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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L359	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Harpyionycteris whiteheadi	Harpyionycteris whiteheadi	Harpyionycteris whiteheadi	Harpyionycteris whiteheadi	Harpyionycteris whiteheadi	Harpyionycteris whiteheadi	Harpyionycteris whiteheadi	Harpyionycteris whiteheadi	Harpyionycteris whiteheadi	Harpyionycteris whiteheadi	Harpyionycteris whiteheadi	Harpyionycteris whiteheadi	Harpyionycteris whiteheadi	Harpyionycteris whiteheadi	Harpyionycteris whiteheadi		[MSW3] Does not include celebensis; see comments under that species.; [HMW] Harpyionycternis whiteheadi Thomas, 1896 , “Mindoro [ Island ], Philippines , alt. 5000 feet [= 1524 m ].” Two subspecies are recognized.; [batnames2022] Does not include celebensis ; see comments under that species.; [IUCN] Further study of geographic variation in this species is needed.; [batnames2023] Does not include celebensis ; see comments under that species.; [batnames2025_1.7] Does not include celebensis; see comments under that species.						negrosensis.	celebensis, whiteheadi, negrosensis	whiteheadi, negrosensis		whiteheadi, negrosensis		whiteheadi, negrosensis		whiteheadi, negrosensis	Further study of geographic variation in this species is needed.	whiteheadi, negrosensis		whiteheadi, negrosensis	whiteheadi, negrosensis	negrosensis, whiteheadi		whiteheadi O. Thomas, 1896|negrosensis R. L. Peterson & Fenton, 1970		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Harpy fruit bat	Philippines	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Harpyionycteris whiteheadi	Philippines, Mindoro Isl., 5,000 ft. (1524 m).	Thomas	1896	Ann. Mag. Nat. Hist., ser. 6, 18:244.	Distribution: Known from Celebes and the Philippine islands of Mindanao, Camiguin, Negros, and Mindoro.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Harpy fruit bat	Philippines	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1896	Ann. Mag. Nat. Hist., ser. 6, 18:244.		Philippines.	Philippines, Mindoro Isl, 5000 ft. (1524 m).		THOMAS	1896	Forearm length, 82-93 mm.	Distribution: Known from Celebes and the Philippine islands of Mindanao, Camiguin, Negros, and Mindoro.	Three sub species are currently recognized:	H. w. celebensis (Celebes), H. w. whiteheadi (Philippines ex cept Negros), H. w. negrosensis (Negros).	30	species	H. whiteheadi	THOMAS	1896	Harpyionycteris	genus	Harpyionycteris whiteheadi				Forearm length, 82-93 mm.	Three sub species are currently recognized:		1. H. whiteheadi THOMAS 1896.	1	_H. w. negrosensis_ Peterson & Fenton, 1970; _H. w. whiteheadi_ Thomas, 1896			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Harpyionycteris whiteheadi	Harpyionycteris		whiteheadi	Thomas		1896		Ann. Mag. Nat. Hist., ser. 6	18		244		Harpy Fruit Bat	Philippines, Mindoro Isl, 5,000 ft. (1,524 m).	Philippines except Palawan region.	IUCN/SSC Action Plan (1992) – Not Threatened. IUCN 2003 – Lower Risk (lc).	negrosensis Peterson and Fenton, 1970.	Does not include celebensis; see comments under that species.	03AD87FAFFF4F61B8CB73EA7FD94F4F8	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	106	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFF4F61B8CB73EA7FD94F4F8.xml	Harpyionycteris whiteheadi	Pteropodidae	Harpyionycteris	whiteheadi	Thomas	1896	Harpyionyctére des Philippines @fr | Philippinen-Harpyien-Flughund @de | Harpyionicterio de las Filipinas @es | Harpy Fruit Bat @en	Harpyionycternis whiteheadi Thomas, 1896 , “Mindoro [ Island ], Philippines , alt. 5000 feet [= 1524 m ].” Two subspecies are recognized.	H.w.whiteheadiThomas,1896—Philip-pinesinSLuzon,EVisayas,andMindanaoIsgroups;butnotdefinitiverecordsinCat-anduanes,Marinduque,Bohol,Sibuyan,andTablasIs.Thelattertwoislandsmayholdpopulationsofthesubspeciesnegrosensis. H. w. negrosensis Peterson & Fenton, 1970 — WVisayas (Panay, Negros, and Cebu Is).	Head-body 130-159 mm (tailless), ear 20-25 mm , hindfoot 21- 25 mm , forearm 84-91 mm ; weight 99-140 g . Head is round with strong, conical muzzle; nostrils are shortly tubular; philtrum has two separate deep grooves and one median groove; and lowerlip has large triangular papillae. Eyes are moderately large, somewhat oblique; irises are warm brown. Ears are long, oval, and pale brown, with broadly rounded off tips, narrower in negrosensis. Head pelage is short and chocolatebrown, and head midline is often obvious. Nape and dorsum pelage is long, dense, and uniformly rich buffy brown, as is rest of body including chest, belly, and flanks. Dense fur extends over much of forearm and along hindleg to dorsum of foot. Tibia is relatively very short, calcar is very small, uropatagium is reduced to narrow strip, and foot claws have whitish tips. Wing membranes are broad, from sides of body, blackish brown, speckled with small white spots, and inserted on second toes; index claw is present. Skull has conical rostrum, premaxillae are fused and project forward, with nasal process very thin; orbit is large, with obvious rim; zygomatic root is just above upper alveolar line; zygoma is moderately strong and arched; and braincase is rounded. Dorsally, premaxillae project farther anteriorly than tip of nasals; rostrum 1s long and relatively narrow; nasals are very narrow; paranasal recesses are moderately inflated, reaching tiny postorbital foramen; postorbital process is long and posterolaterally directed; postorbital constriction is well-marked; braincase is oval; and sagittal and nuchal crests are obvious and low. Ventrally, palate is long, flat, and narrow; tooth rows are nearly parallel; post-dental palate is relatively short, ending concavely; and ectotympanic is small and narrow, with anterior extension and spine. Mandible has slanted symphysis; body is somewhat arched; coronoid slopes, with rounded tip; condyle is level with lower alveolar line; and angle is large, round, and ample. There are nine arched interdental palatal ridges, with last two divided medially, plus two post-dental ridges, one in middle and one at end of post-dental palate. Dentition is strong, profusely cuspidate, and strongly procumbent, with upper incisors (likely I*, I' missing) very large, hook-like, with ample base, forwardly converging tip, and deeply concave outer margin; C' projects anteriorly at nearly 45°, is large with convex anterior surface, concave internal surface with middle ridge, strong secondary basal distal cusp, and obvious inner cingulum; P' is small, but crown has distinct central and secondary distal cusps; next premolar (P°) is blade- or canine-like, decurved with basal distal cusp obsolete, and sharp lingual cingulum that is denticulate on its mesial side; next premolars and molars have subdivided labial and lingual main ridges, with additional cusps; more frequently, there are three larger cusps on each ridge with additional smaller cusps, but pattern is exceedingly variable; M? is small but cuspidate; and cheekteeth size is moderate, with oval occlusal outline. Lower incisor (likely I, I, missing) is very small, bifid, and generally missing in adults; C, projects anteriorly (but less than C'), almost straight, with cusp coming out high on mesial side, larger basal distal cusp, and sharp cingulum connecting along labial side of canine; P| is relatively very large, with three-cusp pattern similar to C; posterior cheekteeth have subdivided labial and lingual main ridges, with 3—4 cusps each and strong distal ledge; last molar is equally cuspidate; cheekteeth are rather narrow, with anterior tear-shaped to posterior oval occlusal outline. Chromosomal complement has 2n = 36 and FN = 58, with six pairs of medium to large metacentric to submetacentric, six pairs of medium-sized subtelocentric, and five pairs of medium-sized acrocentric autosomes. X-chromosome is mediumsized subtelocentric, and Y-chromosome small acrocentric.	Primary or lightly disturbed rainforests from sea level up to elevations of c. 1900 m . The Philippine Harpy Fruit Bat is more abundant in montane rainforests and less common in lowland rainforests and mossy forest.	In Panay, diet of the Philippine Harpy Fruit Bat contained at least twelve plant genera in ten families. It probably forages in forest canopies.	Philippine Harpy Fruit Bats are seasonally polyestrous. Females give birth for the first time at c.1 year old, and then twice a year in synchronous birth periods:January— February and July-August. Gestation lasts 4-5 months, and lactation lasts 3-4 months, indicating postpartum estrus or at least mating during lactation in both birth seasons.	No information.	A small roosting colony was found in an understory tree in a forest interior. The Philippine Harpy Fruit Bat produces high-pitched, shrieking whistles when they fly, presumably to keep contact among individuals. On Negros Island, density was estimated at 0-7 ind/ha.	Classified as Least Concern on The IUCN Red List. The Philippine Harpy Fruit Batis locally affected by logging, but populations are presumably stable in montane rainforests that are little disturbed.	Aimeida et al. (2011) | Amador et al. (2018) | Andersen (1912b) | Giannini & Simmons (2007a) | Giannini, Almeida & Simmons (2009) | Giannini, Almeida, Simmons & DeSalle (2006) | Heaney, Balete et al. (1998) | Heaney, Tabaranza et al. (2006) | Heideman (1987) | Heideman & Heaney (1989) | Ong, Rosell-Ambal, Tabaranza, Heaney, Gonzalez et al. (2008) | Paguntalan & Jakosalem (2008) | Peterson & Fenton (1970) | Rickart Rickart, Heaney & Rosenfeld (1989) | Tate (1951)		80. Philippine Harpy Fruit Bat Harpyionycteris whiteheadi French: Harpyionyctére des Philippines / German: Philippinen-Harpyien-Flughund / Spanish: Harpyionicterio de las Filipinas Other common names: Harpy Fruit Bat Taxonomy. Harpyionycternis whiteheadi Thomas, 1896 , “Mindoro [ Island ], Philippines , alt. 5000 feet [= 1524 m ].” Two subspecies are recognized. Subspecies and Distribution. H.w.whiteheadiThomas,1896—Philip-pinesinSLuzon,EVisayas,andMindanaoIsgroups;butnotdefinitiverecordsinCat-anduanes,Marinduque,Bohol,Sibuyan,andTablasIs.Thelattertwoislandsmayholdpopulationsofthesubspeciesnegrosensis. H. w. negrosensis Peterson & Fenton, 1970 — WVisayas (Panay, Negros, and Cebu Is). Descriptive notes. Head-body 130-159 mm (tailless), ear 20-25 mm , hindfoot 21- 25 mm , forearm 84-91 mm ; weight 99-140 g . Head is round with strong, conical muzzle; nostrils are shortly tubular; philtrum has two separate deep grooves and one median groove; and lowerlip has large triangular papillae. Eyes are moderately large, somewhat oblique; irises are warm brown. Ears are long, oval, and pale brown, with broadly rounded off tips, narrower in negrosensis. Head pelage is short and chocolatebrown, and head midline is often obvious. Nape and dorsum pelage is long, dense, and uniformly rich buffy brown, as is rest of body including chest, belly, and flanks. Dense fur extends over much of forearm and along hindleg to dorsum of foot. Tibia is relatively very short, calcar is very small, uropatagium is reduced to narrow strip, and foot claws have whitish tips. Wing membranes are broad, from sides of body, blackish brown, speckled with small white spots, and inserted on second toes; index claw is present. Skull has conical rostrum, premaxillae are fused and project forward, with nasal process very thin; orbit is large, with obvious rim; zygomatic root is just above upper alveolar line; zygoma is moderately strong and arched; and braincase is rounded. Dorsally, premaxillae project farther anteriorly than tip of nasals; rostrum 1s long and relatively narrow; nasals are very narrow; paranasal recesses are moderately inflated, reaching tiny postorbital foramen; postorbital process is long and posterolaterally directed; postorbital constriction is well-marked; braincase is oval; and sagittal and nuchal crests are obvious and low. Ventrally, palate is long, flat, and narrow; tooth rows are nearly parallel; post-dental palate is relatively short, ending concavely; and ectotympanic is small and narrow, with anterior extension and spine. Mandible has slanted symphysis; body is somewhat arched; coronoid slopes, with rounded tip; condyle is level with lower alveolar line; and angle is large, round, and ample. There are nine arched interdental palatal ridges, with last two divided medially, plus two post-dental ridges, one in middle and one at end of post-dental palate. Dentition is strong, profusely cuspidate, and strongly procumbent, with upper incisors (likely I*, I' missing) very large, hook-like, with ample base, forwardly converging tip, and deeply concave outer margin; C' projects anteriorly at nearly 45°, is large with convex anterior surface, concave internal surface with middle ridge, strong secondary basal distal cusp, and obvious inner cingulum; P' is small, but crown has distinct central and secondary distal cusps; next premolar (P°) is blade- or canine-like, decurved with basal distal cusp obsolete, and sharp lingual cingulum that is denticulate on its mesial side; next premolars and molars have subdivided labial and lingual main ridges, with additional cusps; more frequently, there are three larger cusps on each ridge with additional smaller cusps, but pattern is exceedingly variable; M? is small but cuspidate; and cheekteeth size is moderate, with oval occlusal outline. Lower incisor (likely I, I, missing) is very small, bifid, and generally missing in adults; C, projects anteriorly (but less than C'), almost straight, with cusp coming out high on mesial side, larger basal distal cusp, and sharp cingulum connecting along labial side of canine; P| is relatively very large, with three-cusp pattern similar to C; posterior cheekteeth have subdivided labial and lingual main ridges, with 3—4 cusps each and strong distal ledge; last molar is equally cuspidate; cheekteeth are rather narrow, with anterior tear-shaped to posterior oval occlusal outline. Chromosomal complement has 2n = 36 and FN = 58, with six pairs of medium to large metacentric to submetacentric, six pairs of medium-sized subtelocentric, and five pairs of medium-sized acrocentric autosomes. X-chromosome is mediumsized subtelocentric, and Y-chromosome small acrocentric. Habitat. Primary or lightly disturbed rainforests from sea level up to elevations of c. 1900 m . The Philippine Harpy Fruit Bat is more abundant in montane rainforests and less common in lowland rainforests and mossy forest. Food and Feeding. In Panay, diet of the Philippine Harpy Fruit Bat contained at least twelve plant genera in ten families. It probably forages in forest canopies. Breeding. Philippine Harpy Fruit Bats are seasonally polyestrous. Females give birth for the first time at c.1 year old, and then twice a year in synchronous birth periods:January— February and July-August. Gestation lasts 4-5 months, and lactation lasts 3-4 months, indicating postpartum estrus or at least mating during lactation in both birth seasons. Activity patterns. No information. Movements, Home range and Social organization. A small roosting colony was found in an understory tree in a forest interior. The Philippine Harpy Fruit Bat produces high-pitched, shrieking whistles when they fly, presumably to keep contact among individuals. On Negros Island, density was estimated at 0-7 ind/ha. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Philippine Harpy Fruit Batis locally affected by logging, but populations are presumably stable in montane rainforests that are little disturbed. Bibliography. Aimeida et al. (2011), Amador et al. (2018), Andersen (1912b), Giannini & Simmons (2007a), Giannini, Almeida & Simmons (2009), Giannini, Almeida, Simmons & DeSalle (2006), Heaney, Balete et al. (1998), Heaney, Tabaranza et al. (2006), Heideman (1987), Heideman & Heaney (1989), Ong, Rosell-Ambal, Tabaranza, Heaney, Gonzalez et al. (2008), Paguntalan & Jakosalem (2008), Peterson & Fenton (1970), Rickart Rickart, Heaney & Rosenfeld (1989), Tate (1951).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Harpyionycteris whiteheadi	Harpyionycteris		whiteheadi	Thomas	1896	0	Ann. Mag. Nat. Hist.	ser. 6, 18: 244: 244	Harpy Fruit Bat	<b> negrosensis </b>Peterson and Fenton, 1970.	Philippines, Mindoro Isl, 5,000 ft. 1,524 m	Philippines except Palawan region.	Not listed.	Least Concern	Does not include celebensis ; see comments under that species.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Harpyionycteris whiteheadi	23	Philippine Harpy Fruit Bat	Harpy Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	HARPYIONYCTERINAE	HARPYIONYCTERINI	Harpyionycteris	NA	whiteheadi	O. Thomas	1896	0						"Mindoro [Island], Philippines, alt. 5000 feet [= 1524 m]."			whiteheadi O. Thomas, 1896|negrosensis R. L. P. Peterson & Fenton, 1970	NA	NA	Philippines	Asia	Indomalaya	LC	0	0	0	Harpyionycteris_whiteheadi	0	sciname match	Harpyionycteris_whiteheadi	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	9740	Harpyionycteris whiteheadi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Harpyionycteris	whiteheadi	Thomas, 1896	Further study of geographic variation in this species is needed.	20000000	Harpyionycteris whiteheadi	Least Concern		2021	2020-12-15 00:00:00 UTC	3.1	English	Harpyionycteris whiteheadi is considered Least Concern, as it is relatively widespread and common in suitable habitats. It is unlikely that there have been substantial population declines in the past. There has been significant lowland deforestation, but this species uses a range of forest conditions, including montane forests which remain more intact. Populations on particular islands such as Cebu are highly threatened.	Harpyionycteris whiteheadi occurs in lowland and montane forest, in primary, lightly disturbed and secondary habitats. It has been found in an old reforestation area planted with exotic species but adjacent to secondary lowland forest, lowland and in montane forest. (Fidelino et al. 2020). These are fairly large bats that fly moderately high and are known to roost in trees, but not caves. They might depend on the fruits of viney pandans (Freycinetia spp.) (Heaney 1984, Heaney et al. 1989, Heideman and Heaney 1989, Rickart et al. 1993).	Harpyionycteris whiteheadi is rarely if ever hunted. Although not considered to be a major threat this species is vulnerable to deforestation (Heaney et al. 1998), which has occurred particularly in the lowlands rather than in the species' preferred montane habitat. However, local (island) populations can be threatened by deforestation on islands where limited forests remain (e.g., Cebu).	Populations of the Harpy Fruit Bat are considered to be stable at present, their main habitat is montane forest, which is mostly intact (Heaney et al. 1998). It is unlikely that there has been a substantial population decline in the past and although this species is not abundant it is considered uncommon to common in lowland forest and moderately common in montane forest (L. Heaney pers. comm. 2006). In a 2003 mist net survey on Mount Apo, Mindanao, H. whiteheadi was uncommon in lower elevation mossy-montane forest that had been subjected to habitat alteration (Godfrey et al. unpublished report). Recent surveys between 2014-2016 indicate that the species is relatively common in the lower montane forest in northern Negros at 1,125 m and in a lowland evergreen forest at 1,075 m in southern Negros Island (Fidelino et al. 2020). But relatively uncommon in the montane forest in Mt Apo between 1,100 and 1,500 m (Duya pers. Comm) and in the lowland secondary forest at 765 m in Leyte (Fidelino et al. 2020). The species occurs in low numbers in reforestation areas in Leyte (Fidelino et al. 2020). Heaney et al. (2016) reported the species to have moderately low density in southern Negros, estimated at 0.7/ha at 850-1,100 m. One individual was captured in Mt Masulong, Mindoro Island, out of 145 net-nights in a montane forest at 1,010 m (Ong et al. 2013).	Stable	Harpyionycteris whiteheadi is endemic to the Philippines, where it is widespread excluding the Palawan Faunal Region and the Batanes/Babuyan group of islands. It has been recorded from Biliran, Camiguin, Cebu, Leyte, Luzon (Camarines Sur, there are no records from northern Luzon (L. Heaney pers. comm. 2006)), Maripipi, Masbate, Mindanao (Agusan del Norte, Bukidnon, Davao Oriental, Lanao del Norte, Lanao del Sur, Misamis Occidental, Misamis Oriental, Southern Cotabato, Suriago del Sur and Zamboanga del Norte provinces), Mindoro Oriental (Mt. Masulong, Roxas (Ong et al. 2013), Anahawin, River Calintaan (Gonzales and Dans 1995), Negros, and Samar. Its elevation range is from sea level to 1,800 m.		Terrestrial	This species occurs in some protected areas (Mt. Apo Natural Park, Mt. Kanlaon Natural Park). Additional research is needed on the species distribution, population status and trends, and threats.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Harpyionycteris		whiteheadi	Thomas	1896	0	Ann. Mag. Nat. Hist.	ser. 6, 18: 244: 244	Harpy Fruit Bat	<b> negrosensis </b>Peterson and Fenton, 1970.	Philippines, Mindoro Isl, 5,000 ft. 1,524 m	Philippines except Palawan region.	Not listed.	Least Concern	Does not include celebensis ; see comments under that species.	Harpyionycteris whiteheadi	1004406	23	Philippine Harpy Fruit Bat	Harpy Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	HARPYIONYCTERINAE	HARPYIONYCTERINI	Harpyionycteris	NA	whiteheadi	O. Thomas	1896	0						"Mindoro [Island], Philippines, alt. 5000 feet [= 1524 m]."			whiteheadi O. Thomas, 1896|negrosensis R. L. P. Peterson & Fenton, 1970	NA	NA				Philippines	Asia	Indomalaya	LC	0	0	0	Harpyionycteris_whiteheadi	0	sciname match	Harpyionycteris_whiteheadi	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Harpyionycteris_whiteheadi	1004406	23	Philippine Harpy Fruit Bat	Harpy Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Harpyionycterinae	Harpyionycterini	Harpyionycteris	NA	whiteheadi	O. Thomas	0	Harpyionycteris Whiteheadi	Thomas, O. 1896-09-01. On mammals from Celebes, Borneo, and the Philippines recently received at the British Museum. Annals and Magazine of Natural History (6)18(105):241-250.	https://www.biodiversitylibrary.org/page/15628431	BMNH:Mamm:1897.5.2.7	holotype	https://data.nhm.ac.uk/object/216fdbe7-f976-4b24-9324-551d75aa0ceb	"Mindoro [Island], Philippines, alt. 5000 feet [= 1524 m]."			NA	NA				Philippines	Asia	Indomalaya	LC	0	0	0	Harpyionycteris_whiteheadi	0	sciname match	Harpyionycteris_whiteheadi	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Harpyionycteris		whiteheadi	Thomas	1896	0	Ann. Mag. Nat. Hist.	ser. 6, 18: 244: 244	Harpy Fruit Bat	negrosensis Peterson and Fenton, 1970.	Philippines, Mindoro Isl, 5,000 ft. 1,524 m	Philippines except Palawan region.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/9740/22045044/' target='_blank'>Least Concern</a>	Does not include celebensis; see comments under that species.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Harpyionycteris whiteheadi; Harpyionycteris whiteheadi; Harpyionycteris whiteheadi; Harpyionycteris whiteheadi; Harpyionycteris whiteheadi; Harpyionycteris whiteheadi; whiteheadi; negrosensis; whiteheadi; negrosensis; negrosensis; whiteheadi; negrosensis; Harpyionyctére des Philippines; Philippinen-Harpyien-Flughund; Harpyionicterio de las Filipinas; Harpy Fruit Bat; Philippine Harpy Fruit Bat; Harpy Fruit Bat; Harpy Fruit Bat; Harpy Fruit Bat; H. whiteheadi