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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L338	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Glischropus tylopus	Glischropus tylopus	Glischropus tylopus	Glischropus tylopus	Glischropus tylopus	Glischropus tylopus	Glischropus tylopus	Glischropus tylopus	Glischropus tylopus	Glischropus tylopus	Glischropus tylopus	Glischropus tylopus	Glischropus tylopus	Glischropus tylopus	Glischropus tylopus		[MSW3] See Corbet and Hill (1992) and Flannery (1995b).; [HMW] Vesperugo tylopus Dobson, 1875 , “ North Borneo ,” Malaysia . This speciesis sister to G. aquilus , although it is morphologically most similar to G. javanus . The name batjanus from Bacan Island in the Moluccas has been regarded as a distinct species or a subspecies of G. tylopus , but is here regarded as synonym of G. tylopus . Monotypic.; [batnames2022] See Corbet and Hill (1992) and Flannery (1995 b ).; [IUCN] The species was formerly considered to North of the Kra Isthmus in Thailand, but the form there is now recognised as a distinct species, Glischropus bucephalus (Csorba 2011).; [batnames2023] See Corbet and Hill (1992) and Flannery (1995 b ).; [batnames2025_1.7] See Corbet and Hill (1992) and Flannery (1995b).						batjanus.	batjanus, tylopus	tylopus, batjanus			batjanus	tylopus, batjanus		tylopus, batjanus	The species was formerly considered to North of the Kra Isthmus in Thailand, but the form there is now recognised as a distinct species, Glischropus bucephalus (Csorba 2011).	tylopus, batjanus		tylopus, batjanus 	tylopus, batjanus 	batjanus, tylopus		tylopus (Dobson, 1875)|batjanus (Matschie, 1901)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Thick-thumbed pipistrelle	Burma – Sumatra, Borneo, Palawan	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Glischropus tylopus	Malaysia, Sabah.	Dobson	1875	Proc. Zool. Soc. Lond., 1875:473.	Distribution: Ranging from Burma and Thailand to Sumatra, Borneo, and Palawan, also the northern Moluccas.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Thick-thumbed pipistrelle	Burma – Sumatra, Borneo, Palawan, N Moluccas	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Dobson	1875	Proc. Zool. Soc. Lond., 1875:473.		Burma, Thailand, W Malaysia, Borneo, SW Philippines, Sumatra and N Molucca Isis.	Malaysia, Sabah.		DOBSON	1875	Size relatively small (forearm length, 28-30 mm). Braincase not flattened.	Distribution: Ranging from Burma and Thailand to Sumatra, Borneo, and Palawan, also the northern Moluccas.	Two subspecies:	G. t. batjanus (northern Moluccas), G. t. tylopus (remainder of range).	119	species	G. tylopus	DOBSON	1875	Glischropus	genus	Glischropus tylopus				Size relatively small (forearm length, 28-30 mm). Braincase not flattened.	Two subspecies:		1. G. tylopus (DOBSON 1875).	1	_G. t. batjanus_ (Matschie, 1901); _G. t. tylopus_ (Dobson, 1875)			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Pipistrellini	Glischropus tylopus	Glischropus		tylopus	Dobson		1875		Proc. Zool. Soc. Lond.	1875		473		Common Thick-thumbed Bat	Malaysia, N Boreneo, Sabah.	Burma, Thailand, W Malaysia, Borneo, Palawan (Philippines), Sumatra and N Molucca Isls.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	batjanus Marschie, 1901.	See Corbet and Hill (1992) and Flannery (1995b).	4C3D87E8FFFE6A41FA889524188EB1DE	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	763	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFFE6A41FA889524188EB1DE.xml	Glischropus tylopus	Vespertilionidae	Glischropus	tylopus	Dobson	1875	Pipistrelle de Dobson @fr | Gewohnliche Dickdaumenfledermaus @de | Pipistrela de Dobson @es	Vesperugo tylopus Dobson, 1875 , “ North Borneo ,” Malaysia . This speciesis sister to G. aquilus , although it is morphologically most similar to G. javanus . The name batjanus from Bacan Island in the Moluccas has been regarded as a distinct species or a subspecies of G. tylopus , but is here regarded as synonym of G. tylopus . Monotypic.	Malay Peninsula, N Sumatra, Borneo, Palawan I ( Philippines ), and Bacan I ( Moluccas ).	Head-body 39 mm (holotype), tail 32-39 mm, ear 8-11 mm, hindfoot 6-6 mm (holotype), forearm 27-30-9 mm; weight 3-5-5 g. All species of Glischropus have thick thumb and foot padsthat rely on friction for holding onto insides of bamboo stems, rocks, and large leaves. Pelage is reddish yellow without any banding both dorsally and ventrally. Ears are moderately sized for the genus, being rounded and dark brown. Face is also dark brown. Tragus is relatively narrow with a broadly rounded tip and is angled slightly forward. Skull is flatter than in the Indochinese Thick-thumbed Bat ( Glischropus bucephalus ) and the Dark Thick-thumbed Bat ( G. aquilus ); the species can be distinguished from the Javan Thick-thumbed Bat ( G. javanus ) byits shorter forearm (generally less than 30 mm ), and from the Dark Thick-thumbed Bat byits lighter and more reddish yellow pelage. I’ is much shorter than secondary cusp of I? and displaced outward; main cusp of I barely exceeds smaller cusplet of I; P! is small but in tooth row: lower molars are nyctalodont. Chromosomal complement has 2n = 30 (males) or 2n = 31 (females) and FN = 50; the species shows an X-autosome translocation in which a small acrocentric chromosome has been transferred to the X-chromosome.	Throughoutits range, the Common Thick-thumbed Bat is generally found in forested regions with abundant bamboo, primarily at lower elevations, below 1000 m .	Common Thick-thumbed Bats are insectivorous.	In the Malay Peninsula, a female gave birth to a single young in mid-April.	Roosts have been found most commonly in dead or damaged bamboo stands as well as in rock crevices and banana leaves. Start frequency has been recorded at 95 kHz followed by a steep FM sweep ending in a QCF sweep at 49-45 kHz. Call durations were recorded at 2-7 milliseconds.	Common Thick-thumbed Bats have been found in a group of twelve in bamboos in Brunei . They may share roosts with species of Tylonycteris , although this is unconfirmed.	Classified as Least Concern on The IUCN Red List. The Common Thick-thumbed Bat is relatively widespread but difficult to find throughoutits range. It can be found in regenerated forests after disturbance. Relatively little information is available on the species’ ecology, and further research is needed.	Csorba (2011) | Csorba et al. (2015) | Heaney et al. (1998) | Heller (1989) | Heller & Volleth (1989) | Kofron (1994) | Mohd-Hanif et al. (2015) | Rosell-Ambal et al. (2008) | Thewissen & Etnier (1995) | Volleth & Yong (1987) | Volleth et al. (2001)	https://zenodo.org/record/6397774/files/figure.png	7. Common Thick-thumbed Bat Glischropus tylopus French: Pipistrelle de Dobson / German: Gewohnliche Dickdaumenfledermaus / Spanish: Pipistrela de Dobson Taxonomy. Vesperugo tylopus Dobson, 1875 , “ North Borneo ,” Malaysia . This speciesis sister to G. aquilus , although it is morphologically most similar to G. javanus . The name batjanus from Bacan Island in the Moluccas has been regarded as a distinct species or a subspecies of G. tylopus , but is here regarded as synonym of G. tylopus . Monotypic. Distribution. Malay Peninsula, N Sumatra, Borneo, Palawan I ( Philippines ), and Bacan I ( Moluccas ). Descriptive notes. Head-body 39 mm (holotype), tail 32-39 mm, ear 8-11 mm, hindfoot 6-6 mm (holotype), forearm 27-30-9 mm; weight 3-5-5 g. All species of Glischropus have thick thumb and foot padsthat rely on friction for holding onto insides of bamboo stems, rocks, and large leaves. Pelage is reddish yellow without any banding both dorsally and ventrally. Ears are moderately sized for the genus, being rounded and dark brown. Face is also dark brown. Tragus is relatively narrow with a broadly rounded tip and is angled slightly forward. Skull is flatter than in the Indochinese Thick-thumbed Bat ( Glischropus bucephalus ) and the Dark Thick-thumbed Bat ( G. aquilus ); the species can be distinguished from the Javan Thick-thumbed Bat ( G. javanus ) byits shorter forearm (generally less than 30 mm ), and from the Dark Thick-thumbed Bat byits lighter and more reddish yellow pelage. I’ is much shorter than secondary cusp of I? and displaced outward; main cusp of I barely exceeds smaller cusplet of I; P! is small but in tooth row: lower molars are nyctalodont. Chromosomal complement has 2n = 30 (males) or 2n = 31 (females) and FN = 50; the species shows an X-autosome translocation in which a small acrocentric chromosome has been transferred to the X-chromosome. Habitat. Throughoutits range, the Common Thick-thumbed Bat is generally found in forested regions with abundant bamboo, primarily at lower elevations, below 1000 m . Food and Feeding. Common Thick-thumbed Bats are insectivorous. Breeding. In the Malay Peninsula, a female gave birth to a single young in mid-April. Activity patterns. Roosts have been found most commonly in dead or damaged bamboo stands as well as in rock crevices and banana leaves. Start frequency has been recorded at 95 kHz followed by a steep FM sweep ending in a QCF sweep at 49-45 kHz. Call durations were recorded at 2-7 milliseconds. Movements, Home range and Social organization. Common Thick-thumbed Bats have been found in a group of twelve in bamboos in Brunei . They may share roosts with species of Tylonycteris , although this is unconfirmed. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Common Thick-thumbed Bat is relatively widespread but difficult to find throughoutits range. It can be found in regenerated forests after disturbance. Relatively little information is available on the species’ ecology, and further research is needed. Bibliography. Csorba (2011), Csorba et al. (2015), Heaney et al. (1998), Heller (1989), Heller & Volleth (1989), Kofron (1994), Mohd-Hanif et al. (2015), Rosell-Ambal et al. (2008), Thewissen & Etnier (1995), Volleth & Yong (1987), Volleth et al. (2001).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Glischropus tylopus	Glischropus		tylopus	Dobson	1875	0	Proc. Zool. Soc. Lond.	1882:53:00	Common Thick-thumbed Bat	<b> batjanus </b>Marschie, 1901.	Malaysia, N Boreneo, Sabah.	Burma, Thailand, W Malaysia, Borneo, Palawan (Philippines), Sumatra and N Molucca Isls.	Not listed.	Least Concern	See Corbet and Hill (1992) and Flannery (1995 b ).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Glischropus tylopus	23	Common Thick-thumbed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PIPISTRELLINI	Glischropus	NA	tylopus	Dobson	1875	1	Vesperugo_tylopus	Dobson, G. E. (1875). Descriptions of new or little-known species of bats of the genus Vesperugo. Proceedings of the Zoological Society of London, 1875, 473.	https://www.biodiversitylibrary.org/item/90418#page/607/mode/1up	BM 1871.2.10.2		"North Borneo," Malaysia.			tylopus (Dobson, 1875)|batjanus (Matschie, 1901)	NA	NA	Thailand|Malaysia|Indonesia|Brunei|Philippines	Asia|Oceania	Indomalaya	LC	0	0	0	Glischropus_tylopus	0	sciname match	Glischropus_tylopus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Glischropus tylopus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Glischropus	tylopus	(Dobson, 1875)	The species was formerly considered to North of the Kra Isthmus in Thailand, but the form there is now recognised as a distinct species, Glischropus bucephalus (Csorba 2011).	20000000	Glischropus tylopus	Least Concern		2021	2021-04-23 00:00:00 UTC	3.1	English	Glischropus tylopus is assessed as Least Concern in view of its wide distribution, presumed large stable population, and because it is unlikely to qualify for listing in a more threatened category.	This insectivorous bat species roosts in rock crevices, in hollow bamboo, and in banana leaves in Peninsular Malaysia (Kofron 1994). It appears to inhabit both primary and secondary forests, often with a high proportion of bamboo (Mohd-Hanif et al. 2015).	There are no known major threats to this species.	There are limited data on the species overall population size, but the species population is suspected to be stable. The species is uncommon throughout most of its range although it can sometimes be locally common. Roosts up to 12 individuals together have been found in bamboo (Kofron 1994).	Stable	The species occurs in extreme southern Thailand (South of the Kra Isthmus), Peninsular Malaysia, Borneo (Malaysia, Indonesia, Brunei), extreme northern Sumatra, Bacan (Maluku) in Indonesia, and Palawan in the Philippines. Is it considered allopatric with other species in the genus such as G. javanus , G. bucephalus and G. aquilus (Csorba 2011, Csorba et al. 2015, Flannery 1995, Heaney et al. 1998).	The species is not known to be hunted, used, or traded.	Terrestrial	There are no species-specific conservation initiatives, although the species is found in a number of protected areas throughout its range. Research is needed to refines its distribution, elevational range, ecology, and population status and trends.	Australasian|Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Glischropus		tylopus	Dobson	1875	0	Proc. Zool. Soc. Lond.	1882:53:00	Common Thick-thumbed Bat	<b> batjanus </b>Marschie, 1901.	Malaysia, N Boreneo, Sabah.	Burma, Thailand, W Malaysia, Borneo, Palawan (Philippines), Sumatra and N Molucca Isls.	Not listed.	Least Concern	See Corbet and Hill (1992) and Flannery (1995 b ).	Glischropus tylopus	1005600	23	Common Thick-thumbed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PIPISTRELLINI	Glischropus	NA	tylopus	Dobson	1875	1	Vesperugo_tylopus	Dobson, G. E. (1875). Descriptions of new or little-known species of bats of the genus Vesperugo. Proceedings of the Zoological Society of London, 1875, 473.	https://www.biodiversitylibrary.org/item/90418#page/607/mode/1up	BM 1871.2.10.2		"North Borneo," Malaysia.			tylopus (Dobson, 1875)|batjanus (Matschie, 1901)	NA	NA				Thailand|Malaysia|Indonesia|Brunei|Philippines	Asia|Oceania	Indomalaya	LC	0	0	0	Glischropus_tylopus	0	sciname match	Glischropus_tylopus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Glischropus_tylopus	1005600	23	Common Thick-thumbed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Pipistrellini	Glischropus	NA	tylopus	Dobson	1	Vesperugo tylopus	Dobson, G.E. 1875-10. Descriptions of new or little-known species of bats of the genus _Vesperugo_. Proceedings of the Zoological Society of London 1875(3):470-474.	https://www.biodiversitylibrary.org/page/28501722	BMNH:Mamm:1871.2.10.2	holotype	https://data.nhm.ac.uk/object/ba972ec7-ff6f-4dbd-bb52-3003d7390043	"North Borneo," Malaysia.			NA	NA				Thailand|Malaysia|Indonesia|Brunei|Philippines	Asia|Oceania (Continent)	Indomalaya	LC	0	0	0	Glischropus_tylopus	0	sciname match	Glischropus_tylopus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Glischropus		tylopus	Dobson	1875	0	Proc. Zool. Soc. Lond.	1882:53:00	Common Thick-thumbed Bat	batjanus Marschie, 1901.	Malaysia, N Boreneo, Sabah.	Burma, Thailand, W Malaysia, Borneo, Palawan (Philippines), Sumatra and N Molucca Isls.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/81187867/22105878/' target='_blank'>Least Concern</a>	See Corbet and Hill (1992) and Flannery (1995b).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Glischropus tylopus; Glischropus tylopus; Glischropus tylopus; Glischropus tylopus; Glischropus tylopus; Glischropus tylopus; tylopus; batjanus; batjanus; batjanus; tylopus; batjanus; Pipistrelle de Dobson; Gewohnliche Dickdaumenfledermaus; Pipistrela de Dobson; Common Thick-thumbed Bat; Common Thick-thumbed Bat; Common Thick-thumbed Bat; G. tylopus
