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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L354	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Harpiocephalus harpia	Harpiocephalus harpia	Harpiocephalus harpia	Harpiocephalus harpia	Harpiocephalus harpia	Harpiocephalus harpia	Harpiocephalus harpia	Harpiocephalus harpia	Harpiocephalus harpia	Harpiocephalus harpia	Harpiocephalus harpia	Harpiocephalus harpia	Harpiocephalus harpia	Harpiocephalus harpia	Harpiocephalus harpia		[MSW2] Includes mordax, which may be a separate species; see Hill and Francis (1984).; [MSW3] Does not include mordax; see Hill and Francis (1984) Corbet and Hill (1992), and Hendrichsen et al. (2002b). Reviewed in part by Bates and Harrison (1997) and Hendrichsen et al. (2001b). The Taiwan record, if valid, has not been allocated to subspecies.; [HMW] Vespertilio harpia Temminck, 1840 , “ I'lle de Java [ Indonesia ] ... les caverns de la partie sud du Volcan de Guédé [= caves of the southern part of Mount Gede].” Based on lectotype selection, restricted by A. M. Husson in 1955 10.*S.E. side of Mount Gede, W. Java .” Genetically, Harpiocephalus appears to be sister to the rest of Murininae or imbedded within Murina and related to M. kontumensis, or to M. leucogaster and M. shuipuensis ; exact placement currently uncertain. The form mordax is a synonym; it has sometimes been considered a distinct species based on differing skull size, but this difference actually represents sexual dimorphism, the name mordax having been applied to the larger females. Four subspecies recognized.; [batnames2022] Includes mordax ; see Matveev (2005). Reviewed in part by Bates and Harrison (1997) and Hendrichsen et al. (2001 b ). The Taiwan record, if valid, has not been allocated to subspecies.; [MDD2022] includes mordax as a synonym; [IUCN] Earlier the taxon ;madrassius ;Thomas, 1923 was considered as a subspecies of this taxon (Ellerman and Morrison-Scott 1951), but has been synonymized with ;Harpiocephalus harpia lasyurus ;(Hodgson, 1847) (Das 1986; Bates and Harrison 1997). Simmons (2005), however, recognizes ;madrassius ;Thomas, 1923 as valid subspecies (Srinivasuluet al. ;in press). ;Also, although frequently accepted as a separate species (Corbet and Hill 1992, Simmons 2005), Matveev (2005) revealed that differences between ;H. harpia ;and ;H. mordax ; are only cases of extreme sexual dimorphism where male and female specimens differ in size and cranial shape. Molecular markers clearly demonstrated that the studied male and female specimens (thought as representatives of ;H. harpia ;and ;H. mordax , respectively) are conspecifics. This well expressed sexual dimorphism is further evidenced by Chen ;et al. ;(2015) and Son ;et al. ; (2016).; [MDD2023] includes mordax as a synonym; [MDD2025_2.0] includes mordax as a synonym; [batnames2025_1.7] Includes mordax; see Matveev (2005). Reviewed in part by Bates and Harrison (1997) and Hendrichsen et al. (2001b). The Taiwan record, if valid, has not been allocated to subspecies. See also Lin et al. (2006) and Son et al. (2015).; [MDD2025_2.2] includes mordax as a synonym						lasyurus, madrassius, mordax, pearsonii, rufulus, rufus.	madrassius, lasyurus, rufulus, harpia	harpia, lasyurus, madrassius, rufulus	pearsonii, rufus	harpia, lasyurus, madrassius, rufulus		harpia, lasyurus, madrassius, rufulus	harpia - pearsonii, rufus	harpia, rufus, lasyurus, pearsonii, rufulus, madrassius, mordax	Earlier the taxon ;madrassius ;Thomas, 1923 was considered as a subspecies of this taxon (Ellerman and Morrison-Scott 1951), but has been synonymized with ;Harpiocephalus harpia lasyurus ;(Hodgson, 1847) (Das 1986; Bates and Harrison 1997). Simmons (2005), however, recognizes ;madrassius ;Thomas, 1923 as valid subspecies (Srinivasuluet al. ;in press). ;Also, although frequently accepted as a separate species (Corbet and Hill 1992, Simmons 2005), Matveev (2005) revealed that differences between ;H. harpia ;and ;H. mordax ; are only cases of extreme sexual dimorphism where male and female specimens differ in size and cranial shape. Molecular markers clearly demonstrated that the studied male and female specimens (thought as representatives of ;H. harpia ;and ;H. mordax , respectively) are conspecifics. This well expressed sexual dimorphism is further evidenced by Chen ;et al. ;(2015) and Son ;et al. ; (2016).	harpia, lasyurus, madrassius, rufulus	harpia - mordax, pearsonii, rufus	harpia, rufus, lasyurus, pearsonii, rufulus, madrassius, mordax 	harpia, cavernarum, rufus, harpyia, lasyurus, pearsonii, pearsoni, pearsonii, rufulus, madrassius, mordax 	harpia, lasyurus, madrassius, rufulus	harpia - pearsonii, rufus	harpia (Temminck, 1840)|cavernarum (Lesson, 1842) [nomen novum]|rufus J. E. Gray, 1842 [nomen novum]|harpyia (H. R. Schinz, 1844) [incorrect subsequent spelling]|lasyurus (B. H. Hodgson, 1847)|pearsonii (Horsfield, 1851)|pearsoni (E. Blyth, 1855) [incorrect subsequent spelling]|pearsonii (Tomes, 1858) [preoccupied]|lasiura (Fitzinger, 1870) [incorrect subsequent spelling]|rufulus G. M. Allen, 1913|madrassius O. Thomas, 1923|mordax O. Thomas, 1923		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Hairy-winged bat	India – Vietnam, Sumatra, Java, S Moluccas	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Harpiocephalus harpia	Indonesia, Java, Mt. Gede.	Temminck	1840	Monogr. Mamm., 2:219.	Distribution: Essentially same as for genus.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Hairy-winged bat	India – Vietnam, Taiwan, Sumatra, Java, Borneo, S Moluccas	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Temminck	1840	Monogr. Mamm., 2:219.	Includes mordax, which may be a separate species; see Hill and Francis (1984).	India to Taiwan and Vietnam, south to Molucca Isis, Java, and Lesser Sunda Isis.	Indonesia, Java, Mt. Gede.		TEMMINCK	1840	Muzzle not greatly broadened nor zygomata greatly expanded. Incisors and canines not enlarged. Size fairly large (forearm length, 43-50 mm).	Distribution: Essentially same as for genus.	Four subspecies are currently recognized:	H. h. madrassius (southern India), H. h. lasyurus (northeastern India), H. h. rufulus (Thailand, Vietnam), H. h. harpia (Sumatra, Borneo, Java, Moluccas). The Taiwan record, if valid, has not been allocated subspecifically.	133	species	H. harpia	TEMMINCK	1840	Harpiocephalus	genus	Harpiocephalus harpia				Muzzle not greatly broadened nor zygomata greatly expanded. Incisors and canines not enlarged. Size fairly large (forearm length, 43-50 mm).	Four subspecies are currently recognized:		1. H. harpia (TEMMINCK 1840).	1	_H. h. harpia_ (Temminck, 1840) (synonyms: _cavernarum_ (Lesson, 1842), _pearsonii_ (Horsfield, 1851), _pearsonii_ (Tomes, 1858), _rufus_ Gray, 1842); _H. h. lasyurus_ (Hodgson, 1847); _H. h. madrassius_ Thomas, 1923; _H. h. rufulus_ Allen, 1913			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Murininae		Harpiocephalus harpia	Harpiocephalus		harpia	Temminck	y	1840		Monogr. Mamm.	2		219		Lesser Hairy-winged Bat	Indonesia, Java, NE side of Mt. Gede.	S and NE India, S China, Taiwan, Laos and Vietnam, Sumatra, Java, Borneo, S Moluccas, and the Philippines.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	pearsonii Horsfield, 1851; rufus Gray, 1842; lasyurus Hodgson, 1847; madrassius Thomas, 1923; rufulus G. M. Allen, 1913.	Does not include mordax; see Hill and Francis (1984) Corbet and Hill (1992), and Hendrichsen et al. (2002b). Reviewed in part by Bates and Harrison (1997) and Hendrichsen et al. (2001b). The Taiwan record, if valid, has not been allocated to subspecies.	4C3D87E8FF696AD6FF9794591B20B7B6	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	907	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF696AD6FF9794591B20B7B6.xml	Harpiocephalus harpia	Vespertilionidae	Harpiocephalus	harpia		1840	Murine a ailes velues @fr | Haarfllgelfledermaus @de | Ratonero de alas pilosas @es | Lesser Hairy-winged Bat @en	Vespertilio harpia Temminck, 1840 , “ I'lle de Java [ Indonesia ] ... les caverns de la partie sud du Volcan de Guédé [= caves of the southern part of Mount Gede].” Based on lectotype selection, restricted by A. M. Husson in 1955 10.*S.E. side of Mount Gede, W. Java .” Genetically, Harpiocephalus appears to be sister to the rest of Murininae or imbedded within Murina and related to M. kontumensis, or to M. leucogaster and M. shuipuensis ; exact placement currently uncertain. The form mordax is a synonym; it has sometimes been considered a distinct species based on differing skull size, but this difference actually represents sexual dimorphism, the name mordax having been applied to the larger females. Four subspecies recognized.	H.h. harpia Temminck,1840—MalayPeninsula,Sumatra,Java,Bali,Lombok,andthePhilippines(Luzon,Leyte,Negros,Mindanao,andPalawan). H.h. lasyurus Hodgson,1847—NEIndia(Sikkim,WestBengal,Assam,Meghalaya,andMizoram),Bhutan,andNWMyanmar. H.h. madrassius Thomas,1923—SIndia(KeralaandTamilNadu). H. h. rufulus G. M. Allen, 1913 — SC & SE China ( Yunnan , Guangdong , and Fujian ), Vietnam (including Cat Ba I), N Laos , C Myanmar , and Thailand . Also Taiwan (subspecies as yet undetermined).	Head—body 57-75 mm , tail 40-50 mm , ear 15-18 mm , hindfoot 10-14 mm , forearm 44-54- 9 mm ; weight 12-30- 2 g . Males are distinctly smaller than females. Fur is thick and soft; dorsally bright orange (with dark-based hairs) to dull orangish brown (with gray-based hairs), with bright orange individuals commoner in Malaysia and the Philippines ; ventrally lighter and grayer. Dorsal pelage extends densely onto wing bases, legs, and feet, and entire uropatagium and rest of wings and forearmsis lightly haired. Entire face is furred except long, protuberant nostrils (similar to those of Murina ); ears and face are dark brown whereas membranes are blackish brown. Ears are evenly rounded on anterior and posterior borders and generally funnel-shaped; tragus is long, pointed, slightly convex anteriorly, concave on posterior border, and with posterior basal notch. Baculum is forked distally a little before midpoint, the two prongs resembling curved plier handles, being broadest in the middle; main shaft is short with a dorsal ridge and ventral groove. Skull is robust with short, broad, deep rostrum; zygomata are long and relatively strong, with a slight jugal bone; lambdoidal crests are well developed; tympanic bullae and cochleae are small; basioccipital pits are shallow; dentition is robust with massive canines; upper molars are modified and reduced, essentially missing mesostyles, the metacone being the largest cusp; M” is extremely small, being represented by a small “scalelike” tooth that lies against M?; talonids of all three lower molars are reduced. Dental formulais12/3,C1/1,P 2/2, M 3/3 (x2) = 34. Chromosomal complement has 2n = 44 and FN = 52 ( Vietnam , Taiwan , and Guangdong , China ) and apparently n = 40 ( Thailand ).	Understory of lowland dipterocarp and hill forests and some disturbed areas in South-east Asia; also montane forests and valleys with tall trees near water in South Asia. In the Philippines , the species occurs in primary and disturbed lowland, montane, and mossy forests. Recorded at elevations of 300-2480 m in the Philippines , and 185-1538 m in India .	Hairy-winged Bats are known to feed on beetles.	No information.	Call shape is a steep FM sweep with start frequency of 111-5 kHz, end frequency 32-5 kHz, peak frequency 57 kHz, and duration 1 millisecond, based on single recording in Vietnam . Peak frequency of 78-3 kHz was recorded in Guangdong , China .	No information.	Classified as Least Concern on The IUCN Red List. The Hairywinged Bat is widespread butis suffering habitat destruction from agricultural expansion and deforestation throughout its range.	Bates & Harrison (1997) | Chen Bocheng etal. (2015) | Csorba, Bumrungsri, Francis, Bates, Rosell-Ambal et al. (2008) | Das (1986b) | Francis (2008a) | Francis et al. (2010) | Heaney, Balete, Dolar et al. (1998) | Heaney, Balete & Rickart (2016) | Hill & Francis (1984) | Husson (1955) | Kruskop (2013a) | Lin Liangkong et al. (2006) | Matveev (2005) | McBee etal. (1986) | Nguyen Truong Son etal. (2015) | Smith & Xie Yan (2008) | Zhou Quan et al. (2014)	https://zenodo.org/record/6398549/files/figure.png	324. Hairy-winged Bat Harpiocephalus harpia French: Murine a ailes velues / German: Haarfllgelfledermaus / Spanish: Ratonero de alas pilosas Other common names: Lesser Hairy-winged Bat Taxonomy. Vespertilio harpia Temminck, 1840 , “ I'lle de Java [ Indonesia ] ... les caverns de la partie sud du Volcan de Guédé [= caves of the southern part of Mount Gede].” Based on lectotype selection, restricted by A. M. Husson in 1955 10.*S.E. side of Mount Gede, W. Java .” Genetically, Harpiocephalus appears to be sister to the rest of Murininae or imbedded within Murina and related to M. kontumensis, or to M. leucogaster and M. shuipuensis ; exact placement currently uncertain. The form mordax is a synonym; it has sometimes been considered a distinct species based on differing skull size, but this difference actually represents sexual dimorphism, the name mordax having been applied to the larger females. Four subspecies recognized. Subspecies and Distribution. H.h.harpiaTemminck,1840—MalayPeninsula,Sumatra,Java,Bali,Lombok,andthePhilippines(Luzon,Leyte,Negros,Mindanao,andPalawan). H.h.lasyurusHodgson,1847—NEIndia(Sikkim,WestBengal,Assam,Meghalaya,andMizoram),Bhutan,andNWMyanmar. H.h.madrassiusThomas,1923—SIndia(KeralaandTamilNadu). H. h. rufulus G. M. Allen, 1913 — SC & SE China ( Yunnan , Guangdong , and Fujian ), Vietnam (including Cat Ba I), N Laos , C Myanmar , and Thailand . Also Taiwan (subspecies as yet undetermined). Descriptive notes. Head—body 57-75 mm , tail 40-50 mm , ear 15-18 mm , hindfoot 10-14 mm , forearm 44-54- 9 mm ; weight 12-30- 2 g . Males are distinctly smaller than females. Fur is thick and soft; dorsally bright orange (with dark-based hairs) to dull orangish brown (with gray-based hairs), with bright orange individuals commoner in Malaysia and the Philippines ; ventrally lighter and grayer. Dorsal pelage extends densely onto wing bases, legs, and feet, and entire uropatagium and rest of wings and forearmsis lightly haired. Entire face is furred except long, protuberant nostrils (similar to those of Murina ); ears and face are dark brown whereas membranes are blackish brown. Ears are evenly rounded on anterior and posterior borders and generally funnel-shaped; tragus is long, pointed, slightly convex anteriorly, concave on posterior border, and with posterior basal notch. Baculum is forked distally a little before midpoint, the two prongs resembling curved plier handles, being broadest in the middle; main shaft is short with a dorsal ridge and ventral groove. Skull is robust with short, broad, deep rostrum; zygomata are long and relatively strong, with a slight jugal bone; lambdoidal crests are well developed; tympanic bullae and cochleae are small; basioccipital pits are shallow; dentition is robust with massive canines; upper molars are modified and reduced, essentially missing mesostyles, the metacone being the largest cusp; M” is extremely small, being represented by a small “scalelike” tooth that lies against M?; talonids of all three lower molars are reduced. Dental formulais12/3,C1/1,P 2/2, M 3/3 (x2) = 34. Chromosomal complement has 2n = 44 and FN = 52 ( Vietnam , Taiwan , and Guangdong , China ) and apparently n = 40 ( Thailand ). Habitat. Understory of lowland dipterocarp and hill forests and some disturbed areas in South-east Asia; also montane forests and valleys with tall trees near water in South Asia. In the Philippines , the species occurs in primary and disturbed lowland, montane, and mossy forests. Recorded at elevations of 300-2480 m in the Philippines , and 185-1538 m in India . Food and Feeding. Hairy-winged Bats are known to feed on beetles. Breeding. No information. Activity patterns. Call shape is a steep FM sweep with start frequency of 111-5 kHz, end frequency 32-5 kHz, peak frequency 57 kHz, and duration 1 millisecond, based on single recording in Vietnam . Peak frequency of 78-3 kHz was recorded in Guangdong , China . Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Hairywinged Bat is widespread butis suffering habitat destruction from agricultural expansion and deforestation throughout its range. Bibliography. Bates & Harrison (1997), Chen Bocheng etal. (2015), Csorba, Bumrungsri, Francis, Bates, Rosell-Ambal et al. (2008), Das (1986b), Francis (2008a), Francis et al. (2010), Heaney, Balete, Dolar et al. (1998), Heaney, Balete & Rickart (2016), Hill & Francis (1984), Husson (1955), Kruskop (2013a), Lin Liangkong et al. (2006), Matveev (2005), McBee etal. (1986), Nguyen Truong Son etal. (2015), Smith & Xie Yan (2008), Zhou Quan et al. (2014).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Harpiocephalus harpia	Harpiocephalus		harpia	Temminck	1840	1	Monogr. Mamm.	0.2354	Lesser Hairy-winged Bat	 pearsonii Horsfield, 1851; rufus Gray, 1842; <b>lasyurus</b> Hodgson, 1847; <b> madrassius </b> Thomas, 1923; <b>rufulus</b> G. M. Allen, 1913.	Indonesia, Java, NE side of Mt. Gede	S and NE India, Myanmar, S China, Taiwan, Laos, Thailand, Cambodia, Vietnam, Sumatra, Java, Borneo, S Moluccas, and the Philippines.	Not listed.	Least Concern	Includes mordax ; see Matveev (2005). Reviewed in part by Bates and Harrison (1997) and Hendrichsen et al. (2001 b ). The Taiwan record, if valid, has not been allocated to subspecies.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Harpiocephalus harpia	23	Hairy-winged Bat	Lesser Hairy-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MURININAE	NA	Harpiocephalus	NA	harpia	Temminck	1840	1						"l'ÃŽle de Java [Indonesia] … les caverns de la partie sud du Volcan de Guédé [= caves of the southern part of Mount Gede]." Based on lectotype selection, restricted by A. M. Husson in 1955 to "S.E. side of Mount Gede, W. Java."			harpia (Temminck, 1840)|rufus J. E. Gray, 1842|lasyurus (Hodgson, 1847)|pearsonii (Horsfield, 1851)|rufulus G. M. Allen, 1913|madrassius O. Thomas, 1923|mordax O. Thomas, 1923	includes mordax as a synonym	Son, N. T., Csorba, G., Tu, V. T., Thong, V. D., Wu, Y., Harada, M., ... & Motokawa, M. (2015). A new species of the genus Murina (Chiroptera: Vespertilionidae) from the Central Highlands of Vietnam with a review of the subfamily Murininae in Vietnam. Acta Chiropterologica, 17(2), 201-232.	India|Bhutan|Myanmar|China|Taiwan|Laos|Vietnam|Thailand|Malaysia|Indonesia|Philippines	Asia|Oceania	Palearctic|Indomalaya|Oceania	LC	0	0	0	Harpiocephalus_harpia	0	sciname match	Harpiocephalus_harpia	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	100000000	Harpiocephalus harpia	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Harpiocephalus	harpia	(Temminck, 1840)	Earlier the taxon ;madrassius ;Thomas, 1923 was considered as a subspecies of this taxon (Ellerman and Morrison-Scott 1951), but has been synonymized with ;Harpiocephalus harpia lasyurus ;(Hodgson, 1847) (Das 1986; Bates and Harrison 1997). Simmons (2005), however, recognizes ;madrassius ;Thomas, 1923 as valid subspecies (Srinivasuluet al. ;in press). ;Also, although frequently accepted as a separate species (Corbet and Hill 1992, Simmons 2005), Matveev (2005) revealed that differences between ;H. harpia ;and ;H. mordax ; are only cases of extreme sexual dimorphism where male and female specimens differ in size and cranial shape. Molecular markers clearly demonstrated that the studied male and female specimens (thought as representatives of ;H. harpia ;and ;H. mordax , respectively) are conspecifics. This well expressed sexual dimorphism is further evidenced by Chen ;et al. ;(2015) and Son ;et al. ; (2016).	20000000	Harpiocephalus harpia	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	In South Asia, this species is found in montane forests and valleys with tall trees in the vicinity of water (Molur et al . 2002). It feeds on beetles (Bates and Harrison 1997). There is little information on the natural history of this species in China (Smith and Xie 2008). It has been recorded only from tall and hill forest in Peninsula Malaysia. In Philippines, the species is known from primary and disturbed lowland forest (Ingle and Heaney 1992; Rickart et al. 1993) and is believed to probably be dependent on lowland forest.	The species throughout its range is threatened by habitat loss, largely through commercial logging and the conversion of land to agricultural use.	The abundance, population size and trends for this species are not well known. It has not been collected in Kerala (southern India) in surveys between 1990 and 1999 (Molur et al . 2002).	Decreasing	This widespread species has been recorded from southern South Asia, through parts of southern China to insular Southeast Asia. In South Asia, this species is presently known from India (Assam, Kerala, Meghalaya, Mizoram, Sikkim, Tamil Nadu and West Bengal), recorded from 475 to 750 m asl (Molur et al. 2002). This taxon may probably also occur in Bhutan (Molur et al. 2002). In China, it has been recorded from Yunnan, Guangdong, Fujian, and is present on the island of Taiwan (Smith and Xie 2008). In Southeast Asia, it has been recorded from a few scattered localities on the mainland, and from Indonesia (the islands of Sumatra, Java, Bali, Lombok and Ambon) and the Philippines. In the Philippines, it has only been recorded from the islands of Luzon Leyte, Negros, and Panay (Heaney et al. 1998).		Terrestrial	In South Asia, there are no conservation measures in place and the species has not been recorded from any protected areas. Surveys, ecological studies, population monitoring are recommended for this species in South Asia (Molur et al. 2002). It has been recorded from a number of protected areas in Southeast Asia.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 																	Harpiocephalus harpia	1005314	23	Hairy-winged Bat	Lesser Hairy-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MURININAE	NA	Harpiocephalus	NA	harpia	Temminck	1840	1						"l'ÃŽle de Java [Indonesia] … les caverns de la partie sud du Volcan de Guédé [= caves of the southern part of Mount Gede]." Based on lectotype selection, restricted by A. M. Husson in 1955 to "S.E. side of Mount Gede, W. Java."			harpia (Temminck, 1840)|rufus J. E. Gray, 1842|lasyurus (Hodgson, 1847)|pearsonii (Horsfield, 1851)|rufulus G. M. Allen, 1913|madrassius O. Thomas, 1923|mordax O. Thomas, 1923	includes mordax as a synonym	Son, N. T., Csorba, G., Tu, V. T., Thong, V. D., Wu, Y., Harada, M., ... & Motokawa, M. (2015). A new species of the genus Murina (Chiroptera: Vespertilionidae) from the Central Highlands of Vietnam with a review of the subfamily Murininae in Vietnam. Acta Chiropterologica, 17(2), 201-232.				India|Bhutan|Myanmar|China|Taiwan|Laos|Vietnam|Thailand|Malaysia|Indonesia|Philippines	Asia|Oceania	Palearctic|Indomalaya|Oceania	LC	0	0	0	Harpiocephalus_harpia	0	sciname match	Harpiocephalus_harpia	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Harpiocephalus_harpia	1005314	23	Hairy-winged Bat	Lesser Hairy-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Murininae	NA	Harpiocephalus	NA	harpia	Temminck	1	Vespertilio harpia	Temminck, C.J. 1840. Livraison 3. Pp. 141–272 in Temminck, C.J. 1835-1841. Monographies de Mammalogie. Tome second. C. C. van der Hoek, Leiden, 392 pp.	https://archive.org/details/monographiedema00temmgoog/page/140/mode/2up	RMNH.MAM.13470	syntypes	https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.13470.a | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.13470.b	"l'ÃŽle de Java [Indonesia] … les caverns de la partie sud du Volcan de Guédé [= caves of the southern part of Mount Gede]." Based on lectotype selection, restricted by A. M. Husson in 1955 to "S.E. side of Mount Gede, W. Java."			includes mordax as a synonym	Son, N. T., Csorba, G., Tu, V. T., Thong, V. D., Wu, Y., Harada, M., ... & Motokawa, M. (2015). A new species of the genus Murina (Chiroptera: Vespertilionidae) from the Central Highlands of Vietnam with a review of the subfamily Murininae in Vietnam. Acta Chiropterologica, 17(2), 201-232.				India|Bhutan|Myanmar|China|Taiwan|Laos|Vietnam|Thailand|Malaysia|Indonesia|Philippines	Asia|Oceania (Continent)	Palearctic|Indomalaya|Oceania (Biorealm)	LC	0	0	0	Harpiocephalus_harpia	0	sciname match	Harpiocephalus_harpia	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Harpiocephalus		harpia	Temminck	1840	1	Monogr. Mamm.	0.235417	Lesser Hairy-winged Bat	pearsonii Horsfield, 1851; rufus Gray, 1842; lasyurus Hodgson, 1847; madrassius Thomas, 1923; rufulus G. M. Allen, 1913.	Indonesia, Java, NE side of Mt. Gede	S and NE India, Myanmar, S China, Taiwan, Laos, Thailand, Cambodia, Vietnam, Sumatra, Java, Borneo, S Moluccas, and the Philippines.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/99711843/22045367/' target='_blank'>Least Concern</a>	Includes mordax; see Matveev (2005). Reviewed in part by Bates and Harrison (1997) and Hendrichsen et al. (2001b). The Taiwan record, if valid, has not been allocated to subspecies. See also Lin et al. (2006) and Son et al. (2015).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Harpiocephalus harpia; Harpiocephalus harpia; Harpiocephalus harpia; Harpiocephalus harpia; Harpiocephalus harpia; Harpiocephalus harpia; harpia; lasyurus; madrassius; rufulus; pearsonii; rufus; harpia; lasyurus; madrassius; rufulus; lasyurus; madrassius; rufulus; pearsonii; rufus; harpia; rufus; lasyurus; pearsonii; rufulus; madrassius; mordax; Murine a ailes velues; Haarfllgelfledermaus; Ratonero de alas pilosas; Lesser Hairy-winged Bat; Hairy-winged Bat; Lesser Hairy-winged Bat; Lesser Hairy-winged Bat; Lesser Hairy-winged Bat; H. harpia