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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L37	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	Pipistrellus paterculus	Pipistrellus paterculus	Pipistrellus paterculus	Pipistrellus paterculus	Pipistrellus paterculus	Pipistrellus paterculus	Pipistrellus paterculus	Pipistrellus paterculus	Pipistrellus paterculus	Pipistrellus paterculus	Pipistrellus paterculus	Pipistrellus paterculus	Alionoctula paterculus		[MSW2] Subgenus Pipistrellus. Included in abramus by Ellernran and Morrison-Scott (1951), but see Hill and Harrison (1987).; [MSW3] Subgenus Pipistrellus. Included in abramus by Ellerman and Morrison-Scott (1951), but see Hill and Harrison (1987), Corbet and Hill (1992), Bates and Harrison (1997), and Bates et al. (1997), and Hendrichsen et al. (2001b). Also see Lunde et al. (2003a).; [HMW] Pipistrellus paterculus Thomas, 1915 , “Upper Burma [= Myanmar ]. Mt. Popa.” Pipistrellus paterculus has been included as a subspecies or synonym of P. javanicus but is generally recognized as a distinct species based on morphological and genetic data. Two subspecies recognized.; [batnames2022] Subgenus Pipistrellus . Included in abramus by Ellerman and Morrison-Scott (1951), but see Hill and Harrison (1987), Corbetand Hill (1992), Bates and Harrison (1997), and Bates et al. (1997), and Hendrichsen et al. (2001 b ). Also see Lunde et al. (2003 a ).; [IUCN] This taxon was traditionally assigned to the “coromandra ” subgroup of “pipistrellus ” species group (e.g.: Corbet and Hill 1992). According to available molecular genetic data (Roehrs et al. 2010, Benda et al. 2016 orig.), P. tenuis is a part of genetic cluster of Oriental pipistrelles. This cluster is quite distinct from all the West Palearctic pipistrelles and may be referred to as “javanicus ” species group. Earlier treated as subspecies of Pipistrellus abramus (Temminck, 1840) (Ellerman and Morrison-Scott 1951, Hill 1962, Soota and Chaturvedi 1980), it has been accorded specific status by Hill and Harrison (1987).; [batnames2023] Subgenus Pipistrellus . Included in abramus by Ellerman and Morrison-Scott (1951), but see Hill and Harrison (1987), Corbetand Hill (1992), Bates and Harrison (1997), and Bates et al. (1997), and Hendrichsen et al. (2001 b ). Also see Lunde et al. (2003 a ).; [batnames2025_1.7] Subgenus Pipistrellus. Included in abramus by Ellerman and Morrison-Scott (1951), but see Hill and Harrison (1987), Corbetand Hill (1992), Bates and Harrison (1997), and Bates et al. (1997), and Hendrichsen et al. (2001b). Also see Lunde et al. (2003 a).; [MDD2025_2.2] moved from Pipistrellus to Alionoctula						yunnanensis.	yunnanensis	paterculus, yunnanensis		paterculus, yunnanensis		paterculus, yunnanensis		paterculus, yunnanensis	This taxon was traditionally assigned to the “coromandra ” subgroup of “pipistrellus ” species group (e.g.: Corbet and Hill 1992). According to available molecular genetic data (Roehrs et al. 2010, Benda et al. 2016 orig.), P. tenuis is a part of genetic cluster of Oriental pipistrelles. This cluster is quite distinct from all the West Palearctic pipistrelles and may be referred to as “javanicus ” species group. Earlier treated as subspecies of Pipistrellus abramus (Temminck, 1840) (Ellerman and Morrison-Scott 1951, Hill 1962, Soota and Chaturvedi 1980), it has been accorded specific status by Hill and Harrison (1987).	paterculus, yunnanensis		paterculus, yunnanensis	paterculus, yunnanensis	paterculus, yunnanensis		paterculus (O. Thomas, 1915)|yunnanensis (Wang Yingxiang, 1982)						N/A					Distribution: Definitely known only from northern Burma, but has been recorded from Thailand and northern India.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		NE India – S China; ref. 4.123	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1915	J. Bombay Nat. Hist. Soc., 24:32.	Subgenus Pipistrellus. Included in abramus by Ellernran and Morrison-Scott (1951), but see Hill and Harrison (1987).	N India, Burma, Thailand, SW China.	Burma, Mt. Popa.		THOMAS	1915	Size relatively small (forearm length, 29-34 mm). Inner upper incisor bicuspid. Outer upper incisor well developed. Anterior upper premolar more or less displaced medially. Rostrum relatively short and broad. Forehead almost flat. Penis greatly enlarged. A species of uncertain relationships.	Distribution: Definitely known only from northern Burma, but has been recorded from Thailand and northern India.	A subspecies (P. p. yunnanensis), has been described from southwestern China.		113	species	P. paterculus	THOMAS	1915	Pipistrellus	subgenus	Pipistrellus paterculus				Size relatively small (forearm length, 29-34 mm). Inner upper incisor bicuspid. Outer upper incisor well developed. Anterior upper premolar more or less displaced medially. Rostrum relatively short and broad. Forehead almost flat. Penis greatly enlarged. A species of uncertain relationships.	A subspecies (P. p. yunnanensis), has been described from southwestern China.		13. P. paterculus THOMAS 1915 [pipistrellus group],	13	_A. p. paterculus_ (Thomas, 1915); _A. p. yunnanensis_ (Wang, 1982)			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Pipistrellini	Pipistrellus paterculus	Pipistrellus	Pipistrellus	paterculus	Thomas		1915		J. Bombay Nat. Hist. Soc.	24		32		Mount Popa Pipistrelle	Burma, Mt. Popa.	N India, Burma, Thailand, Vietnam, SW China.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).	yunnanensis Wang, 1982.	Subgenus Pipistrellus. Included in abramus by Ellerman and Morrison-Scott (1951), but see Hill and Harrison (1987), Corbet and Hill (1992), Bates and Harrison (1997), and Bates et al. (1997), and Hendrichsen et al. (2001b). Also see Lunde et al. (2003a).	4C3D87E8FFEA6A56FA9392D01DB8B84F	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	781	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFEC6A53FF9092061C26B04E.xml	Pipustrellus paterculus	Vespertilionidae	Pipistrellus	paterculus	Thomas	1915	Pipistrelle du Popa @fr | Mount-Popa-Zwergfledermaus @de | Pipistrela de Popa @es | Paternal Pipistrelle @en	Pipistrellus paterculus Thomas, 1915 , “Upper Burma [= Myanmar ]. Mt. Popa.” Pipistrellus paterculus has been included as a subspecies or synonym of P. javanicus but is generally recognized as a distinct species based on morphological and genetic data. Two subspecies recognized.	P.p.paterculusThomas,1915—N&NEIndia(JammuandKashmir,Bihar,Assam,Nagaland,andManipur),Myanmar,ELaos,Vietnam,andN&SWCambodia;alsoreportedfromNWThailandbutrequiresconfirmation. P. p. yunnanensis Wang Yingxiang, 1982 — SW China ( Yunnan ).	Head-body 42-48 mm, tail 31-38 mm, ear 10-13 mm, hindfoot 6-7 mm, forearm 26-34 mm; weight 4:3-5-7 g. The Mount Popa Pipistrelle is very similar to the Javan Pipistrelle ( P. javanicus ) and the Japanese Pipistrelle ( P. abramus ), distinguished largely by general color, penial structure, and canine shape. Pelage of the Mount Popa Pipistrelle is long, thick, and silky. Dorsum is dark brown (nearly black), with golden brown (or reddish) tipped hairs; venter is paler, with black-based and buffy brown tipped hairs. Ears, face, and membranes are dark brown. Uropatagium stretches from calcar to nearly tail tip (only very tip is free). Baculum is characteristically very large and long (exceeding 9 mm ), with long narrow shaft and strongly bifurcated tip that deflects ventrally; it is similar than that of the Japanese Pipistrelle but straighter and thicker. Skull has robust braincase; zygomatic arches are robust, with small dorsal process on each jugal bone; I? is bicuspid, and I’ is unicuspid and exceeds height of secondary cusp of I*; C' lacks secondary cusp unlike other Asiatic Pipustrellus; P? is small and slightly displaced; and lower molars are nyctalodont.	Primary and secondary forests in lowland and montane regions, limestone karst regions, and disturbed and urban areas from sea level up to elevations of ¢. 2400 m .	No information.	In central Myanmar , a subadult Mount Popa Pipistrelle was captured in lateJune.	The Mount Popa Pipistrelle is nocturnal. Roosts have been recorded in banana and bamboo trees, hollows in trees, holes in tree stumps, roofs of thatched huts, and limestone caves.	No information.	Classified as Least Concern on The IUCN Red List. There are currently no major threats affecting the Mount Popa Pipistrelle, but localized deforestation from logging and agricultural expansion might be a future threat.	Bates & Harrison (1997) | Bates, Molur & Srinivasulu (2008) | Bates, Nwe Tin et al. (2005) | Bumrungsri et al. (2006) | Corbet & Hill (1992) | Francis (2008a) | Furey et al. (2012) | Hill (1962) | Hill & Harrison (1987) | Kruskop (2013a) | Smith & Xie Yan (2008)	https://zenodo.org/record/6397854/files/figure.png	40. Mount Popa Pipistrelle Pipustrellus paterculus French: Pipistrelle du Popa / German: Mount-Popa-Zwergfledermaus / Spanish: Pipistrela de Popa Other common names: Paternal Pipistrelle Taxonomy. Pipistrellus paterculus Thomas, 1915 , “Upper Burma [= Myanmar ]. Mt. Popa.” Pipistrellus paterculus has been included as a subspecies or synonym of P. javanicus but is generally recognized as a distinct species based on morphological and genetic data. Two subspecies recognized. Subspecies and Distribution. P.p.paterculusThomas,1915—N&NEIndia(JammuandKashmir,Bihar,Assam,Nagaland,andManipur),Myanmar,ELaos,Vietnam,andN&SWCambodia;alsoreportedfromNWThailandbutrequiresconfirmation. P. p. yunnanensis Wang Yingxiang, 1982 — SW China ( Yunnan ). Descriptive notes. Head-body 42-48 mm, tail 31-38 mm, ear 10-13 mm, hindfoot 6-7 mm, forearm 26-34 mm; weight 4:3-5-7 g. The Mount Popa Pipistrelle is very similar to the Javan Pipistrelle ( P. javanicus ) and the Japanese Pipistrelle ( P. abramus ), distinguished largely by general color, penial structure, and canine shape. Pelage of the Mount Popa Pipistrelle is long, thick, and silky. Dorsum is dark brown (nearly black), with golden brown (or reddish) tipped hairs; venter is paler, with black-based and buffy brown tipped hairs. Ears, face, and membranes are dark brown. Uropatagium stretches from calcar to nearly tail tip (only very tip is free). Baculum is characteristically very large and long (exceeding 9 mm ), with long narrow shaft and strongly bifurcated tip that deflects ventrally; it is similar than that of the Japanese Pipistrelle but straighter and thicker. Skull has robust braincase; zygomatic arches are robust, with small dorsal process on each jugal bone; I? is bicuspid, and I’ is unicuspid and exceeds height of secondary cusp of I*; C' lacks secondary cusp unlike other Asiatic Pipustrellus; P? is small and slightly displaced; and lower molars are nyctalodont. Habitat. Primary and secondary forests in lowland and montane regions, limestone karst regions, and disturbed and urban areas from sea level up to elevations of ¢. 2400 m . Food and Feeding. No information. Breeding. In central Myanmar , a subadult Mount Popa Pipistrelle was captured in lateJune. Activity patterns. The Mount Popa Pipistrelle is nocturnal. Roosts have been recorded in banana and bamboo trees, hollows in trees, holes in tree stumps, roofs of thatched huts, and limestone caves. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCN Red List. There are currently no major threats affecting the Mount Popa Pipistrelle, but localized deforestation from logging and agricultural expansion might be a future threat. Bibliography. Bates & Harrison (1997), Bates, Molur & Srinivasulu (2008), Bates, Nwe Tin et al. (2005), Bumrungsri et al. (2006), Corbet & Hill (1992), Francis (2008a), Furey et al. (2012), Hill (1962), Hill & Harrison (1987), Kruskop (2013a), Smith & Xie Yan (2008).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Pipistrellus paterculus	Pipistrellus	Pipistrellus	paterculus	Thomas	1915	0	J. Bombay Nat. Hist. Soc.	######	Mount Popa Pipistrelle	<b> yunnanensis </b>Wang, 1982.	Burma, Mt. Popa.	N India, Burma, Thailand, Vietnam, SW China.	Not listed.	Least Concern	Subgenus Pipistrellus . Included in abramus by Ellerman and Morrison-Scott (1951), but see Hill and Harrison (1987), Corbetand Hill (1992), Bates and Harrison (1997), and Bates et al. (1997), and Hendrichsen et al. (2001 b ). Also see Lunde et al. (2003 a ).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Pipistrellus paterculus	23	Mount Popa Pipistrelle	Paternal Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PIPISTRELLINI	Pipistrellus	NA	paterculus	O. Thomas	1915	0	Pipistrellus_paterculus	Thomas, O. (1915). Scientific results from the mammal survey. No. XI. A. -On Pipistrels of the genera Pipistrellus and Scotozous. The Journal of the Bombay Natural History Society, 24, 32.	https://www.biodiversitylibrary.org/item/95302#page/72/mode/1up	BM 1914.7.19.242		"Upper Burma [= Myanmar]. Mt. Popa."			paterculus O. Thomas, 1915|yunnanensis Wang Yingxiang, 1982	NA	NA	India|Myanmar|China|Laos|Vietnam|Cambodia|Thailand?	Asia	Indomalaya|Palearctic	LC	0	0	0	Pipistrellus_paterculus	0	sciname match	Pipistrellus_paterculus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	17356	Pipistrellus paterculus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Pipistrellus	paterculus	Thomas, 1915	This taxon was traditionally assigned to the “coromandra ” subgroup of “pipistrellus ” species group (e.g.: Corbet and Hill 1992). According to available molecular genetic data (Roehrs et al. 2010, Benda et al. 2016 orig.), P. tenuis is a part of genetic cluster of Oriental pipistrelles. This cluster is quite distinct from all the West Palearctic pipistrelles and may be referred to as “javanicus ” species group. Earlier treated as subspecies of Pipistrellus abramus (Temminck, 1840) (Ellerman and Morrison-Scott 1951, Hill 1962, Soota and Chaturvedi 1980), it has been accorded specific status by Hill and Harrison (1987).	20000000	Pipistrellus paterculus	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, it occurs in a number of protected areas, has a tolerance of a degree of habitat modification, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	In South Asia, this species is found in primary and secondary forests. It roosts among banana and bamboo trees, deep holes in trees and roofs of thatched huts in forested areas (Molur et al. 2002). In Myanmar, it is predominantly found in limestone caves, although it has also been recorded from forest areas without karst habitats. In Vietnam it is found in light pristine forests far from karst areas (S. Kruskop pers. comm.). It can be found in disturbed areas including agricultural land (Bates et al. 1997, P. Bates pers. comm.). Nothing is known about its reproduction biology.	In Southeast Asia there are no believed to be any major threats to this species as a whole (P. Bates pers. comm.), though continued deforestation may threaten to some local populations. In South Asia, this species is threatened by deforestation, generally resulting from logging operations and the conversion of land to agricultural and other uses (Molur et al. 2002).	This species is relatively common in many areas in Myanmar (P. Bates pers. comm.). In South Asia, although this is a widely distributed species, a declining trend in its population is being observed (Molur et al. 2002). In suitable habitats in Vietnam this species is inferior in number to other co-occurring pipistrelles, though nothing can be said about the population trends.	Unknown	This species is distributed in northern South Asia, southern China and parts of mainland Southeast Asia. In South Asia, this species is presently known from India (Assam, Bihar, Jammu and Kashmir, Manipur and Nagaland), where it has been recorded from sea level to an elevation of 1,500 m asl (Molur et al. 2002). Mention for Pakistan (Kruskop 2013) based on a point on map published by Corbett and Hill (1992) and represents a definite mistake. In China, it is restricted to Yunnan (Smith and Xie 2008). In Southeast Asia, it has been recorded from Myanmar, northern Thailand (requires confirmation Bumrungsri et al. 2006), Lao PDR and Vietnam. In Vietnam it occurs sporadically, mainly at low elevations (Kruskop 2013).	There are no evidences that this species was used in commercial trade, in local cuisine or in local medicine. Like other insectivorous bats, P. paterculus plays certain role as natural regulator of some harvest and forestry pests.	Terrestrial	It has been recorded from some protected areas in Southeast Asia (e.g. from Binchau-Phuocbuu and Yokdon in Vietnam). In South Asia, there are no direct conservation measures in place for this species and it has not been recorded from any protected areas. Further studies are needed into the distribution, abundance, reproduction and ecology of this species. Populations of this species should be monitored to record changes in abundance and distribution (Molur et al. 2002).	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Pipistrellus	Pipistrellus	paterculus	Thomas	1915	0	J. Bombay Nat. Hist. Soc.	24:32:00	Mount Popa Pipistrelle	<b> yunnanensis </b>Wang, 1982.	Burma, Mt. Popa.	N India, Burma, Thailand, Vietnam, SW China.	Not listed.	Least Concern	Subgenus Pipistrellus . Included in abramus by Ellerman and Morrison-Scott (1951), but see Hill and Harrison (1987), Corbetand Hill (1992), Bates and Harrison (1997), and Bates et al. (1997), and Hendrichsen et al. (2001 b ). Also see Lunde et al. (2003 a ).	Pipistrellus paterculus	1005630	23	Mount Popa Pipistrelle	Paternal Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PIPISTRELLINI	Pipistrellus	NA	paterculus	O. Thomas	1915	0	Pipistrellus_paterculus	Thomas, O. (1915). Scientific results from the mammal survey. No. XI. A. -On Pipistrels of the genera Pipistrellus and Scotozous. The Journal of the Bombay Natural History Society, 24, 32.	https://www.biodiversitylibrary.org/item/95302#page/72/mode/1up	BM 1914.7.19.242		"Upper Burma [= Myanmar]. Mt. Popa."			paterculus O. Thomas, 1915|yunnanensis Wang Yingxiang, 1982	NA	NA				India|Myanmar|China|Laos|Vietnam|Cambodia|Thailand?	Asia	Indomalaya|Palearctic	LC	0	0	0	Pipistrellus_paterculus	0	sciname match	Pipistrellus_paterculus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Alionoctula_paterculus	1005630	23	Mount Popa Pipistrelle	Paternal Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Pipistrellini	Alionoctula	NA	paterculus	O. Thomas	1	Pipistrellus paterculus	Thomas, O. 1915-09-30. On pipistrels of the genera _Pipistrellus_ and _Scotozous_. Journal of the Bombay Natural History Society 24(1):29-34.	https://www.biodiversitylibrary.org/page/30151555	BMNH:Mamm:1914.7.19.242	holotype	https://data.nhm.ac.uk/object/0c3ef46a-78bb-4c83-a87d-14a901f45eb1	"Upper Burma [= Myanmar]. Mt. Popa."			moved from Pipistrellus to Alionoctula	Zhukova, S. S., Yuzefovich, A. P., Lebedev, V. S., & Kruskop, S. V. (2025). Reassessment of the Taxonomic Borders Within Pipistrellus (Chiroptera, Vespertilionidae, Pipistrellini). Diversity, 17(5), 317.				India|Myanmar|China|Laos|Vietnam|Cambodia|Thailand?	Asia	Indomalaya|Palearctic	LC	0	0	0	Pipistrellus_paterculus	0	sciname match	Pipistrellus_paterculus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Pipistrellus	Pipistrellus	paterculus	Thomas	1915	0	J. Bombay Nat. Hist. Soc.	24:32:00	Mount Popa Pipistrelle	yunnanensis Wang, 1982.	Burma, Mt. Popa.	N India, Burma, Thailand, Vietnam, SW China.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/17356/22126738/' target='_blank'>Least Concern</a>	Subgenus Pipistrellus. Included in abramus by Ellerman and Morrison-Scott (1951), but see Hill and Harrison (1987), Corbetand Hill (1992), Bates and Harrison (1997), and Bates et al. (1997), and Hendrichsen et al. (2001b). Also see Lunde et al. (2003 a).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Pipistrellus paterculus; Pipistrellus paterculus; Pipistrellus paterculus; Pipistrellus paterculus; Pipistrellus paterculus; Pipistrellus paterculus; paterculus; yunnanensis; paterculus; yunnanensis; yunnanensis; paterculus; yunnanensis; Pipistrelle du Popa; Mount-Popa-Zwergfledermaus; Pipistrela de Popa; Paternal Pipistrelle; Mount Popa Pipistrelle; Paternal Pipistrelle; Mount Popa Pipistrelle; Mount Popa Pipistrelle; P. paterculus