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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L36	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Pipistrellus angulatus [synonym of]	N/A	Pipistrellus tenuis [synonym of]	Pipistrellus tenuis [synonym of]	Pipistrellus tenuis papuanus	Pipistrellus papuanus	Pipistrellus papuanus	Pipistrellus papuanus	Pipistrellus papuanus	Pipistrellus papuanus	Pipistrellus papuanus	Pipistrellus papuanus	Pipistrellus papuanus	Pipistrellus papuanus	Alionoctula papuana		[MSW3] Subgenus Pipistrellus. Included in tenuis by many authors, but see Kitchener et al. (1986); also see Flannery (1995a, b) and Bonaccorso (1998).; [HMW] Vesperugo papuanus Peters & Doria, 1881 , “Salawati [Island],” West Papua , Indonesia . Pipistrellus papuanus has been included under P. tenuis but is considered a distinct species based on morphology. Monotypic.; [batnames2022] Subgenus Pipistrellus . Included in tenuis by many authors, but see Kitchener et al. (1986); also see Flannery (1995 a , b ) and Bonaccorso (1998).; [IUCN] Koopman in Wilson and Reeder (1993) treated this as a synonym of P. tenuis papuanus whereas others consider it a valid species.; [batnames2023] Subgenus Pipistrellus . Included in tenuis by many authors, but see Kitchener et al. (1986); also see Flannery (1995 a , b ) and Bonaccorso (1998).; [batnames2025_1.7] Subgenus Pipistrellus. Included in tenuis by many authors, but see Kitchener et al. (1986); also see Flannery (1995a, b) and Bonaccorso (1998).; [MDD2025_2.2] moved from Pipistrellus to Alionoctula									orientalis			papuanus 	papuanus - orientalis	papuanus, orientalis	Koopman in Wilson and Reeder (1993) treated this as a synonym of P. tenuis papuanus whereas others consider it a valid species.	papuanus 	papuanus - orientalis	papuanus, orientalis	papuanus, orientalis	papuanus 	papuanus - orientalis	papuana (W. C. H. Peters & Doria, 1881)|orientalis (A. B. Meyer, 1899)						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Pipistrellini	Pipistrellus papuanus	Pipistrellus	Pipistrellus	papuanus	Peters and Doria		1881		Ann. Mus. Stor. Nat. Genova	16		696		Lesser Papuan Pipistrelle	Indonesia, Prov. of Papua, Salawati Isl.	Seram, Aru Isls, Baik-Supiori, New Guinea, New Ireland (Bismarck Arch.), adjacent small islands.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).	orientalis Meyer, 1899.	Subgenus Pipistrellus. Included in tenuis by many authors, but see Kitchener et al. (1986); also see Flannery (1995a, b) and Bonaccorso (1998).	4C3D87E8FFEC6A53FF9092061C26B04E	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	782	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFE36A5CFF449CE81442BD54.xml	Pipustrellus papuanus	Vespertilionidae	Pipustrellus	papuanus		1881	Pipistrelle papoue @fr | Kleine Papua-Zwergfledermaus @de | Pipistrelapequena de Papua @es | Papuan Pipistrelle @en	Vesperugo papuanus Peters & Doria, 1881 , “Salawati [Island],” West Papua , Indonesia . Pipistrellus papuanus has been included under P. tenuis but is considered a distinct species based on morphology. Monotypic.	E New Guinea , Seram, Ambon , Batanta, Salawati, Biak-Supiori, Dolak, New Ireland, and New Britain Is as well as Kai, Aru, Fergusson, and Samarai Is.	Head-body 33.-9-50 mm, tail 22-35 mm, ear 7-5-1383 mm, hindfoot 4-5-9 mm, forearm 27-32-2 mm; weight 3-9-5-8 g. Dorsal pelage of the Lesser Papuan Pipistrelle is dark brown; ventral pelage is pale brown, with some whitish hairs intermixed. Skin on face, ears, arms, and wing membranes are medium brown. Uropatagium is lightly furred, with cinnamon brown hair along venation; forearm is lightly furred under humerus. Ears are narrow and subtriangular, with rounded tips; basal lobe of tragus is not as developed as in the Forest Pipistrelle (P. adamst), but the rest of tragus is like that of latter species. Uropatagium stretches nearly to end of tail (only extreme tip is free), and postcalcarial lobe is elongated. Glans penis is similar to that of the Forest Pipistrelle but with shorter and broader distal projecting lobe and very reduced dorsal distal fold. Baculum is short, with broad base, relatively broad uncurved shaft, and bifurcated tip (for last 10-20% of baculum). Skull is small, fragile, flat, and generally similar to the Northern Pipistrelle ( P. westralis ) in most aspects; I” is usually less than the height of posterior cusp of I* and does not extrude from tooth row; P* is less than one-half the height of P* and slightly extrudes from tooth row; and lower molars are nyctalodont.	Generally associated with towns, villages, and gardens (particularly in lowland areas) and undisturbed lowland forests from sea level up to elevations of ¢. 1300 m .	The Lesser Papuan Pipistrelle is insectivorous.	Pregnant and lactating Lesser Papuan Pipistrelles have been captured in two major peaks in Papua New Guinea : March-April and August-September. Maternity colonies have been observed in August. In Wau, eastern Papua New Guinea , they appear to breed continuously year-round, with pregnant and lactating females and immature individuals being captured year-round. Wau does not vary in temperature and precipitation throughout the year, as do some other regions of New Guinea , which might allow continuous breeding.	Lesser Papuan Pipistrelles emerge shortly before dusk to forage in urban areas and plantations. They roost in buildings and hollows in trees. Call shape is very steep FM.	Lesser Papuan Pipistrelles commonly roost in colonies of several hundred individuals. Colonies of 40-50 individuals have frequently been reported in cavities in coconut trees. It seems to form maternity colonies with hundreds of individuals.	Classified as Least Concern on The IUCN Red List. The Lesser Papuan Pipistrelle has a wide distribution,is tolerant to habitat modification, has a presumably large population, and does not seem to face any major threats. It appears to tolerate humans and might have extended its distribution as humans spread through the region.	Aplin & Opiang (2009) | Bonaccorso (1998) | Bonaccorso & Reardon ( 2008g) | Flannery (1995a, 1995b) | Grinnell & Hagiwara (1972) | Hill & Harrison (1987) | Kitchener et al. (1986) | Woolley & Allison (1996)	https://zenodo.org/record/6397862/files/figure.png	44. Lesser Papuan Pipistrelle Pipustrellus papuanus French: Pipistrelle papoue / German: Kleine Papua-Zwergfledermaus / Spanish: Pipistrela pequena de Papua Other common names: Papuan Pipistrelle Taxonomy. Vesperugo papuanus Peters & Doria, 1881 , “Salawati [Island],” West Papua , Indonesia . Pipistrellus papuanus has been included under P. tenuis but is considered a distinct species based on morphology. Monotypic. Distribution. E New Guinea , Seram, Ambon , Batanta, Salawati, Biak-Supiori, Dolak, New Ireland, and New Britain Is as well as Kai, Aru, Fergusson, and Samarai Is. Descriptive notes. Head-body 33.-9-50 mm, tail 22-35 mm, ear 7-5-1383 mm, hindfoot 4-5-9 mm, forearm 27-32-2 mm; weight 3-9-5-8 g. Dorsal pelage of the Lesser Papuan Pipistrelle is dark brown; ventral pelage is pale brown, with some whitish hairs intermixed. Skin on face, ears, arms, and wing membranes are medium brown. Uropatagium is lightly furred, with cinnamon brown hair along venation; forearm is lightly furred under humerus. Ears are narrow and subtriangular, with rounded tips; basal lobe of tragus is not as developed as in the Forest Pipistrelle (P. adamst), but the rest of tragus is like that of latter species. Uropatagium stretches nearly to end of tail (only extreme tip is free), and postcalcarial lobe is elongated. Glans penis is similar to that of the Forest Pipistrelle but with shorter and broader distal projecting lobe and very reduced dorsal distal fold. Baculum is short, with broad base, relatively broad uncurved shaft, and bifurcated tip (for last 10-20% of baculum). Skull is small, fragile, flat, and generally similar to the Northern Pipistrelle ( P. westralis ) in most aspects; I” is usually less than the height of posterior cusp of I* and does not extrude from tooth row; P* is less than one-half the height of P* and slightly extrudes from tooth row; and lower molars are nyctalodont. Habitat. Generally associated with towns, villages, and gardens (particularly in lowland areas) and undisturbed lowland forests from sea level up to elevations of ¢. 1300 m . Food and Feeding. The Lesser Papuan Pipistrelle is insectivorous. Breeding. Pregnant and lactating Lesser Papuan Pipistrelles have been captured in two major peaks in Papua New Guinea : March-April and August-September. Maternity colonies have been observed in August. In Wau, eastern Papua New Guinea , they appear to breed continuously year-round, with pregnant and lactating females and immature individuals being captured year-round. Wau does not vary in temperature and precipitation throughout the year, as do some other regions of New Guinea , which might allow continuous breeding. Activity patterns. Lesser Papuan Pipistrelles emerge shortly before dusk to forage in urban areas and plantations. They roost in buildings and hollows in trees. Call shape is very steep FM. Movements, Home range and Social organization. Lesser Papuan Pipistrelles commonly roost in colonies of several hundred individuals. Colonies of 40-50 individuals have frequently been reported in cavities in coconut trees. It seems to form maternity colonies with hundreds of individuals. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Lesser Papuan Pipistrelle has a wide distribution,is tolerant to habitat modification, has a presumably large population, and does not seem to face any major threats. It appears to tolerate humans and might have extended its distribution as humans spread through the region. Bibliography. Aplin & Opiang (2009), Bonaccorso (1998), Bonaccorso & Reardon ( 2008g ), Flannery (1995a, 1995b), Grinnell & Hagiwara (1972), Hill & Harrison (1987), Kitchener et al. (1986), Woolley & Allison (1996).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Pipistrellus papuanus	Pipistrellus	Pipistrellus	papuanus	Peters and Doria	1881	0	Ann. Mus. Stor. Nat. Genova	1.15	Lesser Papuan Pipistrelle	 orientalis Meyer, 1899.	Indonesia, Prov. of Papua, Salawati Isl.	Seram, Aru Isls, Baik-Supiori, New Guinea, New Ireland (Bismarck Arch.), adjacent small islands.	Not listed.	Least Concern	Subgenus Pipistrellus . Included in tenuis by many authors, but see Kitchener et al. (1986); also see Flannery (1995 a , b ) and Bonaccorso (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Pipistrellus papuanus	23	Lesser Papuan Pipistrelle	Papuan Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PIPISTRELLINI	Pipistrellus	NA	papuanus	W. Peters & Doria	1881	1	Vesperugo_papuanus	Peters, W. C. H. & Doria, A. (1881). Enumerazione dei Mammiferi raccolti da O. Beccari, L. M. d'Albertis, ed A. A. Bruijn, nella Nuova Guinea propriamente detta. Annali del Museo civico di storia naturale di Genova, 16, 696.	https://www.biodiversitylibrary.org/item/94436#page/708/mode/1up	MSNG 11660		"Salawati [Island]," West Papua, Indonesia.			papuanus (W. Peters & Doria, 1881)|orientalis (A. B. Meyer, 1899)	NA	NA	Indonesia|Papua New Guinea	Oceania	Australasia/Oceania	LC	0	0	0	Pipistrellus_papuanus	0	sciname match	Pipistrellus_papuanus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	17355	Pipistrellus papuanus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Pipistrellus	papuanus	Peters &; Doria, 1881	Koopman in Wilson and Reeder (1993) treated this as a synonym of P. tenuis papuanus whereas others consider it a valid species.	20000000	Pipistrellus papuanus	Least Concern		2020	2019-07-30 00:00:00 UTC	3.1	English	This species is listed as Least Concern in view of its wide distribution, tolerance of habitat modification, large populations, and because its population trend is believed to be stable or increasing. The Papuan Pipistrelle is commensal with humans and may have increased in distribution with the spread of people (Bonaccorso 1998).	This species commonly roosts in buildings, tree hollows, and in small cavities in coconut trees (in groups of up to 50 animals) (Flannery 1995b, Bonaccorso 1998). This species has been recorded from undisturbed lowland forest (D. Wright pers. comm.), but it is more commonly found as a human commensal species that seems to have benefitted from the modification of natural habitat as it is often found in towns, villages, coconut plantations and gardens, particularly in low-lying areas.  It is an aerial insectivore that emerges shortly before dusk to forage in urban areas and plantations. The Papuan Pipistrelle may produce at least two litters of a single young per year in Papua New Guinea, with birth peaks in March/April and August/September (Bonaccorso 1998), though at Wau there was evidence of continuous reproduction and polyoestry (Woolley and Allison 1996).	There appear to be no major threats to this species. It may suffer some localised persecution within towns, villages, and other settlements.	It is a very common species that can be found in colonies of hundreds of individuals. Due to its association with humans in towns and villages populations of this species are likely increasing (Bonaccorso 1998).	Stable	This widespread species is distributed from the islands of Seram, Ambon, Batanta, Dolak, Biak-Supiori, Salawati, the Aru Islands, and the Kai Islands (all Indonesia) in the west of its range, through parts of the island of New Guinea (Indonesia and Papua New Guinea). It is also present on Fergusson Island and the Samarai Islands in Milne Bay Province (Papua New Guinea), on the islands of New Britain and New Ireland in the Bismarck Archipelago (Papua New Guinea), ranging as far east as the island of Bougainville (Papua New Guinea) (Flannery 1995a,b; Bonaccorso 1998). It occurs from sea level to 1,300 m asl.	Not hunted or traded.	Terrestrial	It is presumed to occur in some protected areas.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Pipistrellus	Pipistrellus	papuanus	Peters and Doria	1881	0	Ann. Mus. Stor. Nat. Genova	1.15	Lesser Papuan Pipistrelle	 orientalis Meyer, 1899.	Indonesia, Prov. of Papua, Salawati Isl.	Seram, Aru Isls, Baik-Supiori, New Guinea, New Ireland (Bismarck Arch.), adjacent small islands.	Not listed.	Least Concern	Subgenus Pipistrellus . Included in tenuis by many authors, but see Kitchener et al. (1986); also see Flannery (1995 a , b ) and Bonaccorso (1998).	Pipistrellus papuanus	1005629	23	Lesser Papuan Pipistrelle	Papuan Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PIPISTRELLINI	Pipistrellus	NA	papuanus	W. Peters & Doria	1881	1	Vesperugo_papuanus	Peters, W. C. H. & Doria, A. (1881). Enumerazione dei Mammiferi raccolti da O. Beccari, L. M. d'Albertis, ed A. A. Bruijn, nella Nuova Guinea propriamente detta. Annali del Museo civico di storia naturale di Genova, 16, 696.	https://www.biodiversitylibrary.org/item/94436#page/708/mode/1up	MSNG 11660		"Salawati [Island]," West Papua, Indonesia.			papuanus (W. Peters & Doria, 1881)|orientalis (A. B. Meyer, 1899)	NA	NA				Indonesia|Papua New Guinea	Oceania	Australasia/Oceania	LC	0	0	0	Pipistrellus_papuanus	0	sciname match	Pipistrellus_papuanus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Alionoctula_papuana	1005629	23	Lesser Papuan Pipistrelle	Papuan Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Pipistrellini	Alionoctula	NA	papuana	W. C. H. Peters & Doria	1	Vesperugo papuanus	Peters, W.C.H. and Doria, G. 1881-03-31. Enumerazione dei Mammiferi raccolti da O. Beccari, L. M. d'Albertis, ed A. A. Bruijn, nella Nuova Guinea propriamente detta. Annali del Museo civico di storia naturale di Genova 16:665-707.	https://www.biodiversitylibrary.org/page/29845133	MSNG 11660	holotype		"Salawati [Island]," West Papua, Indonesia.			moved from Pipistrellus to Alionoctula	Zhukova, S. S., Yuzefovich, A. P., Lebedev, V. S., & Kruskop, S. V. (2025). Reassessment of the Taxonomic Borders Within Pipistrellus (Chiroptera, Vespertilionidae, Pipistrellini). Diversity, 17(5), 317.				Indonesia|Papua New Guinea	Oceania (Continent)	Australasia	LC	0	0	0	Pipistrellus_papuanus	0	sciname match	Pipistrellus_papuanus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Pipistrellus	Pipistrellus	papuanus	Peters & Doria	1881	1	Ann. Mus. Stor. Nat. Genova	1.15	Lesser Papuan Pipistrelle	orientalis Meyer, 1899.	Indonesia, Prov. of Papua, Salawati Isl.	Seram, Aru Isls, Baik-Supiori, New Guinea, New Ireland (Bismarck Arch.), adjacent small islands.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/17355/22127056/' target='_blank'>Least Concern</a>	Subgenus Pipistrellus. Included in tenuis by many authors, but see Kitchener et al. (1986); also see Flannery (1995a, b) and Bonaccorso (1998).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Pipistrellus papuanus; Pipistrellus papuanus; Pipistrellus papuanus; Pipistrellus papuanus; Pipistrellus papuanus; Pipistrellus papuanus; orientalis; orientalis; papuanus; orientalis; Pipistrelle papoue; Kleine Papua-Zwergfledermaus; Pipistrelapequena de Papua; Papuan Pipistrelle; Lesser Papuan Pipistrelle; Papuan Pipistrelle; Lesser Papuan Pipistrelle; Lesser Papuan Pipistrelle; P. papuanus