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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L365	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Hipposideros abae	Hipposideros abae	Hipposideros abae	Hipposideros abae	Hipposideros abae	Hipposideros abae	Hipposideros abae	Hipposideros abae	Hipposideros abae	Hipposideros abae	Hipposideros abae	Hipposideros abae	Hipposideros abae	Hipposideros abae	Hipposideros abae		[MSW3] speoris species group.; [HMW] Hipposideros abae J. A. Allen, 1917 , “Aba, Uele district, Belgian Congo [= DR Congo ].” Hipposideros abae was formerly included in the speoris species group, but is now placed in the ruber species group (7 species). Monotypic.; [batnames2022]  speoris species group.; [batnames2023]  speoris species group.; [batnames2025_1.7] speorisspecies group.														abae				abae 	abae 			abae J. A. Allen in J. A. Allen, Lang, & Chapin, 1917		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Aba leaf-nosed bat	Guinea – S Sudan, Uganda	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Hipposideros abae	Zaire, Oriental, Aba.	J. A. Allen	1917	Bull. Am. Mus. Nat. Hist., 37:432.	Distribution: Occurring in woodlands of northern tropical Africa from Guinea (Bissau) to southern Sudan.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Aba leaf-nosed bat	Guinea – S Sudan, Uganda	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	J. A. Allen	1917	Bull. Am. Mus. Nat. Hist., 37:432.		Guinea-Bissau to SW Sudan and Uganda.	Zaire, Oriental, Aba.		J. A. ALLEN	1917	Ears pointed and triangular, but separate. Peri otics relatively small, the width of each approxi mately equal to their distance apart. Upper in cisors strong but weakly bilobed. Crown area of outer lower incisors much greater than that of in ner lower incisors. Noseleaf with three lateral leaf lets. No definite frontal depression. Maxillae not elongated. Posterior noseleaf equal in width to anterior noseleaf, its border more or less semicir cular, supporting septa absent. Rostrum flattened with prominent supraorbital ridges, its eminences moderately inflated. Roofs of narial and mesop terygoid canals not sharply discontinuous. Ear with a small projection at the antitragal fold. Pterygoid wings of skull undeveloped. Anterior noseleaf without median emargination. Vomer not projecting behind palate. Upper canine with a shallow antero-internal groove and low pos terior cusp. Posterior edge of bony palate nearly square, with prominent posterior palatal foramina. Sphenoidal bridge constricted, not con cealing lateral apertues. Size medium (forearm length, 55-66 mm).	Distribution: Occurring in woodlands of northern tropical Africa from Guinea (Bissau) to southern Sudan.	No sub species.		65	species	H. abae	J. A. ALLEN	1917	Hipposideros	genus	Hipposideros abae				Ears pointed and triangular, but separate. Peri otics relatively small, the width of each approxi mately equal to their distance apart. Upper in cisors strong but weakly bilobed. Crown area of outer lower incisors much greater than that of in ner lower incisors. Noseleaf with three lateral leaf lets. No definite frontal depression. Maxillae not elongated. Posterior noseleaf equal in width to anterior noseleaf, its border more or less semicir cular, supporting septa absent. Rostrum flattened with prominent supraorbital ridges, its eminences moderately inflated. Roofs of narial and mesop terygoid canals not sharply discontinuous. Ear with a small projection at the antitragal fold. Pterygoid wings of skull undeveloped. Anterior noseleaf without median emargination. Vomer not projecting behind palate. Upper canine with a shallow antero-internal groove and low pos terior cusp. Posterior edge of bony palate nearly square, with prominent posterior palatal foramina. Sphenoidal bridge constricted, not con cealing lateral apertues. Size medium (forearm length, 55-66 mm).	No sub species.		44. H. abae J. A. ALLEN 1917 [speoris group],	44	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Hipposideros abae	Hipposideros		abae	J. A. Allen		1917		Bull. Am. Mus. Nat. Hist.	37		432		Aba Leaf-nosed Bat	Dem. Rep. Congo, Oriental, Aba.	Guinea-Bissau to SW Sudan and Uganda.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).		speoris species group.	03BD87A2C662A210FF52F514FBEA516B	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	247	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C662A210FF52F514FBEA516B.xml	Hipposideros abae	Hipposideridae	Hipposideros	abae	J. A. Allen	1917	Phyllorhine d'Aba @fr | Aba-Rundblattnase @de | Hiposidérido de Aba @es | Aba Roundleaf Bat @en	Hipposideros abae J. A. Allen, 1917 , “Aba, Uele district, Belgian Congo [= DR Congo ].” Hipposideros abae was formerly included in the speoris species group, but is now placed in the ruber species group (7 species). Monotypic.	Patchily in Africa N of the equator from Guinea Bissau and Guinea E to South Sudan and extreme NW Uganda .	Head—body 60-70 mm, tail 28—40 mm , ear 18-24 mm, hindfoot 7—13 mm , forearm 54—65 mm ; weight 17— 19 g . The Aba Leaf-nosed Bat has short, rounded wings. Muzzle is relatively short with a large but simple noseleaf that has three lateral leaflets. A frontal sac is present in males only. Intemarial septum is not swollen and does not partially cover the nares. Ears are separate, and relatively small. Fur is fine and short, dark brown dorsally, paler ventrally; an orange morph exists, which is bright orange or rusty brown dorsally and paler ventrally.	Inhabits Guinea savanna, including derived savanna, dry forest, and recently cleared rainforest. Apparently avoids intact rainforest.	No information.	Births of single young have been reported in March throughout its range.	The Aba Leaf-nosed Bat roosts during the day in caves, under boulders, and in burrows of mammals.	The Aba Leaf-nosed Bat roosts in small to large colonies of up to several hundred individuals. In one study, sex ratio was 1:2 with females doubling males. The species may share its roost with a number of other Hipposideridae and Rhinolophus .	Classified as Least Concern on The IUCN Red List. The Aba Leaf-nosed Bat is widespread and relatively common with no major known conservation threats.	Decher (1997) | Grubb et al. (1998) | Happold, D.C.D. (1987) | Happold, M. (2013u) | Hill (1963a) | Koch-Weser (1984) | Rosevear (1965) | Schütter et al. (1982)	https://zenodo.org/record/3749145/files/figure.png	55. Aba Leaf-nosed Bat Hipposideros abae French: Phyllorhine d'Aba / German: Aba-Rundblattnase / Spanish: Hiposidérido de Aba Other common names: Aba Roundleaf Bat Taxonomy. Hipposideros abae J. A. Allen, 1917 , “Aba, Uele district, Belgian Congo [= DR Congo ].” Hipposideros abae was formerly included in the speoris species group, but is now placed in the ruber species group (7 species). Monotypic. Distribution. Patchily in Africa N of the equator from Guinea Bissau and Guinea E to South Sudan and extreme NW Uganda . Descriptive notes. Head—body 60-70 mm, tail 28—40 mm , ear 18-24 mm, hindfoot 7—13 mm , forearm 54—65 mm ; weight 17— 19 g . The Aba Leaf-nosed Bat has short, rounded wings. Muzzle is relatively short with a large but simple noseleaf that has three lateral leaflets. A frontal sac is present in males only. Intemarial septum is not swollen and does not partially cover the nares. Ears are separate, and relatively small. Fur is fine and short, dark brown dorsally, paler ventrally; an orange morph exists, which is bright orange or rusty brown dorsally and paler ventrally. Habitat. Inhabits Guinea savanna, including derived savanna, dry forest, and recently cleared rainforest. Apparently avoids intact rainforest. Food and Feeding. No information. Breeding. Births of single young have been reported in March throughout its range. Activity patterns. The Aba Leaf-nosed Bat roosts during the day in caves, under boulders, and in burrows of mammals. Movements, Home range and Social organization. The Aba Leaf-nosed Bat roosts in small to large colonies of up to several hundred individuals. In one study, sex ratio was 1:2 with females doubling males. The species may share its roost with a number of other Hipposideridae and Rhinolophus . Status and Conservation. Classified as Least Concern on The IUCN Red List. The Aba Leaf-nosed Bat is widespread and relatively common with no major known conservation threats. Bibliography. Decher (1997), Grubb eta/. (1998), Happold, D.C.D. (1987), Happold, M. (2013u), Hill (1963a), Koch-Weser (1984), Rosevear (1965), Schütter eta/. (1982).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Hipposideros abae	Hipposideros		abae	J. A. Allen	1917	0	Bull. Am. Mus. Nat. Hist.	44:12:00	Aba Leaf-nosed Bat	None.	Dem. Rep. Congo, Oriental, Aba.	Guinea-Bissau to SW Sudan and Uganda.	Not listed.	Least Concern	 speoris species group.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hipposideros abae	23	Aba Leaf-nosed Bat	Aba Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Hipposideros	NA	abae	J. A. Allen	1917	0	Hipposideros_abae	Allen, J. A. (1917). Part 1. Systematic List. In J. A. Allen, H. Lang & J. P. Chapin. The American Museum Congo Expedition Collection of Bats. Bulletin of the American Museum of Natural History, 37, 432.	http://digitallibrary.amnh.org/bitstream/handle/2246/1068/B37a18.pdf?sequence=1&isAllowed=y	AMNH 49123		"Aba, Uele district, Belgian Congo [= DR Congo]."			abae J. A. Allen, 1917	NA	NA	Guinea-Bissau|Guinea|Sierra Leone|Liberia|Côte d'Ivoire|Burkina Faso|Ghana|Togo|Mali?|Benin?|Nigeria|Cameroon|Chad?|Central African Republic|South Sudan|Democratic Republic of the Congo|Uganda	Africa	Afrotropic	LC	0	0	0	Hipposideros_abae	0	sciname match	Hipposideros_abae	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10109	Hipposideros abae	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Hipposideros	abae	J.A. Allen, 1917		20000000	Hipposideros abae	Least Concern		2017	2016-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, tolerance of a degree of habitat modification, presumed large population, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	It is found in the Guinea savanna zone extending southwards into derived savanna and areas of destroyed rainforest (Koopman, Mumford and Heisterberg 1978; Happold 1987). It does not occur in closed rainforest. It roosts in caves and rocky crevices or on the undersides of boulders, but apparently not in houses (Happold 1987).	There are no known major threats to this species. It appears to have taken advantage of forest clearance by moving into formerly forested land.	There is little information. The Aba roundleaf bat is thought to be sparsely distributed throughout its range, and to roost in small colonies (approximate group size in the dozens, not hundreds).	Unknown	This species is patchily distributed in a range which extends from Guinea-Bissau in west Africa (also the most northerly record) to northern Uganda (also the most eastern and southerly record) (Simmons 2005). It is a lowland species found below 1,000 m. There are records from Guinea-Bissau, Guinea, Sierra Leone, Liberia, Côte d'Ivoire, Burkina Faso, Ghana, Togo, Nigeria, Cameroon, Central African Republic, Sudan, Democratic Republic of Congo and Uganda, and it is to be expected in Mali, Benin and Chad.		Terrestrial	No specific conservation measures are in place, but it occurs in Comoe National Park in Côte d'Ivoire. and presumably also in other protected areas across its wide range.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Hipposideros		abae	J. A. Allen	1917	0	Bull. Am. Mus. Nat. Hist.	44:12:00	Aba Leaf-nosed Bat	None.	Dem. Rep. Congo, Oriental, Aba.	Guinea-Bissau to SW Sudan and Uganda.	Not listed.	Least Concern	 speoris species group.	Hipposideros abae	1004574	23	Aba Leaf-nosed Bat	Aba Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Hipposideros	NA	abae	J. A. Allen	1917	0	Hipposideros_abae	Allen, J. A. (1917). Part 1. Systematic List. In J. A. Allen, H. Lang & J. P. Chapin. The American Museum Congo Expedition Collection of Bats. Bulletin of the American Museum of Natural History, 37, 432.	http://digitallibrary.amnh.org/bitstream/handle/2246/1068/B37a18.pdf?sequence=1&isAllowed=y	AMNH 49123		"Aba, Uele district, Belgian Congo [= DR Congo]."			abae J. A. Allen, 1917	NA	NA				Guinea-Bissau|Guinea|Sierra Leone|Liberia|Côte d'Ivoire|Burkina Faso|Ghana|Togo|Mali?|Benin?|Nigeria|Cameroon|Chad?|Central African Republic|South Sudan|Democratic Republic of the Congo|Uganda	Africa	Afrotropic	LC	0	0	0	Hipposideros_abae	0	sciname match	Hipposideros_abae	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hipposideros_abae	1004574	23	Aba Roundleaf Bat	Aba Roundleaf Bat|Aba Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Hipposideros	NA	abae	J. A. Allen in J. A. Allen, Lang, & Chapin	0	Hipposideros abæ	Allen, J.A., Lang, H. and Chapin, J.P. 1917-09-29. The American Museum Congo Expedition collection of bats. Bulletin of the American Museum of Natural History 37(18):405-563.	https://hdl.handle.net/2246/1068	AMNH M-49123	holotype	http://portal.vertnet.org/o/amnh/mammals?id=urn-catalog-amnh-mammals-m-49123	"Aba, Uele district, Belgian Congo [= DR Congo]."			NA	NA				Guinea-Bissau|Guinea|Sierra Leone|Liberia|Cote d'Ivoire|Burkina Faso|Ghana|Togo|Mali?|Benin?|Nigeria|Cameroon|Chad?|Central African Republic|South Sudan|Democratic Republic of the Congo|Uganda	Africa	Afrotropic	LC	0	0	0	Hipposideros_abae	0	sciname match	Hipposideros_abae	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Hipposideros		abae	J. A. Allen	1917	0	Bull. Am. Mus. Nat. Hist.	44:12:00	Aba Leaf-nosed Bat	None.	Dem. Rep. Congo, Oriental, Aba.	Guinea-Bissau to SW Sudan and Uganda.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10109/22097582/' target='_blank'>Least Concern</a>	speorisspecies group.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hipposideros abae; Hipposideros abae; Hipposideros abae; Hipposideros abae; Hipposideros abae; Hipposideros abae; abae; Phyllorhine d'Aba; Aba-Rundblattnase; Hiposidérido de Aba; Aba Roundleaf Bat; Aba Leaf-nosed Bat; Aba Roundleaf Bat; Aba Leaf-nosed Bat; Aba Leaf-nosed Bat; H. abae