http://www.w3.org/ns/prov#wasDerivedFrom	http://purl.org/dc/elements/1.1/format	name_CH1_1980	name_MSW1_1982	name_CH3_1991	name_MSW2_1993	name_Koopman_1994	name_MSW3_2005	name_HMW_2019	name_BatNames_2022	name_MDD_2022	name_IUCN_2022	name_BatNames_2023	name_MDD_2023	name_MDD_2025_2.0	name_batnames_2025_1.7	name_MDD_2025_2.2	column151	taxonomic_notes_concatenated	column171	synonyms_CH1	subspecies__MSW2	synonyms__MSW1	synonyms_CH3	synonyms_MSW2	subspecies_Koopman94_interpreted	subspecies_MSW3_interpreted	synonym_MSW3_interpreted	subspecies_HMW_interpreted	synonym_HMW_interpreted	subspecies_batnames_interpreted	synonym_batnames_interpreted	synonym_MDD_interpreted	synonym_IUCN_interpreted	subspecies_batnames2023_interpreted	synonym_batnames2023_interpreted	synonym_MDD2023_interpreted	synonym_MDD2025_interpreted	subspecies_batnames2025_interpreted	synonyms_batnames2025_interpreted	nominalNames	column391	docOrigin_CH1	commonName_CH1	distribution_CH1	docOrigin_MSW1	column451	typeLocality_MSW1	authority_MSW1	year_MSW1	citation_MSW1	distribution	comment_MSW1	docOrigin_CH3	commonName_CH3	distribution_CH3	docOrigin_MSW2	authority_MSW2	year_MSW2	citation_MSW2	comments_MSW2	distribution_MSW2	typeLocality_MSW2	docOrigin_Koopman94	authority_Koopman94	year_Koopman94	description_Koopman94	distribution_Koopman94	diversity_Koopman94	subspecies_Koopman94	page	rank	name	authority	year	parent	parent_rank	corrected_name	actual_species_count	claimed_species_count	dental_formula	description	diversity	full_subspecies_text	name_line	species_index	subspecies	synonym	text	docOrigin_MSW3	order_MSW3	family_MSW3	subfamily_MSW3	tribe_MSW3	name_MSW3	genus_MSW3	subgenus_MSW3	species_MSW3	authoritySpeciesAuthor_MSW3	(parentheses (1=author & date in parentheses)_MSW3	authoritySpeciesYear_MSW3	actualDate_MSW3	citation_MSW3	volume_MSW3	issue_MSW3	pages_MSW3	type_species_MSW3	commonName_MSW3	typeLocality_MSW3	distribution_MSW3	status_MSW3	synonym_MSW3	comments_MSW3	docId_HMW	docOrigin_HMW	docISBN_HMW	docName_HMW	docMasterId_HMW	docPageNumber_HMW	derivedFrom_HMW	name_HMW	family_HMW	genus_HMW	species_HMW	authoritySpeciesAuthor_HMW	authoritySpeciesYear	commonNames_HMW	taxonomy_HMW	subspeciesAndDistribution_HMW	descriptiveNotes_HMW	habitat_HMW	foodAndFeeding_HMW	breeding_HMW	activityPatterns_HMW	movementsHomeRangeAndSocialOrganization_HMW	statusAndConservation_HMW	bibliography_HMW	distributionImageURL_HMW	verbatimText_HMW	docOrigin_batnames	family_batnames	name_batnames	genus_batnames	subgenus_batnames	species_batnames	authoritySpeciesAuthor_batnames	date_batnames	parentheses_batnames (1=author & date in parentheses)	citation_batnames	docPageNumber_batnames	common Name_batnames	synonyms_batnames	type_locality_batnames	Distribution_batnames	CITES_batnames	IUCN_batnames	comments_batnames	docOrigin_MDD	name_MDD	phylosort_MDD	mainCommonName_MDD	otherCommonNames_MDD	subclass_MDD	infraclass_MDD	magnorder_MDD	superorder_MDD	order_MDD	suborder_MDD	infraorder_MDD	parvorder_MDD	superfamily_MDD	family_MDD	subfamily_MDD	tribe_MDD	genus_MDD	subgenus_MDD	specificEpithet_MDD	authoritySpeciesAuthor_MDD	authoritySpeciesYear_MDD	authorityParentheses_MDD	originalNameCombination_MDD	authoritySpeciesCitation_MDD	authoritySpeciesLink_MDD	holotypeVoucher_MDD	holotypeVoucherURIs_MDD	typeLocality_MDD	typeLocalityLatitude_MDD	typeLocalityLongitude_MDD	nominalNames_MDD	taxonomyNotes_MDD	taxonomyNotesCitation_MDD	countryDistribution_MDD	continentDistribution_MDD	biogeographicRealm_MDD	iucnStatus_MDD	extinct_MDD	domestic_MDD	flagged_MDD	CMW_sciName_MDD	diffSinceCMW_MDD	MSW3_matchtype_MDD	MSW3_sciName_MDD	diffSinceMSW3_MDD	docOrigin_IUCN	internalTaxonId_IUCN	NAME_IUCN	kingdomName_IUCN	phylumName_IUCN	className_IUCN	orderName_IUCN	familyName_IUCN	genusName_IUCN	speciesName_IUCN	authoritySpeciesAuthorYear_IUCN	taxonomicNotes_IUCN	assessmentId_IUCN	scientificName_IUCN	redlistCategory_IUCN	redlistCriteria_IUCN	yearPublished_IUCN	assessmentDate_IUCN	criteriaVersion_IUCN	language_IUCN	rationale_IUCN	habitat_IUCN	threats_IUCN	population_IUCN	populationTrend_IUCN	range_IUCN	useTrade_IUCN	systems_IUCN	conservationActions_IUCN	realm_IUCN	yearLastSeen_IUCN	possiblyExtinct_IUCN	possiblyExtinctInTheWild_IUCN	scopes_IUCN	docOrigin_batnames2023	FAMILY_batnames2023	GENUS_batnames2023	SUBGENUS_batnames2023	SPECIES_batnames2023	authoritySpeciesAuthor_batnames2023	authoritySpeciesYearbatnames2023	PARENTHESES_batnames2023 (1=AUTHOR & DATE IN PARENTHESES)	CITATION_batnames2023	PAGES_batnames2023	COMMON NAME_batnames2023	SYNONYMS_batnames2023	TYPE LOCALITY_batnames2023	DISTRIBUTION_batnames2023	CITES_batnames2023	IUCN_batnames2023	COMMENTS_batnames2023	name MDD2023	id_MDD2023	phylosort_MDD2023	mainCommonName_MDD2023	otherCommonNames_MDD2023	subclass_MDD2023	infraclass_MDD2023	magnorder_MDD2023	superorder_MDD2023	order_MDD2023	suborder_MDD2023	infraorder_MDD2023	parvorder_MDD2023	superfamily_MDD2023	Family_mdd2023	subfamily_MDD2023	tribe_MDD2023	genus_MDD2023	subgenus_MDD2023	specificEpithet_MDD2023	authoritySpeciesAuthor_MDD2023	authoritySpeciesYear_MDD2023	authorityParentheses_MDD2023	originalNameCombination_MDD2023	authoritySpeciesCitation_MDD2023	authoritySpeciesLink_MDD2023	holotypeVoucher_MDD2023	holotypeVoucherURIs_MDD2023	typeLocality_MDD2023	typeLocalityLatitude_MDD2023	typeLocalityLongitude_MDD2023	nominalNames_MDD2023	taxonomyNotes_MDD2023	taxonomyNotesCitation_MDD2023	distributionNotes_MDD2023	distributionNotesCitation_MDD2023	subregionDistribution_MDD2023	countryDistribution_MDD2023	continentDistribution_MDD2023	biogeographicRealm_MDD2023	iucnStatus_MDD2023	extinct_MDD2023	domestic_MDD2023	flagged_MDD2023	CMW_sciName_MDD2023	diffSinceCMW_MDD2023	MSW3_matchtype_MDD2023	MSW3_sciName_MDD2023	diffSinceMSW3_MDD2023	docOrigin_MDD2025	sciName	id	phylosort	mainCommonName	otherCommonNames	subclass	infraclass	magnorder	superorder	order	suborder	infraorder	parvorder	superfamily	family	subfamily	tribe	genus	subgenus	specificEpithet	authoritySpeciesAuthor	authorityParentheses	originalNameCombination	authoritySpeciesCitation	authoritySpeciesLink	typeVoucher	typeKind	typeVoucherURIs	typeLocality	typeLocalityLatitude	typeLocalityLongitude	taxonomyNotes	taxonomyNotesCitation	distributionNotes	distributionNotesCitation	subregionDistribution	countryDistribution	continentDistribution	biogeographicRealm	iucnStatus	extinct	domestic	flagged	CMW_sciName	diffSinceCMW	MSW3_matchtype	MSW3_sciName	diffSinceMSW3	docOrigin_batnames2025	Family	Genus	Subgenus	Species	Author	Date	Parentheses (1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L368	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Hipposideros armiger	Hipposideros armiger	Hipposideros armiger	Hipposideros armiger	Hipposideros armiger	Hipposideros armiger	Hipposideros armiger	Hipposideros armiger	Hipposideros armiger	Hipposideros armiger	Hipposideros armiger	Hipposideros armiger	Hipposideros armiger	Hipposideros armiger	Hipposideros armiger		[MSW2] Includes terasensis, but see Yoshiyuki (1991).; [MSW3] armiger species group. Includes terasensis, but see Yoshiyuki (1991a) and Pavlinov et al. (1995b). Reviewed in part by Kock (1996) Bates and Harrison (1997), Sinha (1999), and Hendrichsen et al. (2001b).; [HMW] Rhinolphus [sic] armiger Hodgson, 1835 , “Nipal [= Nepal ].” Hipposideros armiger is in the armiger species group. Four subspecies recognized.; [batnames2022]  armiger species group. Includes terasensis, but see Yoshiyuki (1991a) and Pavlinov et al. (1995b). Reviewed in part by Kock (1996) Bates and Harrison (1997), Sinha (1999), and Hendrichsen et al. (2001b).; [MDD2022] previously included the name swinhoei as a synonym, but it has been shown to represent H. pratti and is the oldest name available for that species; [batnames2023]  armiger species group. Includes terasensis, but see Yoshiyuki (1991a) and Pavlinov et al. (1995b). Reviewed in part by Kock (1996) Bates and Harrison (1997), Sinha (1999), and Hendrichsen et al. (2001b).; [MDD2023] previously included the name swinhoei as a synonym, but it has been shown to represent H. pratti and is the oldest name available for that species; [MDD2025_2.0] previously included the name swinhoei as a synonym, but it has been shown to represent H. pratti and is the oldest name available for that species; [batnames2025_1.7] armiger species group. Includes terasensis, but see Yoshiyuki (1991a) and Pavlinov et al. (1995b). Reviewed in part by Kock (1996) Bates and Harrison (1997), Sinha (1999), and Hendrichsen et al. (2001b).; [MDD2025_2.2] previously included the name swinhoei as a synonym, but it has been shown to represent H. pratti and is the oldest name available for that species						debilis, swinhoei, terasensis, tranninhensis.	armiger, terasensis, tranninhensis	armiger, fujianensis, terasensis, tranninhensis	debilis, swinhoei	armiger, fujianensis, terasensis, traninhensis		armiger, fujianensis, terasensis, tranninhensis	armiger - debilis, swinhoei	armiger, debilis, terasensis, tranninhensis, fujianensis		armiger, fujianensis, terasensis, tranninhensis	armiger - debilis	armiger, debilis, terasensis, tranninhensis, fujianensis	armiger, armigera, debilis, terasensis, tranninhensis, fujianensis, traninhensis	armiger, fujianensis, terasensis, tranninhensis	armiger - debilis	armiger (B. H. Hodgson, 1835)|armigera (W. C. H. Peters, 1871) [incorrect subsequent spelling]|debilis Andersen, 1906|typicus Andersen, 1906 [nomen novum]|terasensis Kishida, 1924|tranninhensis Bourret, 1942|fujianensis Zhen Xiuyun, 1987|traninhensis D. E. Wilson & Mittermeier, 2019 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Himalayan leaf-nosed bat N India – Malaya, Taiwan	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Hipposideros armiger	Nepal.	Hodgson	1835	J. Asiat. Soc. Bengal, 4:699.	Distribution: The species ranges from northern In dia and southern China south to Malaya.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Himalayan leaf-nosed bat	N India – Malaya, Taiwan	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Hodgson	1835	J. Asiat. Soc. Bengal, 4:699.	Includes terasensis, but see Yoshiyuki (1991).	N India, Nepal, Burma, S China, Vietnam, Laos, Thailand, Malay Peninsula, Taiwan.	Nepal.		HODGSON	1835	Ears pointed and triangular, with a small pro jection at the antitragal fold, but separate. Peri otics relatively small, the width of each approxi mately equal to their distance apart. Upper in cisors strong with two subequal lobes. Crown area of outer lower incisors much greater than that of inner lower incicisors. Noseleaf with three large and one very small lateral leaflet. Frontal depres sion absent. Maxillae not elongated. Posterior noseleaf narrower than anterior noseleaf with a slightly lobate border. Rostrum flattened, its eminences not inflated. A distinct discontinuity between roofs of narial and mesopterygoid ca nals. Size large (forearm length, 79-100 mm).	Distribution: The species ranges from northern In dia and southern China south to Malaya.	Three subspecies are currently recognized:	H. a. armiger (entire range except Taiwan and Indo-China), H. a. terasensis (Taiwan), H. a. tranninhensis (Vietnam, Laos).	65	species	H. armiger	HODGSON	1835	Hipposideros	genus	Hipposideros armiger				Ears pointed and triangular, with a small pro jection at the antitragal fold, but separate. Peri otics relatively small, the width of each approxi mately equal to their distance apart. Upper in cisors strong with two subequal lobes. Crown area of outer lower incisors much greater than that of inner lower incicisors. Noseleaf with three large and one very small lateral leaflet. Frontal depres sion absent. Maxillae not elongated. Posterior noseleaf narrower than anterior noseleaf with a slightly lobate border. Rostrum flattened, its eminences not inflated. A distinct discontinuity between roofs of narial and mesopterygoid ca nals. Size large (forearm length, 79-100 mm).	Three subspecies are currently recognized:		42. H. armiger (HODGSON 1835) [armiger group].	42	_H. a. armiger_ (Hodgson, 1835) (synonyms: _debilis_ Andersen, 1906, _typicus_ Andersen, 1906); _H. a. fujianensis_ Zhen, 1987; _H. a. terasensis_ Kishida, 1924; _H. a. tranninhensis_ Bourret, 1942			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Hipposideros armiger	Hipposideros		armiger	Hodgson	y	1835		J. Asiat. Soc. Bengal	4		699		Great Leaf-nosed Bat	Nepal.	N India, Nepal, Burma, S and SE China, Vietnam, Laos, Cambodia, Thailand, Malay Peninsula, Taiwan.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	debilis K. Andersen, 1906; swinhoei Peters, 1871; fujianensis Zhen, 1987; terasensis Kishida, 1924; tranninhensis Bourret, 1942.	armiger species group. Includes terasensis, but see Yoshiyuki (1991a) and Pavlinov et al. (1995b). Reviewed in part by Kock (1996) Bates and Harrison (1997), Sinha (1999), and Hendrichsen et al. (2001b).	03BD87A2C67CA20FF899F1F9FC2A4A21	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	237	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C67CA20FF899F1F9FC2A4A21.xml	Hipposideros armiger	Hipposideridae	Hipposideros	armiger	Hodgson	1835	Phyllorhine de @fr | 'Himalaya @en | Grosse Himalaya-Rundblattnase @de | Hiposidérido de Himalaya @es | Great Leaf-nosed Bat @en | Great Roundleaf Bat @en	Rhinolphus [sic] armiger Hodgson, 1835 , “Nipal [= Nepal ].” Hipposideros armiger is in the armiger species group. Four subspecies recognized.	H. a. armiger Hodgson, 1835 -N India , Nepal , Myanmar , C & S China (including Hainan I), and mainland SE Asia except Vietnam . H. a. fujianensis Zhen, 1987 -SE China ( Fujian ). H. a. terasensis Kishida, 1924 — Taiwan I. On following pages: 28. Griffin's Leaf-nosed Bat ( Hipposideros griffin/}; 29. Pendlebury's Leaf-nosed Bat ( Hipposideros grandis }; 32. Horsfield's Leaf-nosed Bat ( Hipposideros larvatus }; 33. Maduran Leaf-nosed Bat ( Hipposideros madurae sumbae }; 36. Shield-faced Leaf-nosed Bat ( Hipposideros lylei }; 37. Pratt's Leaf-nosed Bat ( Hipposideros pratti }; 38. 40. Diadem Leaf-nosed Bat ( Hipposideros diadema }; 41. Fierce Leaf-nosed Bat ( Hipposideros dinops }; 42. Crested Leaf-nosed Bat ( Hipposideros lankadiva }; 45. Large Asian Leaf-nosed Bat ( Hipposideros lekaguli }; 46. Peleng Leaf-nosed pendleburyi ); 30. Lesser Leaf-nosed Bat ( Hipposideros turpis }; 31. Grand Leaf-nosed Bat ( Hipposideros }; 34. Sorensen's Leaf-nosed Bat ( Hipposideros sorensenl); 35. Sumban Leaf-nosed Bat ( Hipposideros Shield-nosed Leaf-nosed Bat ( Hipposideros scutinares }; 39. Makira Leaf-nosed Bat ( Hipposideros demissus }; Leaf-nosed Bat ( Hipposideros inexpectatus }; 43. Arnhem Leaf-nosed Bat ( Hipposideros inornatus }; 44. Indian Bat ( Hipposideros pelingensis }. H. a. traninhensis Bourret, 1942 — Vietnam .	Head—body 80— 110 mm , tail 48-70 mm, ear 30-35 mm, hindfoot 13-17 mm, forearm 85—103 mm ; weight 44-67 g. Males are usually larger than females. Ears of the Great Himalayan Leaf-nosed Bat are large and have a pointed tip. Hair is long and soft, and dark brown. Tail is long, with tip free from interfemoral membrane. Noseleaf is rather small and does not cover muzzle. There are four supplementary leaflets, outer one being very small. Intermediate leaf is thick and swollen. Posterior leaf is thick but narrower than anterior leaf. Adult males have a thick, swollen structure behind posterior leaf. Skull is large with well-developed sagittal crest. Rostrum and supraorbital region are gready inflated. C1 and j are heavily built. P2 is small and extruded from tooth row, so that C1 and P4 are in contact. Chromosomal complement has 2n = 32 and FN = 60.	The Great Himalayan Leaf-nosed Bat forages in primary and disturbed forests, seeking its food in the forest itself or in other vegetation near caves. It is also recorded in orchards and rubber plantations.	The Great Himalayan Leaf-nosed Bat feeds on insects, mostly o leoptera and Hymenoptera, in forest understory and around trees. It sometimes flies high and fast in open spaces, perhaps while commuting to and from foraging sites.	Females were found pregnant from January to early May. Young Great Himalayan Leaf-nosed Bats were seen attached to females in the roost from February to June.	The Great Himalayan Leaf-nosed Bat often roosts in large numbers in caves. Its echolocation calls are typical F components terminating with a FM tail, with the frequency of the F part of 65—75 kHz.	The Great Himalayan Leaf-nosed Bat forms small to very large colonies in caves. Each individual usually has a roosting space of c. 10-20 cm from other individuals. It is often found in mixed colonies with other large Hipposideros species , such as Diadem Leaf-nosed Bats ( H. diadema ), Large Asian Leaf-nosed Bats ( H. lekaguli ), or Shield-feced Leaf-nosed Bats ( H. lylei ).	Classified as Least Concern on The IUCNRed List. The Great Himalayan Leaf-nosed Bat is widespread and rather common in cave habitats. It has been recorded from several protected areas throughout its range.	Bates, Bumrungsri, Francis & Csorba (2008) | Corbet & Hill (1992) | Francis (2008a) | Hughes et al. (2010) | Lekagul & McNeely (1988) | Simmons (2005)	https://zenodo.org/record/6470379/files/figure.png	27. Great Himalayan Leaf-nosed Bat Hipposideros armiger French: Phyllorhine de I'Himalaya I German: Grosse Himalaya-Rundblattnase / Spanish: Hiposidérido de Himalaya Other common names: Great Leaf-nosed Bat , Great Roundleaf Bat Taxonomy. Rhinolphus [sic] armiger Hodgson, 1835 , “Nipal [= Nepal ].” Hipposideros armiger is in the armiger species group. Four subspecies recognized. Subspecies and Distribution. H. a. armiger Hodgson, 1835 -N India , Nepal , Myanmar , C & S China (including Hainan I), and mainland SE Asia except Vietnam . H. a. fujianensis Zhen, 1987 -SE China ( Fujian ). H. a. terasensis Kishida, 1924 — Taiwan I. On following pages: 28. Griffin's Leaf-nosed Bat ( Hipposideros griffin/}; 29. Pendlebury's Leaf-nosed Bat ( Hipposideros grandis }; 32. Horsfield's Leaf-nosed Bat ( Hipposideros larvatus }; 33. Maduran Leaf-nosed Bat ( Hipposideros madurae sumbae }; 36. Shield-faced Leaf-nosed Bat ( Hipposideros lylei }; 37. Pratt's Leaf-nosed Bat ( Hipposideros pratti }; 38. 40. Diadem Leaf-nosed Bat ( Hipposideros diadema }; 41. Fierce Leaf-nosed Bat ( Hipposideros dinops }; 42. Crested Leaf-nosed Bat ( Hipposideros lankadiva }; 45. Large Asian Leaf-nosed Bat ( Hipposideros lekaguli }; 46. Peleng Leaf-nosed pendleburyi ); 30. Lesser Leaf-nosed Bat ( Hipposideros turpis }; 31. Grand Leaf-nosed Bat ( Hipposideros }; 34. Sorensen's Leaf-nosed Bat ( Hipposideros sorensenl); 35. Sumban Leaf-nosed Bat ( Hipposideros Shield-nosed Leaf-nosed Bat ( Hipposideros scutinares }; 39. Makira Leaf-nosed Bat ( Hipposideros demissus }; Leaf-nosed Bat ( Hipposideros inexpectatus }; 43. Arnhem Leaf-nosed Bat ( Hipposideros inornatus }; 44. Indian Bat ( Hipposideros pelingensis }. H. a. traninhensis Bourret, 1942 — Vietnam . Descriptive notes. Head—body 80— 110 mm , tail 48-70 mm, ear 30-35 mm, hindfoot 13-17 mm, forearm 85—103 mm ; weight 44-67 g. Males are usually larger than females. Ears of the Great Himalayan Leaf-nosed Bat are large and have a pointed tip. Hair is long and soft, and dark brown. Tail is long, with tip free from interfemoral membrane. Noseleaf is rather small and does not cover muzzle. There are four supplementary leaflets, outer one being very small. Intermediate leaf is thick and swollen. Posterior leaf is thick but narrower than anterior leaf. Adult males have a thick, swollen structure behind posterior leaf. Skull is large with well-developed sagittal crest. Rostrum and supraorbital region are gready inflated. C1 and j are heavily built. P2 is small and extruded from tooth row, so that C1 and P4 are in contact. Chromosomal complement has 2n = 32 and FN = 60. Habitat. The Great Himalayan Leaf-nosed Bat forages in primary and disturbed forests, seeking its food in the forest itself or in other vegetation near caves. It is also recorded in orchards and rubber plantations. Food and Feeding. The Great Himalayan Leaf-nosed Bat feeds on insects, mostly o leoptera and Hymenoptera, in forest understory and around trees. It sometimes flies high and fast in open spaces, perhaps while commuting to and from foraging sites. Breeding. Females were found pregnant from January to early May. Young Great Himalayan Leaf-nosed Bats were seen attached to females in the roost from February to June. Activity patterns. The Great Himalayan Leaf-nosed Bat often roosts in large numbers in caves. Its echolocation calls are typical F components terminating with a FM tail, with the frequency of the F part of 65—75 kHz. Movements, Home range and Social organization. The Great Himalayan Leaf-nosed Bat forms small to very large colonies in caves. Each individual usually has a roosting space of c. 10-20 cm from other individuals. It is often found in mixed colonies with other large Hipposideros species , such as Diadem Leaf-nosed Bats ( H. diadema ), Large Asian Leaf-nosed Bats ( H. lekaguli ), or Shield-feced Leaf-nosed Bats ( H. lylei ). Status and Conservation. Classified as Least Concern on The IUCNRed List. The Great Himalayan Leaf-nosed Bat is widespread and rather common in cave habitats. It has been recorded from several protected areas throughout its range. Bibliography. Bates, Bumrungsri, Francis & Csorba (2008), Corbet & Hill (1992), Francis (2008a), Hughes eta/. (2010), Lekagul & McNeely (1988), Simmons (2005).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Hipposideros armiger	Hipposideros		armiger	Hodgson	1835	1	J. Asiat. Soc. Bengal	0.6521	Great Leaf-nosed Bat	 debilis K. Andersen, 1906; swinhoei Peters, 1871; <b> fujianensis </b> Zhen, 1987; <b> terasensis </b> Kishida, 1924; <b> tranninhensis </b> Bourret, 1942.	Nepal.	N India, Nepal, Burma, S China, Vietnam, Laos, Cambodia, Thailand, Malay Peninsula, Taiwan.	Not listed.	Least Concern	 armiger species group. Includes terasensis, but see Yoshiyuki (1991a) and Pavlinov et al. (1995b). Reviewed in part by Kock (1996) Bates and Harrison (1997), Sinha (1999), and Hendrichsen et al. (2001b).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hipposideros armiger	23	Great Himalayan Leaf-nosed Bat	Great Leaf-nosed Bat|Great Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Hipposideros	NA	armiger	Hodgson	1835	1	Rhinolophus_armiger	Hodgson, B. H. (1835). Synopsis of the Vespertilionidae of Nipal. Journal of the Asiatic Society of Bengal, 4, 699.	https://www.biodiversitylibrary.org/item/114405#page/839/mode/1up	BM 1843.1.12.133 [lectotype]		"Nipal [= Nepal]."			armiger (Hodgson, 1835)|debilis K. Andersen, 1906|terasensis Kishida, 1924|tranninhensis Bourret, 1942|fujianensis Zhen Xiuyun, 1987	previously included the name swinhoei as a synonym, but it has been shown to represent H. pratti and is the oldest name available for that species	Kruskop, S. V. (2014). Valid name for the Pratt's leaf-nosed bat, Hipposideros pratti (Hipposideridae, Chiroptera, Mammalia). Russian Journal of Theriology. Ð ÑƒÑÑÐºÐ¸Ð¹ Ñ‚ÐµÑ€Ð¸Ð¾Ð»Ð¾Ð³Ð¸Ñ‡ÐµÑÐºÐ¸Ð¹ Ð¶ÑƒÑ€Ð½Ð°Ð», 13(2), 105-108.	India|Nepal|Myanmar|China|Taiwan|Thailand|Cambodia|Vietnam|Laos|Malaysia	Asia	Indomalaya|Palearctic	LC	0	0	0	Hipposideros_armiger	0	sciname match	Hipposideros_armiger	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10110	Hipposideros armiger	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Hipposideros	armiger	(Hodgson, 1835)		20000000	Hipposideros armiger	Least Concern		2020	2018-09-27 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, it occurs in a number of protected areas, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	In South Asia, this usually low flying species is generally a high altitude species found in montane and bamboo forests (Mistry 1995). It has been recorded roosting either singly or in colonies of several individuals and shares its roosts with other species of bats in subterranean caves, lofts of houses, verandahs of old houses, old temples. It breeds once a year and gives birth to two young (Bates and Harrison 1997). In China, it is considered to be a species found in a variety of habitats. They are known to inhabit caves and a variety of man-made structures. Colonies can number in the hundreds of individuals, and they co-occur with species of Rhinolophus and other bats. In Southeast Asia, the species has been primarily recorded near caves but occurs in quite distant areas from the roosts when foraging.	In South Asia, this species is threatened by deforestation, generally resulting from logging operations and the conversion of land for agricultural purposes, from mining activities, and disturbance to roosting sites in caves (Molur et al. 2002). In Southeast Asia, cave disturbance is occurring throughout the species' range and it is hunted (presumably for food) in Lao PDR, Viet Nam and Thailand.	It appears to be fairly common throughout its range.	Unknown	This widespread species has been recorded from India and Nepal, eastwards into central and southeastern China, and from much of peninsular Southeast Asia. In South Asia, it has been reported from India (Assam, Manipur, Meghalaya, Sikkim, Uttaranchal and West Bengal) and Nepal (Central, Eastern and Western Nepal) in South Asia (Molur et al . 2002). In China, it is distributed south of the Yangtze river, and has been recorded in Jiangxi, Zhejiang, Guangdong, Hong Kong, Macao, Guangxi, Hainan, Hunan, Jiangsu, Anhui, Yunnan, Sichuan, Shaanxi, Guizhou, Fujian and the island of Taiwan. In Southeast Asia, it ranges from Myanmar in the west, through to Thailand and the Malay Peninsula. In South Asia, it has been recorded at elevations of 1,000 to 2,031 m asl (Molur et al. 2002). This species is also found in limestone caves in Mon State, Southern part of Myanmar and the elevation is around 20 m asl. (Myo and Soisook in prep.).		Terrestrial	In South Asia, there are no direct conservation measures in place. The species has been recorded from protected areas in India like Mahanandi Wildlife Sanctuary in West Bengal. In Southeast Asia, it occurs in protected areas throughout its range. In parts of its range further studies are needed into the distribution, abundance, breeding biology and general ecology of this species. Populations of this species should be monitored to record changes in abundance and distribution (Molur et al. 2002).	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Hipposideros		armiger	Hodgson	1835	1	J. Asiat. Soc. Bengal	0.652083	Great Leaf-nosed Bat	 debilis K. Andersen, 1906; swinhoei Peters, 1871; <b> fujianensis </b> Zhen, 1987; <b> terasensis </b> Kishida, 1924; <b> tranninhensis </b> Bourret, 1942.	Nepal.	N India, Nepal, Burma, S China, Vietnam, Laos, Cambodia, Thailand, Malay Peninsula, Taiwan.	Not listed.	Least Concern	 armiger species group. Includes terasensis, but see Yoshiyuki (1991a) and Pavlinov et al. (1995b). Reviewed in part by Kock (1996) Bates and Harrison (1997), Sinha (1999), and Hendrichsen et al. (2001b).	Hipposideros armiger	1004577	23	Great Himalayan Leaf-nosed Bat	Great Leaf-nosed Bat|Great Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Hipposideros	NA	armiger	Hodgson	1835	1	Rhinolophus_armiger	Hodgson, B. H. (1835). Synopsis of the Vespertilionidae of Nipal. Journal of the Asiatic Society of Bengal, 4, 699.	https://www.biodiversitylibrary.org/item/114405#page/839/mode/1up	BM 1843.1.12.133 [lectotype]		"Nipal [= Nepal]."			armiger (Hodgson, 1835)|debilis K. Andersen, 1906|terasensis Kishida, 1924|tranninhensis Bourret, 1942|fujianensis Zhen Xiuyun, 1987	previously included the name swinhoei as a synonym, but it has been shown to represent H. pratti and is the oldest name available for that species	Kruskop, S. V. (2014). Valid name for the Pratt's leaf-nosed bat, Hipposideros pratti (Hipposideridae, Chiroptera, Mammalia). Russian Journal of Theriology. Ð ÑƒÑÑÐºÐ¸Ð¹ Ñ‚ÐµÑ€Ð¸Ð¾Ð»Ð¾Ð³Ð¸Ñ‡ÐµÑÐºÐ¸Ð¹ Ð¶ÑƒÑ€Ð½Ð°Ð», 13(2), 105-108.				India|Nepal|Myanmar|China|Taiwan|Thailand|Cambodia|Vietnam|Laos|Malaysia	Asia	Indomalaya|Palearctic	LC	0	0	0	Hipposideros_armiger	0	sciname match	Hipposideros_armiger	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hipposideros_armiger	1004577	23	Great Himalayan Roundleaf Bat	Great Leaf-nosed Bat|Great Roundleaf Bat|Great Himalayan Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Hipposideros	NA	armiger	B. H. Hodgson	1	Rhinolophus armiger	Hodgson, B.H. 1835. Synopsis of the VespertilionidÃ¦ of Nipal. Journal of the Asiatic Society of Bengal (2)4:699-701.	https://www.biodiversitylibrary.org/page/37189992	BMNH:Mamm:1843.1.12.132	lectotype	https://data.nhm.ac.uk/object/f7d1dbe7-17a5-4d4d-81f3-e108c2ec2529	"Nipal [= Nepal]."			previously included the name swinhoei as a synonym, but it has been shown to represent H. pratti and is the oldest name available for that species	Kruskop, S. V. (2014). Valid name for the Pratt's leaf-nosed bat, Hipposideros pratti (Hipposideridae, Chiroptera, Mammalia). Russian Journal of Theriology. Ð ÑƒÑÑÐºÐ¸Ð¹ Ñ‚ÐµÑ€Ð¸Ð¾Ð»Ð¾Ð³Ð¸Ñ‡ÐµÑÐºÐ¸Ð¹ Ð¶ÑƒÑ€Ð½Ð°Ð», 13(2), 105-108.				India|Nepal|Myanmar|China|Taiwan|Thailand|Cambodia|Vietnam|Laos|Malaysia	Asia	Indomalaya|Palearctic	LC	0	0	0	Hipposideros_armiger	0	sciname match	Hipposideros_armiger	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Hipposideros		armiger	Hodgson	1835	1	J. Asiat. Soc. Bengal	0.652083	Great Leaf-nosed Bat	debilis K. Andersen, 1906; swinhoei Peters, 1871; fujianensis Zhen, 1987; terasensis Kishida, 1924; tranninhensis Bourret, 1942.	Nepal.	N India, Nepal, Burma, S China, Vietnam, Laos, Cambodia, Thailand, Malay Peninsula, Taiwan.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10110/22097743/' target='_blank'>Least Concern</a>	armiger species group. Includes terasensis, but see Yoshiyuki (1991a) and Pavlinov et al. (1995b). Reviewed in part by Kock (1996) Bates and Harrison (1997), Sinha (1999), and Hendrichsen et al. (2001b).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hipposideros armiger; Hipposideros armiger; Hipposideros armiger; Hipposideros armiger; Hipposideros armiger; Hipposideros armiger; armiger; fujianensis; terasensis; tranninhensis; debilis; swinhoei; armiger; fujianensis; terasensis; traninhensis; fujianensis; terasensis; tranninhensis; debilis; swinhoei; armiger; debilis; terasensis; tranninhensis; fujianensis; Phyllorhine de; 'Himalaya; Grosse Himalaya-Rundblattnase; Hiposidérido de Himalaya; Great Leaf-nosed Bat; Great Roundleaf Bat; Great Himalayan Leaf-nosed Bat; Great Leaf-nosed Bat; Great Roundleaf Bat; Great Leaf-nosed Bat; Great Leaf-nosed Bat; H. armiger
