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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L357	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Harpiola isodon	Harpiola isodon	Harpiola isodon	Harpiola isodon	Harpiola isodon	Harpiola isodon	Harpiola isodon	Harpiola isodon	Harpiola isodon		[HMW] Harpiola isodon Haochih et al., 2006 , “ Hualien County , Jhuosi Township, Yuli Wildlife Refuge [ Taiwan ], 23°32'N , 121°15'E ,. 2,000 m asl.” Limited genetic data place this species within Murina , close to M. harrisoni , M. hutton, and M. ussuriensis , making Murina paraphyletic and calling the generic validity of Harpiola into doubt. Monotypic.; [MDD2022] recently described; [IUCN] Distinct from its only congeneric, Harpiola grisea , from India by dental features (Kuo et al. 2006) and probably also by craniodental dimensions (Kruskop et al. 2006).; [MDD2023] recently described; [MDD2025_2.0] recently described; [batnames2025_1.7] For the record from China see Li et al. (2024). A specimen referred to as Harpiola cf. isodon has been reported from Vietnam; see Kruskop et al. (2006).; [MDD2025_2.2] recently described														isodon	Distinct from its only congeneric, Harpiola grisea , from India by dental features (Kuo et al. 2006) and probably also by craniodental dimensions (Kruskop et al. 2006).			isodon	isodon			isodon Kuo Haochih, Fang Yinping, Csorba, & Lee Lingling, 2006						N/A																																								NA																											4C3D87E8FF696AD7FA959E7E1D4BB858	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	907	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF696AD7FA959E7E1D4BB858.xml	Harpiola isodon	Vespertilionidae	Harpiola	isodon	Haochih	2006	Murine isodonte @fr | Goldgelbe Formosa-Rohrennase @de | Ratonero isodonte @es | Golden-haired Tube-nosed Bat @en | Taiwan Tube-nosed Bat @en	Harpiola isodon Haochih et al., 2006 , “ Hualien County , Jhuosi Township, Yuli Wildlife Refuge [ Taiwan ], 23°32'N , 121°15'E ,. 2,000 m asl.” Limited genetic data place this species within Murina , close to M. harrisoni , M. hutton, and M. ussuriensis , making Murina paraphyletic and calling the generic validity of Harpiola into doubt. Monotypic.	NW & C Vietnam (Hoang Lien National Park and Ngoc Linh National Reserve) and Taiwan .	Head-body 43-53 mm , tail 24- 5-36 mm , ear 12- 4-15 mm , hindfoot 5-:8-8- 1 mm , forearm 30-37- 3 mm ; weight 3-4- 8 g . Fur is long and woolly; dorsally shiny golden yellow (hairs dark-brown-based with bright yellow bands and a dark brown tip, and dark brown guard hairs with shiny golden yellow tips mixed throughout); ventrally browner (hairs dark brown exceptfor light brown terminal two-thirds), sometimes with grayish white hairs along midline of chest and abdomen. Bright dorsal hairs continue onto bones ofwings, thumb, uropatagium, and feet. Last vertebra oftail extends beyond uropatagium. Wings and ears are dark brown, and face is furred except for long, tubular nostrils. Ears have distinct notch at upperthird of posterior border; tragus is moderately long but wide basally, gradually tapering to a backward-curved tip, with small, toothlike projection on outer margin. Skull is small, with narrow, shallow rostrum; braincase is elevated above rostrum with a distinctly higher mid-portion than the lambda; I* is bicuspid and enlarged; P* and P, are well developed, and are subequal to the canines and second premolars of the corresponding tooth row; C,is strongly bifid. Chromosomal complement has 2n = 44 and FN = 50 ( Vietnam ). On following pages: 327. Rufous Tube-nosed Bat ( Murina leucogasten; 328. Bicolored Tube-nosed Bat ( Murina bicolon); 329. Rongjiang Tube-nosed Bat ( Murina rongjiangensis); 330. Fang He Tube-nosed Bat ( Murina fanjingshanensis); 331. Shuipu Tube-nosed Bat ( Murina shuipuensis ); 332. Dusky Tube-nosed Bat ( Murina fusca ); 333. Hilgendorf's Tube-nosed Bat ( Murina hilgendorf); 334. Little Tube-nosed Bat ( Murina aurata ); 335. Jaintia Tube-nosed Bat ( Murina jaintiana); 336. Rainforest Tube-nosed Bat ( Murina pluvialis ); 337. Fea's Tube-nosed Bat ( Murina feae ); 338. Beelzebub Tube-nosed Bat ( Murina beelzebub ); 339. Lorelie’s Tube-nosed Bat ( Murina lorelieae ); 340. Vietnamese Tube-nosed Bat Murina harpioloides ); 341. Golden-haired Tube-nosed Bat ( Murina chrysochaetes ); 342. Harrison's Tube-nosed Bat ( Murina harrisoni ); 343. Ussuri Tube-nosed Bat ( Murina ussuriensis ); 344. Gloomy Tube-nosed Bat ( Murina tenebrosa ); 345. Ryukyu Tube-nosed Bat ( Murina ryukyuana ); 346. Hutton's Tube-nosed Bat ( Murina huttonii ); 347. Taiwan Tube-nosed Bat ( Murina puta ).	Coniferous plantations or mixed forests of coniferous and broadleaved trees in Taiwan . It has been captured over streams in deciduous moss forests in Vietnam . Recorded at elevations of 1000-2400 m in Taiwan , and 1950-2250 m in Vietnam .	It has been seen foraging over stream backwaters and shallows flying c.0-2- 2 m above water or ground. It is a fast and very maneuverable flier.	A pregnant female with one embryo was captured in May in Taiwan .	No information.	Two specimens were collected in a tunnel, where they may have been roosting.	Classified as Data Deficient on The IUCNRed List. Very little is currently known aboutits ecology. This species might be threatened by deforestation.	Kruskop (2013a) | Kruskop & Shchinov (2010) | Kruskop et al. (2006) | Kuo Haochih et al. (2006) | Nguyen Truong Son etal. (2015) | Smith & Johnston (2008c)	https://zenodo.org/record/6398553/files/figure.png	326. Formosan Tube-nosed Bat Harpiola isodon French: Murine isodonte / German: Goldgelbe Formosa-Rohrennase / Spanish: Ratonero isodonte Other common names: Golden-haired Tube-nosed Bat , Taiwan Tube-nosed Bat Taxonomy. Harpiola isodon Haochih et al., 2006 , “ Hualien County , Jhuosi Township, Yuli Wildlife Refuge [ Taiwan ], 23°32'N , 121°15'E ,. 2,000 m asl.” Limited genetic data place this species within Murina , close to M. harrisoni , M. hutton, and M. ussuriensis , making Murina paraphyletic and calling the generic validity of Harpiola into doubt. Monotypic. Distribution. NW & C Vietnam (Hoang Lien National Park and Ngoc Linh National Reserve) and Taiwan . Descriptive notes. Head-body 43-53 mm , tail 24- 5-36 mm , ear 12- 4-15 mm , hindfoot 5-:8-8- 1 mm , forearm 30-37- 3 mm ; weight 3-4- 8 g . Fur is long and woolly; dorsally shiny golden yellow (hairs dark-brown-based with bright yellow bands and a dark brown tip, and dark brown guard hairs with shiny golden yellow tips mixed throughout); ventrally browner (hairs dark brown exceptfor light brown terminal two-thirds), sometimes with grayish white hairs along midline of chest and abdomen. Bright dorsal hairs continue onto bones ofwings, thumb, uropatagium, and feet. Last vertebra oftail extends beyond uropatagium. Wings and ears are dark brown, and face is furred except for long, tubular nostrils. Ears have distinct notch at upperthird of posterior border; tragus is moderately long but wide basally, gradually tapering to a backward-curved tip, with small, toothlike projection on outer margin. Skull is small, with narrow, shallow rostrum; braincase is elevated above rostrum with a distinctly higher mid-portion than the lambda; I* is bicuspid and enlarged; P* and P, are well developed, and are subequal to the canines and second premolars of the corresponding tooth row; C,is strongly bifid. Chromosomal complement has 2n = 44 and FN = 50 ( Vietnam ). On following pages: 327. Rufous Tube-nosed Bat ( Murina leucogasten; 328. Bicolored Tube-nosed Bat ( Murina bicolon); 329. Rongjiang Tube-nosed Bat ( Murina rongjiangensis); 330. Fang He Tube-nosed Bat ( Murina fanjingshanensis); 331. Shuipu Tube-nosed Bat ( Murina shuipuensis ); 332. Dusky Tube-nosed Bat ( Murina fusca ); 333. Hilgendorf's Tube-nosed Bat ( Murina hilgendorf); 334. Little Tube-nosed Bat ( Murina aurata ); 335. Jaintia Tube-nosed Bat ( Murina jaintiana); 336. Rainforest Tube-nosed Bat ( Murina pluvialis ); 337. Fea's Tube-nosed Bat ( Murina feae ); 338. Beelzebub Tube-nosed Bat ( Murina beelzebub ); 339. Lorelie’s Tube-nosed Bat ( Murina lorelieae ); 340. Vietnamese Tube-nosed Bat Murina harpioloides ); 341. Golden-haired Tube-nosed Bat ( Murina chrysochaetes ); 342. Harrison's Tube-nosed Bat ( Murina harrisoni ); 343. Ussuri Tube-nosed Bat ( Murina ussuriensis ); 344. Gloomy Tube-nosed Bat ( Murina tenebrosa ); 345. Ryukyu Tube-nosed Bat ( Murina ryukyuana ); 346. Hutton's Tube-nosed Bat ( Murina huttonii ); 347. Taiwan Tube-nosed Bat ( Murina puta ). Habitat. Coniferous plantations or mixed forests of coniferous and broadleaved trees in Taiwan . It has been captured over streams in deciduous moss forests in Vietnam . Recorded at elevations of 1000-2400 m in Taiwan , and 1950-2250 m in Vietnam . Food and Feeding. It has been seen foraging over stream backwaters and shallows flying c.0-2- 2 m above water or ground. It is a fast and very maneuverable flier. Breeding. A pregnant female with one embryo was captured in May in Taiwan . Activity patterns. No information. Movements, Home range and Social organization. Two specimens were collected in a tunnel, where they may have been roosting. Status and Conservation. Classified as Data Deficient on The IUCNRed List. Very little is currently known aboutits ecology. This species might be threatened by deforestation. Bibliography. Kruskop (2013a), Kruskop & Shchinov (2010), Kruskop et al. (2006), Kuo Haochih et al. (2006), Nguyen Truong Son etal. (2015), Smith & Johnston (2008c).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Harpiola isodon	Harpiola		isodon	Kuo, Fang, Csorba, & Lee	2006	0	Acta Chiropterol.	8(1): 13	Golden-haired Tube-nosed Bat	None.	Taiwan, Hualien County, Jhuosi Township, Yuli, Wildlife Refuge,	Taiwan	Not listed.	Least Concern		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Harpiola isodon	23	Formosan Tube-nosed Bat	Golden-haired Tube-nosed Bat|Taiwan Tube-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MURININAE	NA	Harpiola	NA	isodon	Kuo Haochih, Fang Yinping, Csorba, & Lee Lingling	2006	0	Harpiola_isodon	Kuo, H. C., Fang, Y. P., Csorba, G., & Lee, L. L. (2006). The definition of Harpiola (Vespertilionidae: Murininae) and the description of a new species from Taiwan. Acta Chiropterologica, 8(1), 13.	https://bioone.org/journals/acta-chiropterologica/volume-8/issue-1/1733-5329(2006)8%5B11%3ATDOHVM%5D2.0.CO%3B2/The-definition-of-Harpiola-Vespertilionidae--Murininae-and-the-description/10.3161/1733-5329(2006)8[11:TDOHVM]2.0.CO;2.short	NTU 1998.5.3		"Hualien County, Jhuosi Township, Yuli Wildlife Refuge [Taiwan], 23°32'N, 121°15'E, 2,000 m a.s.l."	23.533	121.25	isodon Kuo Haochih, Fang Yinping, Csorba, & Lee Lingling, 2006	recently described	Kuo, H. C., Fang, Y. P., Csorba, G., & Lee, L. L. (2006). The definition of Harpiola (Vespertilionidae: Murininae) and the description of a new species from Taiwan. Acta Chiropterologica, 8(1), 11-19.	Vietnam|Taiwan	Asia	Palearctic	LC	0	0	0	Harpiola_isodon	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136445	Harpiola isodon	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Harpiola	isodon	Kuo, Fang, Csorba &; Lee, 2006	Distinct from its only congeneric, Harpiola grisea , from India by dental features (Kuo et al. 2006) and probably also by craniodental dimensions (Kruskop et al. 2006).	20000000	Harpiola isodon	Least Concern		2020	2018-09-01 00:00:00 UTC	3.1	English	This species is listed as Least Concern. It was only sporadically reported from the mainland Asia but was regularly recorded in Taiwan. The population in Taiwan is believed not to undergo rapid decline, as the forest habitats of this species there are often under protections.	This species occurs in various types of forests with more or less closed canopy, including those dominated by planted coniferous trees. Individuals were observed foraging 0.5-2m above water surface of a small stream near forest in Vietnam (Kruskop and Shchinov 2010). In Taiwan, two collected specimens were obtained from a tunnel (Kuo et al. 2006). Individuals have been observed hibernating in abandoned tunnels in the mountain area with an elevation over 2,300m from November to early March (H.-C. Chang pers. comm.). Pregnant females were recorded in May while nursing ones were recorded from May to July (Kruskop and Shchinov 2010, Cheng et al. 2017b, H.-C. Kuo pers. obs.).	This species appears to be more abundant in Taiwan than in the mainland Asia although further survey work in the latter area is needed for a confirmation. In Taiwan, this species is adapted to different types of forests which are in general under protection. Accordingly, the island’s population may not be under any major threat. In Vietnam, it has been recorded from Hoang Lien Son National Park (Kruskop and Shchinov 2010). Deforestation and transformation might threaten the country’s populations.	The global population is unknown as there are currently no data regarding the population sizes and trends of this species.	Unknown	Disjunctly distributed in Taiwan and Vietnam. In Taiwan, this species occurs in mountain regions with elevations of up to 2,6oo m asl. An estimated extent of occurrence (EOO) in Taiwan is around 13,000 km² (based on total area within a minimum convex polygon around known occurrences there). In Vietnam, this species is currently only reported from two locations (both with elevations of around 2,000 m asl): one in the Ngoc Linh Mt. in Central Vietnam (Kruskop et al. 2006) and the other one near Hoang Lien Son National Park in the Northwestern Vietnam (Kruskop and Shchinov 2010).		Terrestrial	Present in many protected areas in Taiwan, including national parks, nature reserves, wildlife refuges and major wildlife habitats. Neither listed as a protected species in Taiwan (Cheng et al. 2017a) nor in Vietnam (Vu Dinh Thong, pers. com.). Studies on its distribution in Vietnam and other countries in the mainland, roost use in the winter and influences of deforestation are recommended.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Harpiola		isodon	Kuo, Fang, Csorba, & Lee	2006	0	Acta Chiropterol.	8(1): 13	Golden-haired Tube-nosed Bat	None.	Taiwan, Hualien County, Jhuosi Township, Yuli, Wildlife Refuge,	Taiwan	Not listed.	Least Concern		Harpiola isodon	1005316	23	Formosan Tube-nosed Bat	Golden-haired Tube-nosed Bat|Taiwan Tube-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MURININAE	NA	Harpiola	NA	isodon	Kuo Haochih, Fang Yinping, Csorba, & Lee Lingling	2006	0	Harpiola_isodon	Kuo, H. C., Fang, Y. P., Csorba, G., & Lee, L. L. (2006). The definition of Harpiola (Vespertilionidae: Murininae) and the description of a new species from Taiwan. Acta Chiropterologica, 8(1), 13.	https://bioone.org/journals/acta-chiropterologica/volume-8/issue-1/1733-5329(2006)8%5B11%3ATDOHVM%5D2.0.CO%3B2/The-definition-of-Harpiola-Vespertilionidae--Murininae-and-the-description/10.3161/1733-5329(2006)8[11:TDOHVM]2.0.CO;2.short	NTU 1998.5.3		"Hualien County, Jhuosi Township, Yuli Wildlife Refuge [Taiwan], 23°32'N, 121°15'E, 2,000 m a.s.l."	23.53333	121.25	isodon Kuo Haochih, Fang Yinping, Csorba, & Lee Lingling, 2006	recently described	Kuo, H. C., Fang, Y. P., Csorba, G., & Lee, L. L. (2006). The definition of Harpiola (Vespertilionidae: Murininae) and the description of a new species from Taiwan. Acta Chiropterologica, 8(1), 11-19.				Vietnam|Taiwan	Asia	Palearctic	LC	0	0	0	Harpiola_isodon	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Harpiola_isodon	1005316	23	Formosan Tube-nosed Bat	Golden-haired Tube-nosed Bat|Taiwan Tube-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Murininae	NA	Harpiola	NA	isodon	Kuo Haochih, Fang Yinping, Csorba, & Lee Lingling	0	Harpiola isodon	Kuo, H.-C., Fang, Y.-P., Csorba, G. and Lee, L.-L. 2006-04. The definition of _Harpiola_ (Vespertilionidae: Murininae) and the description of a new species from Taiwan. Acta Chiropterologica 8(1):11-19.	https://doi.org/10.3161/1733-5329(2006)8[11:TDOHVM]2.0.CO;2	ZMNTU 1998.5.3	holotype		"Hualien County, Jhuosi Township, Yuli Wildlife Refuge [Taiwan], 23°32'N, 121°15'E, 2,000 m a.s.l."	23.53333	121.25	recently described	Kuo, H. C., Fang, Y. P., Csorba, G., & Lee, L. L. (2006). The definition of Harpiola (Vespertilionidae: Murininae) and the description of a new species from Taiwan. Acta Chiropterologica, 8(1), 11-19.				Vietnam|Taiwan	Asia	Palearctic	LC	0	0	0	Harpiola_isodon	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Harpiola		isodon	Kuo, Fang, Csorba & Lee	2006	0	Acta Chiropterol.	8(1): 13	Golden-haired Tube-nosed Bat	None.	Taiwan, Hualien County, Jhuosi Township, Yuli, Wildlife Refuge,	Taiwan, Yunnan (China)	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136445/21983827/' target='_blank'>Least Concern</a>	For the record from China see Li et al. (2024). A specimen referred to as Harpiola cf. isodon has been reported from Vietnam; see Kruskop et al. (2006).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Harpiola isodon; Harpiola isodon; Harpiola isodon; Harpiola isodon; Harpiola isodon; isodon; Murine isodonte; Goldgelbe Formosa-Rohrennase; Ratonero isodonte; Golden-haired Tube-nosed Bat; Taiwan Tube-nosed Bat; Formosan Tube-nosed Bat; Golden-haired Tube-nosed Bat; Taiwan Tube-nosed Bat; Golden-haired Tube-nosed Bat; H. isodon