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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L356	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Murina grisea	Murina grisea	Murina grisea	Murina grisea	Murina grisea	Murina grisea	Harpiola grisea	Harpiola grisea	Harpiola grisea	Harpiola grisea	Harpiola grisea	Harpiola grisea	Harpiola grisea	Harpiola grisea	Harpiola grisea		[MSW2] Subgenus Harpiola.; [MSW3] Subgenus Harpiola. Reviewed by Bates and Harrison (1997) and Bhattacharyya (2002).; [HMW] Murina grisea Peters, 1872 , Jeripanee, 5500 ft. (= 1676 m ), Mussooree, Dehra Dun, Uttar Pradesh , India . Traditionally considered a subgenus of Murina , Harpiola is now tentatively treated as a distinct genus, based on morphology, following the discovery of more specimens of H. grisea in Mizoram and of a second species, H. isodon , in Taiwan and Vietnam ; nevertheless, Harpiola is phylogenetically imbedded within Murina and may be best retained there; further study needed. The Mizoram specimens resemble H. isodon in fur coloration; further sampling throughout both species’ ranges is needed to understand the full variability. Monotypic.; [batnames2022] Formerly included in Murina as subgenus Harpiola; see Kuo, Fnag, Csorba, and Lee (2006) . Reviewed by Bates and Harrison(1997) and Bhattacharyya (2002).; [MDD2022] moved from Murina to Harpiola; [IUCN] <p>Although Harpiola is considered as a subgenus of the genus Murina by Simmons (2005), Kuo et al. (2006) resurrected its generic status after examining a series of specimens collected from Taiwan.</p>; [batnames2023] Formerly included in Murina as subgenus Harpiola; see Kuo, Fnag, Csorba, and Lee (2006) . Reviewed by Bates and Harrison(1997) and Bhattacharyya (2002).; [MDD2023] moved from Murina to Harpiola; [MDD2025_2.0] moved from Murina to Harpiola; [batnames2025_1.7] Formerly included in Murina as subgenus Harpiola; see Kuo, Fnag, Csorba, and Lee (2006). Reviewed by Bates and Harrison(1997) and Bhattacharyya (2002).; [MDD2025_2.2] moved from Murina to Harpiola														grisea	<p>Although Harpiola is considered as a subgenus of the genus Murina by Simmons (2005), Kuo et al. (2006) resurrected its generic status after examining a series of specimens collected from Taiwan.</p>			grisea 	grisea 			grisea (W. C. H. Peters, 1872)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Peters' tube-nosed bat	NW India	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Murina grisea	India, Uttar Pradesh, Dehra Dun, Mussooree, Jeripanee, 5500 ft. (1676 m).	Peters	1872	Monatsb. Preuss. Akad. Wiss. Berlin, p. 258.	Distribution: Same as for subgenus.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Peters' tube-nosed bat	NW India	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Peters	1872	Monatsb. K. Preuss. Akad. Wiss. Berlin, 1872:258.	Subgenus Harpiola.	NW Himalayas.	India, Uttar Pradesh, Dehra Dun, Mussooree, Jeripanee, 5,500 ft. (1,676 m).		PETERS	1872	Size fairly small (forearm length, 33 mm).	Distribution: Same as for subgenus.	No subspecies.		133	species	M. grisea	PETERS	1872	Harpiola	subgenus	Murina grisea				Size fairly small (forearm length, 33 mm).	No subspecies.		14. M. grisea PETERS 1872.	14	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Murininae		Murina grisea	Murina	Harpiola	grisea	Peters		1872		Monatsb. K. Preuss. Akad. Wiss. Berlin	1872		258		Peters's Tube-nosed Bat	India, Uttar Pradesh, Dehra Dun, Mussooree, Jeripanee, 5,500 ft. (1,676 m).	NW Himalayas, Mizoram (India).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Endangered.		Subgenus Harpiola. Reviewed by Bates and Harrison (1997) and Bhattacharyya (2002).	4C3D87E8FF696AD6FA9A97F61611BCD1	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	907	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF696AD6FA9A97F61611BCD1.xml	Harpiola grisea	Vespertilionidae	Harpiola	grisea	Peters	1872	Murine de @fr | 'Himalaya @en | Graue Rohrennase @de | Ratonero del Himalaya @es | Peters's Tube-nosed Bat @en	Murina grisea Peters, 1872 , Jeripanee, 5500 ft. (= 1676 m ), Mussooree, Dehra Dun, Uttar Pradesh , India . Traditionally considered a subgenus of Murina , Harpiola is now tentatively treated as a distinct genus, based on morphology, following the discovery of more specimens of H. grisea in Mizoram and of a second species, H. isodon , in Taiwan and Vietnam ; nevertheless, Harpiola is phylogenetically imbedded within Murina and may be best retained there; further study needed. The Mizoram specimens resemble H. isodon in fur coloration; further sampling throughout both species’ ranges is needed to understand the full variability. Monotypic.	N & NE India ( Uttarakhand and Mizoram ).	Head-body c. 45 mm , tail 25 mm , ear 12- 1 mm , hindfoot 6 mm , forearm 32-4-32- 8 mm . Dorsal pelage is dark grayish brown (hairs with yellowish-brown tips), ventral pelage is lighter and grayer (hairs with ashy gray tips). Ears have narrowly rounded tip and distinct notch on posterior border. Skull is small, with narrow, shallow rostrum; braincase is elevated above rostrum with distinctly higher mid-portion than the lambda; I? is bicuspid and enlarged; P* and P, are well developed and subequalto the canines and second premolars of the corresponding tooth row; C,is strongly bifid; distinguished from the Formosan Tube-nosed Bat ( H. isodon ) by its relatively narrow P*, being almost as long as wide, M' with no postcingular platform, and M* without any trace of a mesostyle. Dental formula for both species of Harpiolais12/3,C1/1,P 2/2, M 3/3 (x2) = 34.	Two Gray Tube-nosed Bats were caught over a small pool in dense forest with thick undergrowth; the speciesis also known from montane forest. Recorded at 1367-1692 m .	No information.	No information.	No information.	No information.	Classified as Data Deficient on The IUCN Red List. Known only from a few specimens from Mizoram and the type locality. Agricultural expansion and human settlements may be threats.	Bates & Harrison (1997) | Bhattacharyya (2002) | Csorba, Molur, Srinivasulu & Chakravarty (2016) | Kuo Haochih et al. (2006) | Molur etal. (2002) | Thomas (1915d)	https://zenodo.org/record/6398551/files/figure.png	325. Gray Tube-nosed Bat Harpiola grisea French: Murine de I'Himalaya / German: Graue Rohrennase / Spanish: Ratonero del Himalaya Other common names: Peters's Tube-nosed Bat Taxonomy. Murina grisea Peters, 1872 , Jeripanee, 5500 ft. (= 1676 m ), Mussooree, Dehra Dun, Uttar Pradesh , India . Traditionally considered a subgenus of Murina , Harpiola is now tentatively treated as a distinct genus, based on morphology, following the discovery of more specimens of H. grisea in Mizoram and of a second species, H. isodon , in Taiwan and Vietnam ; nevertheless, Harpiola is phylogenetically imbedded within Murina and may be best retained there; further study needed. The Mizoram specimens resemble H. isodon in fur coloration; further sampling throughout both species’ ranges is needed to understand the full variability. Monotypic. Distribution. N & NE India ( Uttarakhand and Mizoram ). Descriptive notes. Head-body c. 45 mm , tail 25 mm , ear 12- 1 mm , hindfoot 6 mm , forearm 32-4-32- 8 mm . Dorsal pelage is dark grayish brown (hairs with yellowish-brown tips), ventral pelage is lighter and grayer (hairs with ashy gray tips). Ears have narrowly rounded tip and distinct notch on posterior border. Skull is small, with narrow, shallow rostrum; braincase is elevated above rostrum with distinctly higher mid-portion than the lambda; I? is bicuspid and enlarged; P* and P, are well developed and subequalto the canines and second premolars of the corresponding tooth row; C,is strongly bifid; distinguished from the Formosan Tube-nosed Bat ( H. isodon ) by its relatively narrow P*, being almost as long as wide, M' with no postcingular platform, and M* without any trace of a mesostyle. Dental formula for both species of Harpiolais12/3,C1/1,P 2/2, M 3/3 (x2) = 34. Habitat. Two Gray Tube-nosed Bats were caught over a small pool in dense forest with thick undergrowth; the speciesis also known from montane forest. Recorded at 1367-1692 m . Food and Feeding. No information. Breeding. No information. Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The IUCN Red List. Known only from a few specimens from Mizoram and the type locality. Agricultural expansion and human settlements may be threats. Bibliography. Bates & Harrison (1997), Bhattacharyya (2002), Csorba, Molur, Srinivasulu & Chakravarty (2016), Kuo Haochih et al. (2006), Molur etal. (2002), Thomas (1915d).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Harpiola grisea	Harpiola	Harpiola	grisea	Peters	1872	0	Monatsb. K. Preuss. Akad. Wiss. Berlin	1876:18:00	Peters' Tube-nosed Bat	None.	India, Uttar Pradesh, Dehra Dun, Mussooree, Jeripanee, 5,500 ft. (1,676 m).	NW Himalayas, Mizoram (India).	Not listed.	Data Deficient	Formerly included in Murina as subgenus Harpiola; see Kuo, Fnag, Csorba, and Lee (2006) . Reviewed by Bates and Harrison(1997) and Bhattacharyya (2002).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Harpiola grisea	23	Gray Tube-nosed Bat	Peters's Tube-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MURININAE	NA	Harpiola	NA	grisea	W. Peters	1872	1	Murina_grisea	Peters, W. C. H. (1872). Ãœber neue Flederthiere (Phyllorhina micropus, Harpyiocephalus Huttonii, Murina grisea, Vesperugo micropus, Vesperus (Marsipolaemus) albigularis, Vesperus propinquus, tenuipinnis). Monatsberichte der Königlich Preussischen Akademie der Wissenschaften zu Berlin, 1872, 258.	https://www.biodiversitylibrary.org/item/110470#page/294/mode/1up	BM 1879.11.21.117		Jeripanee, 5500 ft. (= 1676 m), Mussooree, Dehra Dun, Uttar Pradesh, India.			grisea (W. Peters, 1872)	moved from Murina to Harpiola	Kuo, H. C., Fang, Y. P., Csorba, G., & Lee, L. L. (2006). The definition of Harpiola (Vespertilionidae: Murininae) and the description of a new species from Taiwan. Acta Chiropterologica, 8(1), 11-19.	India	Asia	Indomalaya	DD	0	0	0	Harpiola_grisea	0	manual	Murina_grisea	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	13941	Harpiola grisea	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Harpiola	grisea	(Peters, 1872)	<p>Although Harpiola is considered as a subgenus of the genus Murina by Simmons (2005), Kuo et al. (2006) resurrected its generic status after examining a series of specimens collected from Taiwan.</p>	20000000	Harpiola grisea	Data Deficient		2016	2015-12-26 00:00:00 UTC	3.1	English	this species is assessed as Data Deficient in view of the absence of recent information on its extent of occurrence, ecological requirements, threats and conservation status.	<p><span lang="EN-US">Little is known about the habitat or ecology of this species except that it has been recorded in thick montane forests (Molur et al. 2002). Bhattacharya (2002) caught two individuals at a small pool of water in dense forest with thick undergrowth.<a name="_GoBack"></a></p>	<p><span lang="EN-US">This species is likely threatened by deforestation, generally resulting from logging operations and the conversion of land for human settlements and tourism in Mussourie (Molur et al . 2002). The threats affecting the species in Mizoram are not known (Bhattacharya 2002). Diversion of forest land for shifting cultivation and conversion to oil palm plantations is on the rise and may be a potential threat (Chakravarty pers. comm).</p>	The abundance, population size and trends for this species are not known (Molur et al. 2002).	Unknown	<p><span lang="EN-US">This species is endemic to India and has been recorded from Sairep in Lunglei district of Mizoram and Jharipani near Mussourie, Uttarakhand (Bhattacharyya 2002, Molur et al . 2002). It has been recorded from elevations above 1,360 m asl. The area of occupancy is not estimated as there is limited information on the species distribution and taxonomy (Molur et al. 2002).</p>		Terrestrial	<p><span lang="EN-US">There are no conservation measures in place, and the species has not been reported from any protected area as yet, although suitable habitats exists at Benog Wildlife Sancutary in Mussourie, Phawngpui National Park and Ngenpui and Khawnglung Wildlife Sanctuaries in the vicinity of Lunglei district (Chakravorty pers. comm). It has not been recorded again from the type locality, Widespread surveys are urgently recommended to identify existing populations which need to be followed up with taxonomic, ecological and population studies (Molur et al. 2002).</p>	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Harpiola	Harpiola	grisea	Hutton	1872	0	In Peters Monatsb. K. Preuss. Akad. Wiss. Berlin	1876:18:00	Peters' Tube-nosed Bat	None.	India, Uttar Pradesh, Dehra Dun, Mussooree, Jeripanee, 5,500 ft. (1,676 m).	NW Himalayas, Mizoram (India).	Not listed.	Data Deficient	Formerly included in Murina as subgenus Harpiola; see Kuo, Fnag, Csorba, and Lee (2006) . Reviewed by Bates and Harrison(1997) and Bhattacharyya (2002).	Harpiola grisea	1005315	23	Gray Tube-nosed Bat	Peters's Tube-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MURININAE	NA	Harpiola	NA	grisea	W. Peters	1872	1	Murina_grisea	Peters, W. C. H. (1872). Ãœber neue Flederthiere (Phyllorhina micropus, Harpyiocephalus Huttonii, Murina grisea, Vesperugo micropus, Vesperus (Marsipolaemus) albigularis, Vesperus propinquus, tenuipinnis). Monatsberichte der Königlich Preussischen Akademie der Wissenschaften zu Berlin, 1872, 258.	https://www.biodiversitylibrary.org/item/110470#page/294/mode/1up	BM 1879.11.21.117		Jeripanee, 5500 ft. (= 1676 m), Mussooree, Dehra Dun, Uttar Pradesh, India.			grisea (W. Peters, 1872)	moved from Murina to Harpiola	Kuo, H. C., Fang, Y. P., Csorba, G., & Lee, L. L. (2006). The definition of Harpiola (Vespertilionidae: Murininae) and the description of a new species from Taiwan. Acta Chiropterologica, 8(1), 11-19.				India	Asia	Indomalaya	DD	0	0	0	Harpiola_grisea	0	manual	Murina_grisea	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Harpiola_grisea	1005315	23	Gray Tube-nosed Bat	Peters's Tube-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Murininae	NA	Harpiola	NA	grisea	W. C. H. Peters	1	Murina grisea	Peters, W.C.H. 1872. Ãœber neue Flederthiere (_Phyllorhina micropus_, _Harpyiocephalus Huttonii_, _Murina grisea_, _Vesperugo micropus_, _Vesperus_ (_Marsipolaemus_) _albigularis_, _Vesperus propinquus_, _tenuipinnis_). Monatsberichte der Königlichen Preussischen Akademie der Wissenschaften zu Berlin 1872:256-264.	https://www.biodiversitylibrary.org/page/35721188	BMNH:Mamm:1879.11.21.117	holotype	https://data.nhm.ac.uk/object/cd1cc17e-dcb7-4d32-b6e5-17b93d061a67	Jeripanee, 5500 ft. (= 1676 m), Mussooree, Dehra Dun, Uttar Pradesh, India.			moved from Murina to Harpiola	Kuo, H. C., Fang, Y. P., Csorba, G., & Lee, L. L. (2006). The definition of Harpiola (Vespertilionidae: Murininae) and the description of a new species from Taiwan. Acta Chiropterologica, 8(1), 11-19.				India	Asia	Indomalaya	DD	0	0	0	Harpiola_grisea	0	manual	Murina_grisea	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Harpiola	Harpiola	grisea	Hutton	1872	0	In Peters Monatsb. K. Preuss. Akad. Wiss. Berlin	1876:18:00	Peters' Tube-nosed Bat	None.	India, Uttar Pradesh, Dehra Dun, Mussooree, Jeripanee, 5,500 ft. (1,676 m).	NW Himalayas, Mizoram (India).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/13941/22093890/' target='_blank'>Data Deficient</a>	Formerly included in Murina as subgenus Harpiola; see Kuo, Fnag, Csorba, and Lee (2006). Reviewed by Bates and Harrison(1997) and Bhattacharyya (2002).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Murina grisea; Harpiola grisea; Harpiola grisea; Harpiola grisea; Harpiola grisea; Harpiola grisea; grisea; Murine de; 'Himalaya; Graue Rohrennase; Ratonero del Himalaya; Peters's Tube-nosed Bat; Gray Tube-nosed Bat; Peters's Tube-nosed Bat; Peters's Tube-nosed Bat; Peters' Tube-nosed Bat; H. grisea