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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L362	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	Hesperoptenus gaskelli	Hesperoptenus gaskelli	Hesperoptenus gaskelli	Hesperoptenus gaskelli	Hesperoptenus gaskelli	Hesperoptenus gaskelli	Hesperoptenus gaskelli	Hesperoptenus gaskelli	Hesperoptenus gaskelli	Hesperoptenus gaskelli	Hesperoptenus gaskelli	Hesperoptenus gaskelli	Hesperoptenus gaskelli		[MSW2] Subgenus Milithronycteris.; [MSW3] Subgenus Milithronycteris.; [HMW] Hesperoptenus (Milithronycteris) gaskelli Hill, 1983 , “ Central Sulawesi [Indoncsial: ... River Ranu, 1°51’ S , 121° 30° E .” Subgenus Milithronycteris . Relationships of H. gaskelli to other Hesperoptenus are unknown because it has not been included in a phylogenetic study. Monotypic.; [batnames2022] Subgenus Milithronycteris .; [IUCN] Hesperoptenus gaskelli is recognized as a species (Simmons and Cirranello 2020).; [batnames2023] Subgenus Milithronycteris .; [batnames2025_1.7] None.														gaskelli	Hesperoptenus gaskelli is recognized as a species (Simmons and Cirranello 2020).			gaskelli 	gaskelli 			gaskelli J. Edwards Hill, 1983						N/A					Distribution: Known only from Celebes.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Sulawesi; ref. 4.12	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Hill	1983	Bull. Brit. Mus. (Nat. Hist.) Zool., 45:169.	Subgenus Milithronycteris.	Sulawesi.	Sulawesi, Central R. Ranu.		HILL	1983	Size medium (forearm length, 38-41 mm). No pad at base of thumb. Outer upper incisor relatively large. Lower incisors considerably imbricated. Fur relatively dark.	Distribution: Known only from Celebes.	No subspecies.		123	species	H. gaskelli	HILL	1983	Milithronycteris	subgenus	Hesperoptenus gaskelli				Size medium (forearm length, 38-41 mm). No pad at base of thumb. Outer upper incisor relatively large. Lower incisors considerably imbricated. Fur relatively dark.	No subspecies.		4. H. gaskelli HILL 1983.	4	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Eptesicini	Hesperoptenus gaskelli	Hesperoptenus	Milithronycteris	gaskelli	Hill		1983		Bull. Brit. Mus. (Nat. Hist.) Zool.	45		169		Gaskell's False Serotine	Indonesia, Sulawesi, Central R. Ranu (01°51'S, 121°30'E).	Sulawesi (known only from the type locality).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.		Subgenus Milithronycteris.	4C3D87E8FFB56A0AFF4C91051C72B6A0	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	836	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFB56A0AFF4C91051C72B6A0.xml	Hesperoptenus gaskelli	Vespertilionidae	Hesperoptenus	gaskelli	Hill	1983	Sérotine de Gaskell @fr | Gaskell-Scheinbreitfligel @de | Hesperopteno de Gaskell @es	Hesperoptenus (Milithronycteris) gaskelli Hill, 1983 , “ Central Sulawesi [Indoncsial: ... River Ranu, 1°51’ S , 121° 30° E .” Subgenus Milithronycteris . Relationships of H. gaskelli to other Hesperoptenus are unknown because it has not been included in a phylogenetic study. Monotypic.	Known only from type locality in EC Sulawesi .	Forearm 40-1 mm (type specimen, juvenile female). Gaskell’s False Serotine resembles Tickell’s False Serotine ( H. tickelli ) and the Large False Serotine ( H. tomesi ) but differs from both in its smaller size and teeth. Color (alcohol specimen) is rich chocolate-brown dorsally and ventrally. Hairs are more brownish at base but otherwise darker and lustrous. Muzzle is low, wide, laterally swollen, and sparsely haired. Internarial region is naked. Ears are thick, fleshy, and subquadrangular. Tragus is rather fleshy and more or less hatched-shaped. Thumb is relatively long, and second phalanx ofthird digit is shorter than first phalanx; upper surface of thumb has thin hairs; upper surface of forearm is naked; upper surface of uropatagium is sparsely haired on proximal one-half; base of tail is similarly furred above and below, with tip protruded beyond uropatagium. Skull is medium-sized, with broad non-inflated braincase; postorbital region is moderately constricted; and supraorbital region is abruptly expanded, with well-developed superior anteorbital tubercles. I? is caniniform and massive, and I’ is more or less ovate. I’ is displaced inward so that I? and C! are in contact. Condylo-canine length is 13-1 mm, and maxillary tooth row is 6 mm .	Holotype was caught in small clearing on a covered (with vegetation) path at the edge of primary forest at 10 m above sea level.	No information.	No information.	Nocturnal. Single specimen was caught in a harp trap with its top 3 m aboveground (bottom of trapping area would then be c.1-1-5 m aboveground).	No information.	Classified as Data Deficient on The IUCN Red List. Gaskell’s False Serotine is known only from one locality in Morowali Nature Reserve.	Boitani et al. (2006) | Conenna et al. (2017) | Corbet & Hill (1992) | Hill (1976, 1983) | Hutson, Kingston & Francis (2008b) | Simmons (2005)	https://zenodo.org/record/6398107/files/figure.png	161. Gaskell’s False Serotine Hesperoptenus gaskelli French: Sérotine de Gaskell / German: Gaskell-Scheinbreitfligel / Spanish: Hesperopteno de Gaskell Taxonomy. Hesperoptenus (Milithronycteris) gaskelli Hill, 1983 , “ Central Sulawesi [Indoncsial: ... River Ranu, 1°51’ S , 121° 30° E .” Subgenus Milithronycteris . Relationships of H. gaskelli to other Hesperoptenus are unknown because it has not been included in a phylogenetic study. Monotypic. Distribution. Known only from type locality in EC Sulawesi . Descriptive notes. Forearm 40-1 mm (type specimen, juvenile female). Gaskell’s False Serotine resembles Tickell’s False Serotine ( H. tickelli ) and the Large False Serotine ( H. tomesi ) but differs from both in its smaller size and teeth. Color (alcohol specimen) is rich chocolate-brown dorsally and ventrally. Hairs are more brownish at base but otherwise darker and lustrous. Muzzle is low, wide, laterally swollen, and sparsely haired. Internarial region is naked. Ears are thick, fleshy, and subquadrangular. Tragus is rather fleshy and more or less hatched-shaped. Thumb is relatively long, and second phalanx ofthird digit is shorter than first phalanx; upper surface of thumb has thin hairs; upper surface of forearm is naked; upper surface of uropatagium is sparsely haired on proximal one-half; base of tail is similarly furred above and below, with tip protruded beyond uropatagium. Skull is medium-sized, with broad non-inflated braincase; postorbital region is moderately constricted; and supraorbital region is abruptly expanded, with well-developed superior anteorbital tubercles. I? is caniniform and massive, and I’ is more or less ovate. I’ is displaced inward so that I? and C! are in contact. Condylo-canine length is 13-1 mm, and maxillary tooth row is 6 mm . Habitat. Holotype was caught in small clearing on a covered (with vegetation) path at the edge of primary forest at 10 m above sea level. Food and Feeding. No information. Breeding. No information. Activity patterns. Nocturnal. Single specimen was caught in a harp trap with its top 3 m aboveground (bottom of trapping area would then be c.1-1-5 m aboveground). Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The IUCN Red List. Gaskell’s False Serotine is known only from one locality in Morowali Nature Reserve. Bibliography. Boitani et al. (2006), Conenna et al. (2017), Corbet & Hill (1992), Hill (1976, 1983), Hutson, Kingston & Francis (2008b), Simmons (2005).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Hesperoptenus gaskelli	Hesperoptenus	Milithronycteris	gaskelli	Hill	1983	0	Bull. Br. Mus. Nat. Hist. Zool.	47:49:00	Gaskell's False Serotine	None.	Indonesia, Sulawesi, Central R. Ranu (01°51'S, 121°30'E).	Sulawesi (known only from the type locality).	Not listed.	Data Deficient	Subgenus Milithronycteris .	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hesperoptenus gaskelli	23	Gaskell's False Serotine		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	EPTESICINI	Hesperoptenus	Milithronycteris	gaskelli	J. Edwards Hill	1983	0						"Central Sulawesi [Indonesia]: … River Ranu, 1°51' S, 121° 30' E."	-1.85	121.5	gaskelli J. Edwards Hill, 1983	NA	NA	Indonesia	Asia	Australasia/Oceania	DD	0	0	0	Hesperoptenus_gaskelli	0	sciname match	Hesperoptenus_gaskelli	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	9977	Hesperoptenus gaskelli	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Hesperoptenus	gaskelli	Hill, 1983	Hesperoptenus gaskelli is recognized as a species (Simmons and Cirranello 2020).	20000000	Hesperoptenus gaskelli	Data Deficient		2021	2021-07-23 00:00:00 UTC	3.1	English	Hesperoptenus gaskelli is assessed as Data Deficient as the species is known only from one site in Sulawesi, Indonesia. Limited research in the area has not confirmed new records. Additional research is warranted to confirm its distribution, population status and trends, threats, and basic ecology.	The habitat and ecology of the insectivorous Hesperoptenus gaskelli is largely unknown as it has only been encountered one time. A single specimen was captured at the edge of primary forest in a small clearing.	The threats to this species are unknown as the species has only been captured one time. Given it was captured at the edge of lowland primary forest, it is inferred that the extensive deforestation occurring in Sulawesi due to commercial logging (Supriatna et al. 2020) may pose a significant threat to the species.	There are no population size and trends data available for this species as it is known from only one specimen. It may be threatened and of latent risk due to habitat loss within the region (Davidson et al. 2017).	Unknown	Limited survey efforts in Sulawesi have only documented this species from the type locality Morowali Nature Reserve Ranu Ri, Sulawesi (01°51'S, 121°30'E), Indonesia (Le Saout et al. 2013). It is inferred to be a low elevation species with a distribution up to 500 m asl.	This species is not known to be hunted, used, or traded.	Terrestrial	The species has been only documented within Morowali Nature Reserve ; in central Sulawesi, Indonesia (Le Saout et al. 2013). Timely and comprehensive research is warranted to established its distribution, population status and trends, threats, and ecology.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Hesperoptenus	Milithronycteris	gaskelli	Hill	1983	0	Bull. Br. Mus. Nat. Hist. Zool.	47:49:00	Gaskell's False Serotine	None.	Indonesia, Sulawesi, Central R. Ranu (01°51'S, 121°30'E).	Sulawesi (known only from the type locality).	Not listed.	Data Deficient	Subgenus Milithronycteris .	Hesperoptenus gaskelli	1005550	23	Gaskell's False Serotine		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	EPTESICINI	Hesperoptenus	Milithronycteris	gaskelli	J. Edwards Hill	1983	0						"Central Sulawesi [Indonesia]: … River Ranu, 1°51' S, 121° 30' E."	-1.85	121.5	gaskelli J. Edwards Hill, 1983	NA	NA				Indonesia	Asia	Australasia/Oceania	DD	0	0	0	Hesperoptenus_gaskelli	0	sciname match	Hesperoptenus_gaskelli	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hesperoptenus_gaskelli	1005550	23	Gaskell's False Serotine		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Nycticeiini	Hesperoptenus	NA	gaskelli	J. Edwards Hill	0	Hesperoptenus (Milithronycteris) gaskelli	Hill, J.E. 1983. Bats (Mammalia: Chiroptera) from Indo-Australia. Bulletin of the British Museum (Natural History) (Zoology)45(3):103-208.	https://www.biodiversitylibrary.org/page/2356815	BMNH:Mamm:1982.135	holotype	https://data.nhm.ac.uk/object/0f5947a1-5077-4a0a-a490-2a45b40d3604 | https://data.nhm.ac.uk/object/fe461e67-9b3e-4807-a649-27e0a74bd74d	"Central Sulawesi [Indonesia]: … River Ranu, 1°51' S, 121° 30' E."	-1.85	121.5	NA	NA				Indonesia	Asia	Australasia	DD	0	0	0	Hesperoptenus_gaskelli	0	sciname match	Hesperoptenus_gaskelli	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Hesperoptenus		gaskelli	Hill	1983	0	Bull. Br. Mus. Nat. Hist. Zool.	47:49:00	Gaskell's False Serotine	None.	Indonesia, Sulawesi, Central R. Ranu (01°51'S, 121°30'E).	Sulawesi (known only from the type locality).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/9977/22076119/' target='_blank'>Data Deficient</a>	None.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hesperoptenus gaskelli; Hesperoptenus gaskelli; Hesperoptenus gaskelli; Hesperoptenus gaskelli; Hesperoptenus gaskelli; Hesperoptenus gaskelli; gaskelli; Sérotine de Gaskell; Gaskell-Scheinbreitfligel; Hesperopteno de Gaskell; Gaskell's False Serotine; Gaskell's False Serotine; Gaskell's False Serotine; H. gaskelli