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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L351	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Barticonycteris daviesi	Micronycteris daviesi	Barticonycteris daviesi	Micronycteris daviesi	Micronycteris daviesi	Glyphonycteris daviesi	Glyphonycteris daviesi	Glyphonycteris daviesi	Glyphonycteris daviesi	Glyphonycteris daviesi	Glyphonycteris daviesi	Glyphonycteris daviesi	Glyphonycteris daviesi	Glyphonycteris daviesi	Glyphonycteris daviesi		[MSW2] Subgenus Barticonycteris. Formerly included in genus Barticonycteris; see Koopman (1978ib:4).; [MSW3] Formerly included in the monotypic genus Barticonycteris; see Koopman (1978b) and Simmons (1996). Reviewed by Pine et al. (1996).; [HMW] Barticonycteris daviesi Hill, 1965, Guyana , Cuyuni-Mazaruni , “Forest reserve 24 miles [= 39 km ] from Bartica, along the Potaro road [ Cuyuni-Mazaruni ], British Guiana [= Guyana ].” Glyphonycteris daviesi was originally described as the only species of Barticonycteris and later was included in Micronycteris , subgenus Glyphonycteris . Monotypic.; [batnames2022] Formerly included in the monotypic genus Barticonycteris ; see Koopman (1978 b ) and Simmons (1996). Reviewed by Pine et al.(1996). Unvouchered in Honduras (Turcios-Casco et al., 2020).; [IUCN] Formerly included in the monotypic genus Barticonycteris .; [batnames2023] Formerly included in the monotypic genus Barticonycteris ; see Koopman (1978 b ) and Simmons (1996). Reviewed by Pine et al.(1996). Unvouchered in Honduras (Turcios-Casco et al., 2020).; [batnames2025_1.7] Formerly included in the monotypic genus Barticonycteris; see Koopman (1978b) and Simmons (1996). Reviewed by Pine et al.(1996). Unvouchered in Honduras (Turcios-Casco et al., 2020).														daviesi	Formerly included in the monotypic genus Barticonycteris .			daviesi 	daviesi 			daviesi (J. Edwards Hill, 1964)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Costa Rica, E Peru, Guyana	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Micronycteris daviesi	Guyana, Essequibo Prov., Potaro road, 24 mi. (39 km) from Bartica.	Hill	1964	Mammalia, 28:557.	Distribution: Same as for subgenus.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Davies' large-eared bat	Costa Rica, Panama, E Peru, Guyana, Surinam, Brazil; refs. 4.42, 43	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Hill	1964	Mammalia, 28:557.	Subgenus Barticonycteris. Formerly included in genus Barticonycteris; see Koopman (1978ib:4).	Costa Rica to Peru and French Guiana.	Guyana, Essequibo Prov., Potaro road, 24 mi. (39 km) from Bartica.		HILL	1964	Size relatively large (forearm length, 53-58 mm; condlyobasal length, 22-24 mm).	Distribution: Same as for subgenus.	No subspecies.		74	species	M. daviesi	HILL	1964	Barticonycteris	subgenus	Micronycteris daviesi				Size relatively large (forearm length, 53-58 mm; condlyobasal length, 22-24 mm).	No subspecies.		10. M. daviesi (HILL 1964).	10	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Phyllostominae		Glyphonycteris daviesi	Glyphonycteris		daviesi	Hill	y	1964		Mammalia	28		557		Graybeard Bat	Guyana, Essequibo Prov., Potaro road, 24 mi. (39 km) from Bartica.	Honduras south to Peru, the Guianas, Brazil, and Bolivia; Trinidad.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt) as Micronycteris daviesi.		Formerly included in the monotypic genus Barticonycteris; see Koopman (1978b) and Simmons (1996). Reviewed by Pine et al. (1996).	03A687BCFF87FF86168DFA93FE20F3B6	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	538	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFF87FF86168DFA93FE20F3B6.xml	Glyphonycteris daviesi	Phyllostomidae	Glyphonycteris	daviesi		1964	Glyphonyctere de Davies @fr | Davies-Gro Rohrblattnase @de | Glifonicterio de Davies @es | Davies's Graybeard Bat @en | Graybeard Bat @en	Barticonycteris daviesi Hill, 1965, Guyana , Cuyuni-Mazaruni , “Forest reserve 24 miles [= 39 km ] from Bartica, along the Potaro road [ Cuyuni-Mazaruni ], British Guiana [= Guyana ].” Glyphonycteris daviesi was originally described as the only species of Barticonycteris and later was included in Micronycteris , subgenus Glyphonycteris . Monotypic.	Honduras , Costa Rica , Panama , E Colombia , Venezuela , the Guianas, Brazil , Ecuador , E Peru , and N Bolivia ; also on Trinidad I; expected in Nicaragua .	Head-body 61-84 mm, tail 5~ 11 mm , ear 24-31 mm, hindfoot 15-20 mm, forearm 51-59 mm; weight 18-30 g. Davies’s Big-eared Bat is a large species of Glyphonycteris . Fur is long, loose, and woolly. Dorsal hair is dark grayish brown to dark brownish gray; hairs are uniformly colored for most of their length but have slightly paler tips and slightly paler bases. Underside of chin and throat has longer hairs, grayish brown for most oftheir length but tipped with grayish white. Fringe of short whitish hairs occurs under lowerlip. Snout is compressed, lips and upper chin are haired, and lowerlip has pair of smooth pads forming V on the chin. Ears are not connected by interauricular band. Fourth metacarpal is shortest, and fifth is longest; caudal membrane is developed;tail is short, reaching middle of the uropatagium; and calcar is shorter than foot. Dental formulais11/2,C1/1,P 2/3, M 3/3 ( x2 ) = 32. Upperincisors (I')are large (as long as canines) and project forward.	Mainly lowlands and primary wet tropical forest, secondarily little-disturbed forests, and infrequently cultivated areas (always close to primary forests) at elevations of 180-1030 m. Davies's Big-eared Bat has been captured in terra firma and seasonally inundated forest. Colombian specimens came from the Llanos region, characterized by savannas,riparian forests, and palm swamps.	Peculiar dentition of Davies's Big-eared Bat suggests that it might have an unusual diet. It appearsto be a gleaning insectivore.It eats cockroaches, dragonflies, and katydids taken from vegetation or the ground and carried back to a feeding roost before eating. Davies's Big-eared Bat is large enough to capture small vertebrates such as Norops lizards. A small frog was found in one stomach in Panama ; other stomach samples from Brazil included parts of a moth larva (family Lasiocampidae , probably a species of Euglyphis ) and remains of other insects. Perhaps its dentition is adapted to gnaw through some materials to secure some ofits prey or to create its roosts.	Lactating Daviess Big-eared Bats were reported in March ( Panama ) and August ( Brazil and Peru ); a pregnant female with a 33mm fetus was found in August ( Peru ). An adult male, two adult females (one lactating), a subadult male, and a young female were reported in August (Amazonas Department, Peru ). In Bahia , Brazil , two females reported in July were not pregnant or lactating.	Davies's Big-eared Bat is nocturnal. Recorded flight times are early in evening and 21:15 h to dawn. In Peru , it roosts in hollow trees, in one case 3 m aboveground.	Groups of 3-5 Davies's Big-eared Bats have been reported roosting together in Peru .	Classified as Least Concern on The IUCN Red List. In Bolivia , Davies's Big-eared Bat is assessed as endangered. It is rare, patchily distributed, and poorly known. Deforestation and resulting problems of habitat loss and fragmentation probably affect several populations in Central America, Chocé ecoregion in northwestern Ecuador , and northern Bolivia .	Aguirre & Teran (2007) | Gregorin & Rossi (2005) | Hill (1965) | LaVal & Rodriguez-Herrera (2002) | McCarthy & Ochoa (1991) | Medina-Fitoria & Saldana (2012) | Morales-Martinez & Suérez-Castro (2014) | Nogueira, Peracchi & Moratelli (2007) | Pine et al. (1996) | Solari et al. (1999) | Tirira (2017) | Tirira et al. (2016) | Tuttle (1970) | Williams & Genoways (2008)	https://zenodo.org/record/6458833/files/figure.png	112. Davies’s Big-eared Bat Glyphonycteris daviesi French: Glyphonyctere de Davies / German: Davies-GroRohrblattnase / Spanish: Glifonicterio de Davies Other common names: Davies's Graybeard Bat , Graybeard Bat Taxonomy. Barticonycteris daviesi Hill, 1965, Guyana , Cuyuni-Mazaruni , “Forest reserve 24 miles [= 39 km ] from Bartica, along the Potaro road [ Cuyuni-Mazaruni ], British Guiana [= Guyana ].” Glyphonycteris daviesi was originally described as the only species of Barticonycteris and later was included in Micronycteris , subgenus Glyphonycteris . Monotypic. Distribution. Honduras , Costa Rica , Panama , E Colombia , Venezuela , the Guianas, Brazil , Ecuador , E Peru , and N Bolivia ; also on Trinidad I; expected in Nicaragua . Descriptive notes. Head-body 61-84 mm, tail 5~ 11 mm , ear 24-31 mm, hindfoot 15-20 mm, forearm 51-59 mm; weight 18-30 g. Davies’s Big-eared Bat is a large species of Glyphonycteris . Fur is long, loose, and woolly. Dorsal hair is dark grayish brown to dark brownish gray; hairs are uniformly colored for most of their length but have slightly paler tips and slightly paler bases. Underside of chin and throat has longer hairs, grayish brown for most oftheir length but tipped with grayish white. Fringe of short whitish hairs occurs under lowerlip. Snout is compressed, lips and upper chin are haired, and lowerlip has pair of smooth pads forming V on the chin. Ears are not connected by interauricular band. Fourth metacarpal is shortest, and fifth is longest; caudal membrane is developed;tail is short, reaching middle of the uropatagium; and calcar is shorter than foot. Dental formulais11/2,C1/1,P 2/3, M 3/3 ( x2 ) = 32. Upperincisors (I')are large (as long as canines) and project forward. Habitat. Mainly lowlands and primary wet tropical forest, secondarily little-disturbed forests, and infrequently cultivated areas (always close to primary forests) at elevations of 180-1030 m. Davies's Big-eared Bat has been captured in terra firma and seasonally inundated forest. Colombian specimens came from the Llanos region, characterized by savannas,riparian forests, and palm swamps. Food and Feeding. Peculiar dentition of Davies's Big-eared Bat suggests that it might have an unusual diet. It appearsto be a gleaning insectivore.It eats cockroaches, dragonflies, and katydids taken from vegetation or the ground and carried back to a feeding roost before eating. Davies's Big-eared Bat is large enough to capture small vertebrates such as Norops lizards. A small frog was found in one stomach in Panama ; other stomach samples from Brazil included parts of a moth larva (family Lasiocampidae , probably a species of Euglyphis ) and remains of other insects. Perhaps its dentition is adapted to gnaw through some materials to secure some ofits prey or to create its roosts. Breeding. Lactating Daviess Big-eared Bats were reported in March ( Panama ) and August ( Brazil and Peru ); a pregnant female with a 33mm fetus was found in August ( Peru ). An adult male, two adult females (one lactating), a subadult male, and a young female were reported in August (Amazonas Department, Peru ). In Bahia , Brazil , two females reported in July were not pregnant or lactating. Activity patterns. Davies's Big-eared Bat is nocturnal. Recorded flight times are early in evening and 21:15 h to dawn. In Peru , it roosts in hollow trees, in one case 3 m aboveground. Movements, Home range and Social organization. Groups of 3-5 Davies's Big-eared Bats have been reported roosting together in Peru . Status and Conservation. Classified as Least Concern on The IUCN Red List. In Bolivia , Davies's Big-eared Bat is assessed as endangered. It is rare, patchily distributed, and poorly known. Deforestation and resulting problems of habitat loss and fragmentation probably affect several populations in Central America, Chocé ecoregion in northwestern Ecuador , and northern Bolivia . Bibliography. Aguirre & Teran (2007), Gregorin & Rossi (2005), Hill (1965), LaVal & Rodriguez-Herrera (2002), McCarthy & Ochoa (1991), Medina-Fitoria & Saldana (2012), Morales-Martinez & Suérez-Castro (2014), Nogueira, Peracchi & Moratelli (2007), Pine et al. (1996), Solari et al. (1999), Tirira (2017), Tirira et al. (2016), Tuttle (1970), Williams & Genoways (2008).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Glyphonycteris daviesi	Glyphonycteris		daviesi	Hill	1965	1	Mammalia	37:17:00	Graybeard Bat	None.	Guyana, Essequibo Prov., Potaro road, 24 mi. (39 km) from Bartica.	Honduras south to Peru, the Guianas, Brazil, and Bolivia; Trinidad.	Not listed.	Least Concern	Formerly included in the monotypic genus Barticonycteris ; see Koopman (1978 b ) and Simmons (1996). Reviewed by Pine et al.(1996). Unvouchered in Honduras (Turcios-Casco et al., 2020).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Glyphonycteris daviesi	23	Davies's Big-eared Bat	Davies's Graybeard Bat|Graybeard Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	GLYPHONYCTERINAE	NA	Glyphonycteris	NA	daviesi	J. Edwards Hill	1964	1	Barticonycteris_daviesi	Hill, J. E. (1964). Notes on bats from British Guiana, with the description of a new genus and species of Phyllostomidae. Mammalia, 28, 553.	https://www.degruyter.com/view/journals/mamm/28/4/article-p553.xml	BM 1964.767		Guyana, Cuyuni-Mazaruni, "Forest reserve 24 miles [= 39 km] from Bartica, along the Potaro road [Cuyuni-Mazaruni], British Guiana [= Guyana]."			daviesi (J. Edwards Hill, 1964)	NA	NA	Honduras|Costa Rica|Panama|Colombia|Venezuela|Trinidad & Tobago|Guyana|Suriname|French Guiana|Brazil|Ecuador|Peru|Bolivia	North America|South America	Neotropic	LC	0	0	0	Glyphonycteris_daviesi	0	sciname match	Glyphonycteris_daviesi	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	13377	Glyphonycteris daviesi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Glyphonycteris	daviesi	(Hill, 1964)	Formerly included in the monotypic genus Barticonycteris .	20000000	Glyphonycteris daviesi	Least Concern		2018	2018-02-20 00:00:00 UTC	3.1	English	This species is listed as Least Concern in view of its relatively wide distribution and because it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category. However, this species is rare and its habitat type is threatened regionally. This species seems to be locally threatened (as in Bolivia).	The few available data indicates this is a forest interior species. In general, this species is poorly known; probably restricted to mature, evergreen forest. Roosts in hollow trees (Williams and Genoways 2008); occasionally caught in harp traps or mist nets set across forest trails. These bats feed on large insects and occasionally fruit; they forage by gleaning large insects such as cockroaches, dragonflies, and katydids from the vegetation or ground, and they carry their prey back to a feeding roost before eating (Emmons and Feer 1997). A small spotted frog was found in the stomach of one individual from Panama (Reid 2009). Has been found in secondary forested areas near primary forest (J. Ochoa, pers. comm.), but seems to prefer primary forest. Roost in hollow trees; there is a record of males, females and juveniles roosting together in a lowland forest in northern Peru (Solari et al . 1999). Some records of ecoparasites (Solari pers. comm.).	In Bolivia this species is considered vulnerable as it occurs in fragile ecosystems (montane forests and flood plains; Aguirre 1999). Loss of primary forest, and reduction and fragmentation of habitat are localized threats, but there are no major threats throughout its range at present.	This species is relatively rare, which could be either an artifact of sampling methodology or local fluctuations in abundance (Pine et al . 1996). Apparently very rare and patchily distributed (Reid 2009). There are only two instances of roosts for this species (Williams and Genoways 2008), suggesting that individuals have very specific requirements at a local scale.	Unknown	Glyphonycteris daviesi occurs throughout Honduras south to Peru, the Guianas, Brazil, and Bolivia, as well as Trinidad (Simmons 2005). There are recent records from Colombia (Morales-M. and SuÃ¡rez-Castro 2014) and eastern Brazil (Gregorin and Rossi 2005). Range is confined to eastern flanks of the Andes. Known from 0-300 m asl (J. Ochoa, pers. comm.), and in Ecuador up to 500 m (Pine et al. 1996).		Terrestrial	The species is found in protected areas. A priority would be the conservation of habitat, mostly of primary lowland forests. The species occurs in a number of protected areas throughout its geographic range.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Glyphonycteris		daviesi	Hill	1965	1	Mammalia	37:17:00	Graybeard Bat	None.	Guyana, Essequibo Prov., Potaro road, 24 mi. (39 km) from Bartica.	Honduras south to Peru, the Guianas, Brazil, and Bolivia; Trinidad.	Not listed.	Least Concern	Formerly included in the monotypic genus Barticonycteris ; see Koopman (1978 b ) and Simmons (1996). Reviewed by Pine et al.(1996). Unvouchered in Honduras (Turcios-Casco et al., 2020).	Glyphonycteris daviesi	1004920	23	Davies's Big-eared Bat	Davies's Graybeard Bat|Graybeard Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	GLYPHONYCTERINAE	NA	Glyphonycteris	NA	daviesi	J. Edwards Hill	1964	1	Barticonycteris_daviesi	Hill, J. E. (1964). Notes on bats from British Guiana, with the description of a new genus and species of Phyllostomidae. Mammalia, 28, 553.	https://www.degruyter.com/view/journals/mamm/28/4/article-p553.xml	BM 1964.767		Guyana, Cuyuni-Mazaruni, "Forest reserve 24 miles [= 39 km] from Bartica, along the Potaro road [Cuyuni-Mazaruni], British Guiana [= Guyana]."			daviesi (J. Edwards Hill, 1964)	NA	NA				Honduras|Costa Rica|Panama|Colombia|Venezuela|Trinidad & Tobago|Guyana|Suriname|French Guiana|Brazil|Ecuador|Peru|Bolivia	North America|South America	Neotropic	LC	0	0	0	Glyphonycteris_daviesi	0	sciname match	Glyphonycteris_daviesi	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Glyphonycteris_daviesi	1004920	23	Davies's Big-eared Bat	Davies's Graybeard Bat|Graybeard Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Glyphonycterinae	NA	Glyphonycteris	NA	daviesi	J. Edwards Hill	1	Barticonycteris daviesi	Hill, J.E. 1964. Notes on bats from British Guiana, with the description of a new genus and species of Phyllostomidae. Mammalia 28(4):553-572.	https://doi.org/10.1515/mamm.1964.28.4.553	BMNH:Mamm:1964.767	holotype	https://data.nhm.ac.uk/object/3a29d977-6a78-4a7c-bb9c-73233b797e86	Guyana, Cuyuni-Mazaruni, "Forest reserve 24 miles [= 39 km] from Bartica, along the Potaro road [Cuyuni-Mazaruni], British Guiana [= Guyana]."			NA	NA				Honduras|Costa Rica|Panama|Colombia|Venezuela|Trinidad and Tobago|Guyana|Suriname|French Guiana|Brazil|Ecuador|Peru|Bolivia	North America|South America	Neotropic	LC	0	0	0	Glyphonycteris_daviesi	0	sciname match	Glyphonycteris_daviesi	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Glyphonycteris		daviesi	Hill	1965	1	Mammalia	37:17:00	Graybeard Bat	None.	Guyana, Essequibo Prov., Potaro road, 24 mi. (39 km) from Bartica.	Honduras south to Peru, the Guianas, Brazil, and Bolivia; Trinidad.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/13377/22124873/' target='_blank'>Least Concern</a>	Formerly included in the monotypic genus Barticonycteris; see Koopman (1978b) and Simmons (1996). Reviewed by Pine et al.(1996). Unvouchered in Honduras (Turcios-Casco et al., 2020).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Glyphonycteris daviesi; Glyphonycteris daviesi; Glyphonycteris daviesi; Glyphonycteris daviesi; Glyphonycteris daviesi; Glyphonycteris daviesi; daviesi; Glyphonyctere de Davies; Davies-Gro Rohrblattnase; Glifonicterio de Davies; Davies's Graybeard Bat; Graybeard Bat; Davies's Big-eared Bat; Davies's Graybeard Bat; Graybeard Bat; Graybeard Bat; Graybeard Bat; G. daviesi
