with other bat species including Egyptian Slitfaced Bats (Nycteris thebaica), Rhinolophus spp. , Temminck’s Myotis (Myotis tricolor), and Natal Long-fingered Bats (Miniopterus natalensis). Status and Conservation. Classified as Least Concern on The IUCNRed List. Sundevall’s Leaf-nosed Bat has a wide distribution, utilizes many different structures as day roosts, and usually occurs in good numbers. Bibliography. Aldridge & Rautenbach (1987), Bell (1987), Bell & Fenton (1984), Bernard & Cumming (1997), Bernard & Happold (2013b), Bernard & Meester (1982), Bowie eta/. (1999), Cotterill (2001a), Fenton (1985a), Happold & Happold (1990), Hill (1963a), Monadjem & Reside (2008), Monadjem, Reside & Lumsden (2007), Monadjem, Schoeman eta/. (2010), Monadjem, Taylor eta/. (2010), Mutere (1970), O'Shea & Vaughan (1980), Rautenbach (1982), Rautenbach et a/. (1993), Schoeman & Jacobs (2008, 2011), Thorn & Kerbis Peterhans (2009), Vallo eta/. (2008), Van Cakenberghe eta/. (2017), Whitaker & Black (1976).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Hipposideros caffer	Hipposideros		caffer	Sundevall	1846	1	&Ouml;fv. Kongl. Svenska Vet.-Akad. Forhandl. Stockholm	3(4): 118	Sundevall's Leaf-nosed Bat	 aurantiaca De Beaux, 1924; bicornis Heuglin, 1861; gracilis Peters, 1825; <b> angolensis </b> Seabra, 1898; <b>nanus</b> J. A. Allen, 1917; <b>tephrus</b> Cabrera, 1906; braima Monard, 1939.	South Africa, KwaZulu-Natal Prov., near Durban.	SW Arabian Peninsula including Yemen; most of subsaharan Africa except the central forested region; Morocco; Zanzibar and Pemba.	Not listed.	Least Concern	 bicolor species group. Includes tephrus; see Hayman and Hill (1971). Reviewed in part by Harrison and Bates (1991) and Horácek et al. (2000).See Taylor (2000a) for distribution map. Subspecies limits are somewhat unclear, and it is possible that this complex includes more than onespecies.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hipposideros caffer	23	Sundevall's Leaf-nosed Bat	Cape Leaf-nosed Bat|Common African Leaf-nosed Bat|Lesser Leaf-nosed bat|Sundevall's Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Hipposideros	NA	caffer	Sundevall	1846	1	Rhinolophus_caffer	Sundevall, C. J. (1846). Nya Mammalia frÃ¥ n Sydafrika. Öfversigt af Kongl. Vetenskaps-akademiens forhandlingar, 3(4), 118.	https://www.biodiversitylibrary.org/item/20230#page/144/mode/1up	BM 1848.6.2.16, BM 1849.11.22.11 [syntypes]		near Durban, KwaZulu-Natal Province, South Africa.			caffer (Sundevall, 1846)|caffra (W. Peters, 1852)|gracilis (W. Peters, 1852)|patellifera (W. Peters, 1852)|bicornis (Heuglin, 1861)|angolensis (Seabra, 1898)|nanus J. A. Allen, 1917|aurantiaca De Beaux, 1924|braima Monard, 1938	previously included H. tephrus	Vallo, P., Guillén-Servent, A., Benda, P., Pires, D. B., & Koubek, P. (2008). Variation of mitochondrial DNA in the Hipposideros caffer complex (Chiroptera: Hipposideridae) and its taxonomic implications. Acta Chiropterologica, 10(2), 193-206.	Democratic Republic of the Congo|Angola|Zambia|Malawi|Mozambique|Namibia|Botswana|Zimbabwe|South Africa|Eswatini	Africa	Afrotropic|Palearctic	LC	0	0	0	Hipposideros_caffer	0	sciname match	Hipposideros_caffer	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Hipposideros caffer	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Hipposideros	caffer	(Sundevall, 1846)	This species was previously part of the Hipposideros ruber complex. However, Vallo et al.  (2008) have somewhat clarified the taxonomy of this group and show that H. cafer s.s. is restricted to southern African. The northern limits of this species are currently unknown and may include Tanzania and Kenya (Vallo et al.  2008, Monadjem et al.  2013).	20000000	Hipposideros caffer	Least Concern		2020	2019-07-10 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, and because it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	This species occurs in savanna, bushveld and coastal forest, and is usually associated with rivers and other water resources (Taylor 2000), provided there are caves or buildings where it can roost during the day (Happold 1987). Colony size varies from small to medium-sized groups of tens or hundreds of individuals (Taylor 2000), exceptionally up to 500,000 (Brosset 1966; Happold 1987).	Human disturbance to roosting sites (caves) could have a negative effect.	A widely distributed and numerous species. Sundevall's leaf-nosed bats are extremely gregarious and capable of forming huge colonies where there is adequate roosting space (Monadjem et al. 2010). This species also has been found roosting in sinkholes and cavities, including human-made structures, e.g. mines and culverts (Cotterill 2001).	Decreasing	The exact range of this species is not currently known. Hipposideros caffer sensu lato occurs across most of sub-Saharan Africa except for central forested regions as well as in parts of Arabia (Simmons 2005). However, H. caffer s.s. is only known from southern African countries of Botswana, Mozambique, South Africa, Eswatini, Zimbabwe. Populations in Malawi, Zambia, and southern Tanzania may refer to this species, while the taxonomic status of East African records is currently unresolved. Elevation ranges from sea level to 2,500 m in this area.		Terrestrial	No specific conservation measures in place. Occurs in protected areas throughout its range.	Afrotropical|Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Hipposideros		caffer	Sundevall	1846	1	&Ouml;fv. Kongl. Svenska Vet.-Akad. Forhandl. Stockholm	3(4): 118	Sundevall's Leaf-nosed Bat	 aurantiaca De Beaux, 1924; bicornis Heuglin, 1861; gracilis Peters, 1825; <b> angolensis </b> Seabra, 1898; <b>nanus</b> J. A. Allen, 1917; <b>tephrus</b> Cabrera, 1906; braima Monard, 1939.	South Africa, KwaZulu-Natal Prov., near Durban.	SW Arabian Peninsula including Yemen; most of subsaharan Africa except the central forested region; Morocco; Zanzibar and Pemba.	Not listed.	Least Concern	 bicolor species group. Includes tephrus; see Hayman and Hill (1971). Reviewed in part by Harrison and Bates (1991) and Horácek et al. (2000).See Taylor (2000a) for distribution map. Subspecies limits are somewhat unclear, and it is possible that this complex includes more than onespecies.	Hipposideros caffer	1004583	23	Sundevall's Leaf-nosed Bat	Cape Leaf-nosed Bat|Common African Leaf-nosed Bat|Lesser Leaf-nosed bat|Sundevall's Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Hipposideros	NA	caffer	Sundevall	1846	1	Rhinolophus_caffer	Sundevall, C. J. (1846). Nya Mammalia frÃ¥ n Sydafrika. Öfversigt af Kongl. Vetenskaps-akademiens forhandlingar, 3(4), 118.	https://www.biodiversitylibrary.org/item/20230#page/144/mode/1up	BM 1848.6.2.16, BM 1849.11.22.11 [syntypes]		near Durban, KwaZulu-Natal Province, South Africa.			caffer (Sundevall, 1846)|caffra (W. Peters, 1852)|gracilis (W. Peters, 1852)|patellifera (W. Peters, 1852)|bicornis (Heuglin, 1861)|angolensis (Seabra, 1898)|nanus J. A. Allen, 1917|aurantiaca De Beaux, 1924|braima Monard, 1938	previously included H. tephrus	Vallo, P., Guillén-Servent, A., Benda, P., Pires, D. B., & Koubek, P. (2008). Variation of mitochondrial DNA in the Hipposideros caffer complex (Chiroptera: Hipposideridae) and its taxonomic implications. Acta Chiropterologica, 10(2), 193-206.				Democratic Republic of the Congo|Angola|Zambia|Malawi|Mozambique|Namibia|Botswana|Zimbabwe|South Africa|Eswatini	Africa	Afrotropic|Palearctic	LC	0	0	0	Hipposideros_caffer	0	sciname match	Hipposideros_caffer	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hipposideros_caffer	1004583	23	Sundevall's Roundleaf Bat	Cape Leaf-nosed Bat|Common African Leaf-nosed Bat|Lesser Leaf-nosed bat|Sundevall's Roundleaf Bat|Sundevall's Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Hipposideros	NA	caffer	Sundevall	1	Rhinolophus caffer	Sundevall, C.J. 1847. Nya Mammalia frÃ¥n Sydafrika. Öfversigt af Kongl. Vetenskaps-Akademiens Förhandlingar 3:118-121.	https://www.biodiversitylibrary.org/page/2581775	BMNH:Mamm:1848.6.2.16, BMNH:Mamm:1849.11.22.11, RMNH.MAM.27330	syntypes	https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.27330 | https://data.nhm.ac.uk/object/f46a73e9-5e3e-4b28-bf21-3f8ef00a46f5	near Durban, KwaZulu-Natal Province, South Africa.			previously included H. tephrus	Vallo, P., Guillén-Servent, A., Benda, P., Pires, D. B., & Koubek, P. (2008). Variation of mitochondrial DNA in the Hipposideros caffer complex (Chiroptera: Hipposideridae) and its taxonomic implications. Acta Chiropterologica, 10(2), 193-206.				Democratic Republic of the Congo|Angola|Zambia|Malawi|Mozambique|Namibia|Botswana|Zimbabwe|South Africa|Eswatini	Africa	Afrotropic|Palearctic	LC	0	0	0	Hipposideros_caffer	0	sciname match	Hipposideros_caffer	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Hipposideros		caffer	Sundevall	1846	1	&Ouml;fv. Kongl. Svenska Vet.-Akad. Forhandl. Stockholm	3(4): 118	Sundevall's Leaf-nosed Bat	aurantiaca De Beaux, 1924; bicornis Heuglin, 1861; gracilis Peters, 1825; angolensis Seabra, 1898; nanus J. A. Allen, 1917; tephrus Cabrera, 1906; braima Monard, 1939.	South Africa, KwaZulu-Natal Prov., near Durban.	SW Arabian Peninsula including Yemen; most of subsaharan Africa except the central forested region; Morocco; Zanzibar and Pemba.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/80459007/22094271/' target='_blank'>Least Concern</a>	bicolorspecies group. Does not include tephrus; but see Hayman and Hill (1971). Reviewed in part by Harrison and Bates (1991) and Horácek et al. (2000).See Taylor (2000a) for distribution map. Subspecies limits are somewhat unclear, and it is possible that this complex includes more than onespecies.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hipposideros caffer; Hipposideros caffer; Hipposideros caffer; Hipposideros caffer; Hipposideros caffer; Hipposideros caffer; caffer; angolensis; nanus; tephrus; aurantiaca; bicornis; gracilis; tephrus - braima; angolensis; nanus; tephrus; aurantiaca; bicornis; gracilis; tephrus - braima; caffercaffra; gracilis; patellifera; bicornis; angolensis; nanus; aurantiaca; braima; Phyllorhine de Cafrerie; Gewöhnliche Rundblattnase; Hiposidérido de Cafreria; Cape Leaf-nosed Bat; Common African Leaf-nosed Bat; Lesser Leaf-nosed bat; Sundevall’s Roundleaf Bat; Sundevall's Leaf-nosed Bat; Cape Leaf-nosed Bat; Common African Leaf-nosed Bat; Lesser Leaf-nosed bat; Sundevall's Roundleaf Bat; Sundevall's Leaf-nosed Bat; Sundevall's Leaf-nosed Bat; H. caffer