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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L360	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Hesperoptenus blanfordi	Hesperoptenus blanfordi	Hesperoptenus blanfordi	Hesperoptenus blanfordi	Hesperoptenus blanfordi	Hesperoptenus blanfordi	Hesperoptenus blanfordi	Hesperoptenus blanfordi	Hesperoptenus blanfordi	Hesperoptenus blanfordi	Hesperoptenus blanfordi	Hesperoptenus blanfordi	Hesperoptenus blanfordi	Hesperoptenus blanfordi	Hesperoptenus blanfordi		[MSW2] Subgenus Milithronycteris.; [MSW3] Subgenus Milithronycteris.; [HMW] Vesperugo (Hesperoptenus) blanfordi Dobson, 1877 , “Tenasserim,” Myanmar . Subgenus Milithronycteris . See G. doriae . T. Gorfol and colleagues in 2019 found H. blanfordi sister to Arielulus circumdatus . Monotypic.; [batnames2022] Subgenus Milithronycteris .; [batnames2023] Subgenus Milithronycteris .; [batnames2025_1.7] None.														blanfordi				blanfordi	blanfordi, blandfordi			blanfordi (Dobson, 1877)|blandfordi O'Brien, Menninger, & W. G. Nash, 2006 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Blanford's bat	S Burma – Malaya	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Hesperoptenus blanfordi	Burma, Tenasserim.	Dobson	1877	J. Asiat. Soc. Bengal, 46:312.	Distribution: Known from southern Burma, Thailand, Malaya, and Borneo.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Blanford's bat	S Burma – Malaya, Borneo	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Dobson	1877	J. Asiat. Soc. Bengal, 46:312.	Subgenus Milithronycteris.	Burma, Thailand, Malay Peninsula, Borneo.	Burma, Tenasserim.		DOBSON	1877	Size relatively small (forearm length, 24-29 mm). A broad pad present at the base of the thumb. Outer upper incisor greatly reduced. Dentition not especially massive. Lower incisors not imbricated. Fur fairly dark.	Distribution: Known from southern Burma, Thailand, Malaya, and Borneo.	No subspecies.		123	species	H. blanfordi	DOBSON	1877	Milithronycteris	subgenus	Hesperoptenus blanfordi				Size relatively small (forearm length, 24-29 mm). A broad pad present at the base of the thumb. Outer upper incisor greatly reduced. Dentition not especially massive. Lower incisors not imbricated. Fur fairly dark.	No subspecies.		5. H. blanfordi (DOBSON 1877).	5	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Eptesicini	Hesperoptenus blanfordi	Hesperoptenus	Milithronycteris	blanfordi	Dobson	y	1877		J. Asiat. Soc. Bengal	46		312		Blanford's Bat	Burma, E of Moulmein, Tenasserim.	Burma, Thailand, Cambodia, Laos, Malay Peninsula, Borneo.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).		Subgenus Milithronycteris.	4C3D87E8FFB66A09FF7C9B5516FBB054	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	835	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFB66A09FF7C9B5516FBB054.xml	Hesperoptenus blanfordi	Vespertilionidae	Hesperoptenus	blanfordi	Dobson	1877	Sérotine de Blanford @fr | Blanford-Scheinbreitfligel @de | Hesperoptero de Blanford @es | Blanford’s Bat @en | Least False Serotine @en	Vesperugo (Hesperoptenus) blanfordi Dobson, 1877 , “Tenasserim,” Myanmar . Subgenus Milithronycteris . See G. doriae . T. Gorfol and colleagues in 2019 found H. blanfordi sister to Arielulus circumdatus . Monotypic.	SE Myanmar (Tenasserim) S through W and peninsular Thailand to Peninsular Malaysia , with scattered records from C & S Laos (Phou Hin Poun National Biodiversity Conservation Area and Xe Pian National Protected Area), SW & S Cambodia (Cardamom and Elephant Mts), S Vietnam ( Dong Nai and Ba Ria-Vung Tau provinces and Phu Quoc I), and Borneo (Niah National Park in Sarawak , Witti Range and Sepilok in Sabah , and East Kalimantan).	Head-body 37-45 mm, tail 33-34 mm, ear 11-12 mm, hindfoot 4-5-6-6 mm, forearm 24-29 mm; weight 5-5-8 g. Blanford’s False Serotine is the smallest species of Hesperoptenus , appearing similar to bamboo bats (7ylonycteris spp.) with very short forearms and well-developed adhesive pads on thumbs but not on feet (unique among Hesperoptenus ). Pelage is rather sleek, with glossy sheen. Upperparts are dark brown, with reddish tinge, and underparts are slightly paler. Fur on shoulders is sparser but longer and lighter. Ears and muzzle are grayish brown. Internarial region and upper surface of forearm are densely haired. Wing membranes are grayish brown or brown. Ears are short and triangular. Tragus is short, blunt, and anteriorly directed, with length subequal to width. Calcar has well-developed lobe, with transverse septum. Thumb pads are thickened, dark-brown, and kidney-shaped. Head is slightly pointed, with smoothly sloping profile. Nostrils are widely separated. Baculum has long, slender, ventrally fluted shaft, tip is expanded vertically and horizontally, and base is expanded into broad paired flanges separated by shallow V-shaped aperture. There are differences in baculum size between Malayan and Bornean populations, the latter smaller in body and teeth measurements. Skull is very small, with wide flattened braincase and lambdoid ridges but no sagittal crest. Frontal part of face is flat, without any flexure. I? is greatly reduced. I’ is displaced inward,so that I and C' are in contact. Condylo-canine lengths are 11-8-12-5 mm, and maxillary tooth row lengths are 4-2—-4.-4 mm. Chromosomal complement has 2n = 32 and FN = 54 in Malaysia and 2n = 32 and FN = 56 in Thailand .	Wide variety of habitats from lowland wet forests to dry dipterocarp forests and hill forests from sea level up to elevations of ¢. 1300 m . In Vietnam , recorded in heavily disturbed agricultural landscapes and usually associated with small rivers and streams.	Blanford’s False Serotine is insectivorous.	In Vietnam , pregnant Blanford’s False Serotines were recorded at the end of April. Three captive females had one young each.	Blanford’s False Serotines are nocturnal and fly ¢.7-10 m aboveground, usually in open spaces. In Krau Wildlife Reserve, Malaysia , they were mistnetted over rivers. In Malaysia , they roost in entrances of limestone caves in hill forests. Presence of thumb pads suggests they can hang and move on smooth surfaces and might roost in bamboo. As very skillful runners, they often try to reach nearby hideouts quickly. During the day, they preferred to roost in horizontal columns, where they often slept with their bellies up. Unusual for Vespertilionidae , Blanford’s False Serotine has two call types, with similar frequency modulation but different frequencies, which frequently change in a regular way. This alternation was heard in nine of eleven individuals after release; only high-frequency calls could be observed for the other two. High-frequency pulses (durations of 3-9 milliseconds) started at ¢.105 kHz at the maximum and ended between 40 kHz and 45 kHz; low-frequency calls (durations of 3-10 milliseconds) started at c.105 kHz but swept down to 31-37 kHz. Reported lower starting frequencies might result from larger distances between microphone and bat during recording (high-frequency calls with mean peak frequency of 44 kHz and duration of 6-6 milliseconds and low-frequency calls with peak of 36 kHz and duration of 8-3 milliseconds).	Blanford’s False Serotine lives in small colonies, with less than ten individuals.	Classified as Least Concern on The IUCN Red List.	Abramov et al. (2007) | Amador et al. (2018) | Boitani et al. (2006) | Borisenko & Kruskop (2003) | Corbet & Hill (1992) | Francis (2008a) | Francis et al. (2010) | Fukuda et al. (2009) | Gorfdl et al. (2019) | Hassanin et al. (2018) | Heller (1989) | Heller & Volleth (1989) | Hendrichsen, Bates & Hayes (2001) | Hill (1976) | Hill & Francis (1984) | Hutson, Kingston , Francis, Banks & Csorba (2008) | Kingston , Jones et al. (2003) | Kingston , Lim & Zubaid (2006) | Koubinova et al. (2013) | Kruskop (2013a) | Lim et al. (2017) | Matveev (2005) | McBee et al. (1986) | Medway (1983) | Payne et al. (1985) | Pearch & Writer (2009) | Phillipps & Phillipps (2016) | Robinson (1998) | Robinson & Webber (1998) | Shazali et al. (2016) | Simmons (2005) | Volleth etal. (2001)	https://zenodo.org/record/6398103/files/figure.png	159. Blanford’s False Serotine Hesperoptenus blanfordi French: Sérotine de Blanford / German: Blanford-Scheinbreitfligel / Spanish: Hesperoptero de Blanford Other common names: Blanford’s Bat , Least False Serotine Taxonomy. Vesperugo (Hesperoptenus) blanfordi Dobson, 1877 , “Tenasserim,” Myanmar . Subgenus Milithronycteris . See G. doriae . T. Gorfol and colleagues in 2019 found H. blanfordi sister to Arielulus circumdatus . Monotypic. Distribution. SE Myanmar (Tenasserim) S through W and peninsular Thailand to Peninsular Malaysia , with scattered records from C & S Laos (Phou Hin Poun National Biodiversity Conservation Area and Xe Pian National Protected Area), SW & S Cambodia (Cardamom and Elephant Mts), S Vietnam ( Dong Nai and Ba Ria-Vung Tau provinces and Phu Quoc I), and Borneo (Niah National Park in Sarawak , Witti Range and Sepilok in Sabah , and East Kalimantan). Descriptive notes. Head-body 37-45 mm, tail 33-34 mm, ear 11-12 mm, hindfoot 4-5-6-6 mm, forearm 24-29 mm; weight 5-5-8 g. Blanford’s False Serotine is the smallest species of Hesperoptenus , appearing similar to bamboo bats (7ylonycteris spp.) with very short forearms and well-developed adhesive pads on thumbs but not on feet (unique among Hesperoptenus ). Pelage is rather sleek, with glossy sheen. Upperparts are dark brown, with reddish tinge, and underparts are slightly paler. Fur on shoulders is sparser but longer and lighter. Ears and muzzle are grayish brown. Internarial region and upper surface of forearm are densely haired. Wing membranes are grayish brown or brown. Ears are short and triangular. Tragus is short, blunt, and anteriorly directed, with length subequal to width. Calcar has well-developed lobe, with transverse septum. Thumb pads are thickened, dark-brown, and kidney-shaped. Head is slightly pointed, with smoothly sloping profile. Nostrils are widely separated. Baculum has long, slender, ventrally fluted shaft, tip is expanded vertically and horizontally, and base is expanded into broad paired flanges separated by shallow V-shaped aperture. There are differences in baculum size between Malayan and Bornean populations, the latter smaller in body and teeth measurements. Skull is very small, with wide flattened braincase and lambdoid ridges but no sagittal crest. Frontal part of face is flat, without any flexure. I? is greatly reduced. I’ is displaced inward,so that I and C' are in contact. Condylo-canine lengths are 11-8-12-5 mm, and maxillary tooth row lengths are 4-2—-4.-4 mm. Chromosomal complement has 2n = 32 and FN = 54 in Malaysia and 2n = 32 and FN = 56 in Thailand . Habitat. Wide variety of habitats from lowland wet forests to dry dipterocarp forests and hill forests from sea level up to elevations of ¢. 1300 m . In Vietnam , recorded in heavily disturbed agricultural landscapes and usually associated with small rivers and streams. Food and Feeding. Blanford’s False Serotine is insectivorous. Breeding. In Vietnam , pregnant Blanford’s False Serotines were recorded at the end of April. Three captive females had one young each. Activity patterns. Blanford’s False Serotines are nocturnal and fly ¢.7-10 m aboveground, usually in open spaces. In Krau Wildlife Reserve, Malaysia , they were mistnetted over rivers. In Malaysia , they roost in entrances of limestone caves in hill forests. Presence of thumb pads suggests they can hang and move on smooth surfaces and might roost in bamboo. As very skillful runners, they often try to reach nearby hideouts quickly. During the day, they preferred to roost in horizontal columns, where they often slept with their bellies up. Unusual for Vespertilionidae , Blanford’s False Serotine has two call types, with similar frequency modulation but different frequencies, which frequently change in a regular way. This alternation was heard in nine of eleven individuals after release; only high-frequency calls could be observed for the other two. High-frequency pulses (durations of 3-9 milliseconds) started at ¢.105 kHz at the maximum and ended between 40 kHz and 45 kHz; low-frequency calls (durations of 3-10 milliseconds) started at c.105 kHz but swept down to 31-37 kHz. Reported lower starting frequencies might result from larger distances between microphone and bat during recording (high-frequency calls with mean peak frequency of 44 kHz and duration of 6-6 milliseconds and low-frequency calls with peak of 36 kHz and duration of 8-3 milliseconds). Movements, Home range and Social organization. Blanford’s False Serotine lives in small colonies, with less than ten individuals. Status and Conservation. Classified as Least Concern on The IUCN Red List. Bibliography. Abramov et al. (2007), Amador et al. (2018), Boitani et al. (2006), Borisenko & Kruskop (2003), Corbet & Hill (1992), Francis (2008a), Francis et al. (2010), Fukuda et al. (2009), Gorfdl et al. (2019), Hassanin et al. (2018), Heller (1989), Heller & Volleth (1989), Hendrichsen, Bates & Hayes (2001), Hill (1976), Hill & Francis (1984), Hutson, Kingston , Francis, Banks & Csorba (2008), Kingston , Jones et al. (2003), Kingston , Lim & Zubaid (2006), Koubinova et al. (2013), Kruskop (2013a), Lim et al. (2017), Matveev (2005), McBee et al. (1986), Medway (1983), Payne et al. (1985), Pearch & Writer (2009), Phillipps & Phillipps (2016), Robinson (1998), Robinson & Webber (1998), Shazali et al. (2016), Simmons (2005), Volleth etal. (2001).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Hesperoptenus blanfordi	Hesperoptenus	Milithronycteris	blanfordi	Dobson	1877	1	J. Asiat. Soc. Bengal	51:12:00	Blanford's Bat	None.	Burma, E of Moulmein, Tenasserim.	Burma, Thailand, Cambodia, Laos, Malay Peninsula, Borneo.	Not listed.	Least Concern	Subgenus Milithronycteris .	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hesperoptenus blanfordi	23	Blanford's False Serotine	Blanford's Bat|Least False Serotine	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	EPTESICINI	Hesperoptenus	Milithronycteris	blanfordi	Dobson	1877	1	Vesperugo_(Hesperoptenus)_blanfordi	Dobson, G. E. (1877). Notes on a collection of Chiroptera from India and Burma, with descriptions of new species. The Journal of the Asiatic Society of Bengal, 46, 2, 312.	https://www.biodiversitylibrary.org/item/110105#page/324/mode/1up	ZSI 15593		"Tenasserim," Myanmar.			blanfordi (Dobson, 1877)	NA	NA	Myanmar|Thailand|Laos|Cambodia|Vietnam|Malaysia|Indonesia	Asia	Indomalaya	LC	0	0	0	Hesperoptenus_blanfordi	0	sciname match	Hesperoptenus_blanfordi	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	9975	Hesperoptenus blanfordi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Hesperoptenus	blanfordi	(Dobson, 1877)		20000000	Hesperoptenus blanfordi	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern as the species is considered widespread, it occurs in a variety of habitats, there are no known major threats, it occurs in a number of protected areas, and it is unlikely to be declining fast enough for listing in a more threatened category.	This species is found in a wide range of habitats, from lowland wet forest to dry dipterocarp forest and hill forest, and also limestone caves. It is usually associated with small rivers and streams. It usually lives in small colonies, with less than 10 individuals.	There are no major threats to this species. This species is apparently tolerant to a broad range of habitats.	This species is uncommon throughout its scattered distribution through mainland Southeast Asia and Borneo.	Unknown	This species is found from east of Moulmein, Tenasserim (Myanmar), extending to Thailand and Peninsular Malaysia. There are records also from Cambodia (P. Banks pers. comm.), Lao PDR, Viet Nam, and Sabah where it is only known from two localities (the Witti Range and Sepilok), Sarawak known from Tanjung Datu National Park and Niah National Park (P. Rajasegaran pers.com) and east Kalimantan (one specimen, A. Suyanto pers. comm.) in Borneo. In Lao PDR the species is reported, by Robinson and Webber (1998), from Khammouane Limestone National Biodiversity Conservation Area, and from Xe Pian in southern Lao PDR by Robinson (1998). In Viet Nam the species is known from Don Nai province, Vung Tau province and Phu Quoc Island (Abramov et al. 2007 and Kruskop 2013).		Terrestrial	This species is found in a number of protected areas throughout its range.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Hesperoptenus	Milithronycteris	blanfordi	Dobson	1877	1	J. Asiat. Soc. Bengal	51:12:00	Blanford's Bat	None.	Burma, E of Moulmein, Tenasserim.	Burma, Thailand, Cambodia, Laos, Malay Peninsula, Borneo.	Not listed.	Least Concern	Subgenus Milithronycteris .	Hesperoptenus blanfordi	1005548	23	Blanford's False Serotine	Blanford's Bat|Least False Serotine	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	EPTESICINI	Hesperoptenus	Milithronycteris	blanfordi	Dobson	1877	1	Vesperugo_(Hesperoptenus)_blanfordi	Dobson, G. E. (1877). Notes on a collection of Chiroptera from India and Burma, with descriptions of new species. The Journal of the Asiatic Society of Bengal, 46, 2, 312.	https://www.biodiversitylibrary.org/item/110105#page/324/mode/1up	ZSI 15593		"Tenasserim," Myanmar.			blanfordi (Dobson, 1877)	NA	NA				Myanmar|Thailand|Laos|Cambodia|Vietnam|Malaysia|Indonesia	Asia	Indomalaya	LC	0	0	0	Hesperoptenus_blanfordi	0	sciname match	Hesperoptenus_blanfordi	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hesperoptenus_blanfordi	1005548	23	Blanford's False Serotine	Blanford's Bat|Least False Serotine	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Nycticeiini	Hesperoptenus	NA	blanfordi	Dobson	1	Vesperugo (Hesperoptenus) Blanfordi	Dobson, G.E. 1877. Notes on a collection of Chiroptera from India and Burma, with descriptions of new species. Journal of the Asiatic Society of Bengal 46(3):310-313.	https://www.biodiversitylibrary.org/page/35548400	ZSI 15593	holotype		"Tenasserim," Myanmar.			NA	NA				Myanmar|Thailand|Laos|Cambodia|Vietnam|Malaysia|Indonesia	Asia	Indomalaya	LC	0	0	0	Hesperoptenus_blanfordi	0	sciname match	Hesperoptenus_blanfordi	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Hesperoptenus		blanfordi	Dobson	1877	1	J. Asiat. Soc. Bengal	51:12:00	Blanford's Bat	None.	Burma, E of Moulmein, Tenasserim.	Burma, Thailand, Cambodia, Laos, Malay Peninsula, Borneo.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/9975/22076582/' target='_blank'>Least Concern</a>	None.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hesperoptenus blanfordi; Hesperoptenus blanfordi; Hesperoptenus blanfordi; Hesperoptenus blanfordi; Hesperoptenus blanfordi; Hesperoptenus blanfordi; blanfordi; Sérotine de Blanford; Blanford-Scheinbreitfligel; Hesperoptero de Blanford; Blanford’s Bat; Least False Serotine; Blanford's False Serotine; Blanford's Bat; Least False Serotine; Blanford's Bat; Blanford's Bat; H. blanfordi