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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L333	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Glauconycteris variegata	Chalinolobus variegatus	Glauconycteris variegata	Chalinolobus variegatus	Chalinolobus variegatus	Glauconycteris variegata	Glauconycteris variegata	Glauconycteris variegata	Glauconycteris variegata	Glauconycteris variegata	Glauconycteris variegata	Glauconycteris variegata	Glauconycteris variegata	Glauconycteris variegata	Glauconycteris variegata		[MSW2] Subgenus Glauconycteris. Includes machadoi; see Koopman (1971:6).; [MSW3] Does not include machadoi, see Hayman and Hill (1971), but also see Koopman (1971a).; [HMW] Scotophilus variegatus Tomes, 1861 , “Otjoro,” Namibia . Analysis of mitochondrial genes by A. Hassanin and colleagues in 2018 retrieved G. variegata as sister to all other species of Glauconycteris . Two subspecies recognized.; [batnames2022] Does not include machadoi , see Hayman and Hill (1971), but also see Koopman (1971 a ).; [IUCN] Two subspecies are currently recognized, but their validity remains uncertain (Happold 2013): Glauconycteris variegata phalaena from Sudan and Somalia, and G . v . variegata , which is distributed through the rest of the species range; including the assessment region (Skinner &; Chimimba 2005). Glauconycteris machadoi is sometimes considered a melanistic subspecies of G . variegata (Monadjem et al. 2010), but is considered a distinct species by Happold (2013).; [batnames2023] Does not include machadoi , see Hayman and Hill (1971), but also see Koopman (1971 a ).; [batnames2025_1.7] Does not include machadoi, see Hayman and Hill (1971), but also see Koopman (1971a).				machadoi		machadoi, papilio, phalaena.	phalaena, machadoi, variegatus	variegata, phalaena	papilio	variegata, phalaena		variegata, phalaena	variegata- papilio	variegata, papilio, phalaena	Two subspecies are currently recognized, but their validity remains uncertain (Happold 2013): Glauconycteris variegata phalaena from Sudan and Somalia, and G . v . variegata , which is distributed through the rest of the species range; including the assessment region (Skinner &; Chimimba 2005). Glauconycteris machadoi is sometimes considered a melanistic subspecies of G . variegata (Monadjem et al. 2010), but is considered a distinct species by Happold (2013).	variegata, phalaena	variegata- papilio	variegata, papilio, phalaena	variegata, papilio, phalaena	phalaena, variagata	variagata - papilio	variegata (Tomes, 1861)|papilio O. Thomas, 1905|phalaena O. Thomas, 1915		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Butterfly bat	Ghana – Somalia – Namibia, Mozambique	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Chalinolobus variegatus	Namibia, Otjoro.	Tomes	1861	Proc. Zool. Soc. Lond., 1861:36.	Distribution: Ranging widely in the savanna regions of Africa from Senegal to Ethiopia and south to Natal and northern Namibia.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Butterfly bat	Senegal – Somalia – S Africa	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Tomes	1861	Proc. Zool. Soc. Lond., 1861:36.	Subgenus Glauconycteris. Includes machadoi; see Koopman (1971:6).	Senegal to Somalia, south to South Africa.	Namibia, Otjoro.		TOMES	1861	Plagiopatagium and uropatagium pale but with a prominent dark reticulated pattern. Fur varying from light to dark brown without markings. Ear pinna rounded. Size fairly large (forearm length, 38-46 mm). Inner upper incisor varying from unicuspid to bicuspid (but with unequal cusps).	Distribution: Ranging widely in the savanna regions of Africa from Senegal to Ethiopia and south to Natal and northern Namibia.	Three subspecies are here recognized:	C. v. phalaena (central Sudan), C. v. machadoi (east-central Angola), C. v. variegatus (remainder of range).	125	species	C. variegatus	TOMES	1861	Glauconycteris	subgenus	Chalinolobus variegatus				Plagiopatagium and uropatagium pale but with a prominent dark reticulated pattern. Fur varying from light to dark brown without markings. Ear pinna rounded. Size fairly large (forearm length, 38-46 mm). In- ner upper incisor varying from unicuspid to bicuspid (but with unequal cusps).	Three subspecies are here recognized:		13. C. variegatus (TOMES 1861).	13	_G. v. phalaena_ Thomas, 1915; _G. v. variegata_ (Tomes, 1861) (synonyms: _papilio_ Thomas, 1905)			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Vespertilionini	Glauconycteris variegata	Glauconycteris		variegata	Tomes	y	1861		Proc. Zool. Soc. Lond.	1861		36		Variegated Butterfly Bat	Namibia, Otjoro.	Senegal to Somalia, south to South Africa.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc) as Chalinolobus variegatus.	papilio Thomas, 1915; phalaena Thomas, 1915.	Does not include machadoi, see Hayman and Hill (1971), but also see Koopman (1971a).	4C3D87E8FFB26A0DFF8097F61658B87E	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	831	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFB26A0DFF8097F61658B87E.xml	Glauconycteris variegata	Vespertilionidae	Glauconycteris	variegata		1861	Glauconyctere réticulé @fr | Genetzte Schmetterlingsfledermaus @de | Glauconicterioreticulado @es | Butterfly Bat @en | Leaf-winged Bat @en	Scotophilus variegatus Tomes, 1861 , “Otjoro,” Namibia . Analysis of mitochondrial genes by A. Hassanin and colleagues in 2018 retrieved G. variegata as sister to all other species of Glauconycteris . Two subspecies recognized.	G. v. variegata Tomes, 1861 — widely but patchily distributed throughout much of sub-Saharan Africa, from Atlantic coast of Senegal E to Ethiopia and S to NE Namibia , N Botswana , and NE South Africa . G. v. phalaena Thomas, 1915 — SC Sudan and Somalia .	Head-body ¢.48-62 mm, tail 40-53 mm, ear 11-14 mm, hindfoot 7-10 mm, forearm 38-45 mm; weight 5-14 g. Females are apparently larger than males in body dimensions but not body weight. Pelage is dense and soft, with 7-10 mm middorsal hairs. Dorsal fur lacks markings and ranges from pale creamy buff to yellowish fawn, paler on head and neck in some individuals. Hairs are pale creamy buff or yellowish fawn with paler bases or creamy white with brown bases and yellowish fawn tips. Ventral fur ranges from yellowish cream to pure white, suffused with pale gray particularly on throat in some individuals. Ears are light brown, separated, and shortfor a vespertilionid, with outer margins connecting with fleshy lobe near corner of mouth and inner margins with moderately long backward-pointing lobe at base. Tragus has straight anterior margin and convex posterior margin, with inconspicuous lobe at base. Eyes are very small. Nostrils are subcylindrical and open laterally. Well-developed glandular protuberances separate nostrils by deep groove,giving swollen appearance to muzzle. Flight membranes are conspicuously reticulated. Wings and uropatagium are pale yellowish orange, with dark brown pigment outlining bones and venation. Tibia is long (18-21 mm). Baculum is very small and triangular. Skull is moderately large and has high broad braincase, short obtuse rostrum, and domed cranium without sagittal crest. Profile of forehead region is weakly concave compared with other species of Glauconycteris . Supraoccipital crest is poorly developed, and frontal region of skull is not hollowed. Basisphenoid pits are shallow and indistinct. I* is longer than I’, and they vary from unicuspid to tricuspid. I° is almost rudimentary. I is trifid or rudimentary quadrifid, with moderately developed cingulum. Incisors are pressed tightly between canines. I, and I, are much smaller than I, and coneshaped with broad, evenly developed cingula. P* is large, well developed, and carnassial. P is absent. P, and P, are small and conical, with prominent cingula that form anteroposterior cusps. Dental formula for all species of Glauconycterisis 12/3, C1/1,P 1/2, M 3/3 (<2) = 32. Chromosomal complement has 2n = 18 and FN = 32, with two pairs of metacentric, five pairs of submetacentric, and one pair of subtelocentric autosomes. X-chromosomeis medium-sized subtelocentric, and Y-chromosomeis small metacentric.	Mainly savanna habitats, including open woodland savannas and open bush country known from elevations up to ¢. 1000 m . One female Variegated Butterfly Bat was caught in riverine forest near hot, humid, marshy grassland in Ethiopia , and another individual was caughtin dense riparian forest in Pafuri, north-eastern South Africa .	The Variegated Butterfly Bat forages by moderately fast hawking, high aboveground, mainly in uncluttered habitats, but also between trees, around lights, close to the ground, and over water. It eats airborne prey, primarily soft-bodied insects.	The Variegated Butterfly Bat seems to be monoestrous, but seasonal reproductive cycle might vary acrossits distribution. Sexually active males were reported in July and Octoberin southern Uganda . In north-eastern DR Congo (3-4° N), three of ten females were pregnant in March, with parturition predicted in March-April. In Nairobi, Kenya ( 1°17° S ), a lactating female was recorded in April. In Zimbabwe ( 16— 20° S ), two pregnant females each carrying one fetus were recorded in August, and six females each with a newborn were recorded in early November. Breeding season varies across latitudes, ranging from an early boreal cycle (parturition in March-April) in the north to a late austral cycle (parturition around November) for southern populations. In northern latitudes, gestation lasts ¢.3 months, parturition occurs in April, and young are weaned through May. Litter size is 1-2, but usually one.	During the day, the Variegated Butterfly Bat roosts mainly in dense clusters ofleaves in trees. It also roosts in thatch of abandoned huts. It has been found in lychee trees (Lutchi, Sapindaceae ), in a Natal mahogany tree ( Trichilia emetica , Meliaceae ) 7-5 m aboveground, and on a low branch of a mango tree ( Mangifera , Anacardiaceae ). The species remains motionless and is reluctant to fly when disturbed. It has been observed leaving roosts in daylight (two hours before sunset) or in the beginning of the night. Aspect ratio is low, wing loading is low to very low, and wingtips are long and pointed, suggesting efficient low speed flight, but they can fly moderately fast, with moderate maneuverability. Minimum and maximum power speeds are 3-3 m/s and 4-4 m/s, respectively, and mean foraging flight speed is 4 m /s. Variegated Butterfly Bats can take off from ground. They emit audible squeaks (social calls) during flight. Echolocation vocalizationsare highintensity FM calls sweeping from 70 kHz to 30 kHz, sometimes including a second harmonic and rarely a third. Two foraging bats from Zimbabwe had smooth steep-shallow FM search-phase call shape, with mean start frequency of 53-4 kHz, mean end frequency of 33-7 kHz, mean peak frequency of37-7 kHz, mean bandwidth of 19-7 kHz, and mean call duration of 5-3 milliseconds. Approach-phase calls are shorter and broader in bandwidth.	Variegated Butterfly Bats roost alone, in pairs, or in groups of 3-12 individuals.	Classified as Least Concern on The IUCN Red List.	Happold, M. (2013cb) | Fenton (2001) | Hassanin et al. (2018) | Hayman & Hill (1971) | Koopman (1971a, 1993) | Rambaldini (2010) | Simmons (2005) | Thomas (1905a)	https://zenodo.org/record/6612150/files/figure.png	144. Variegated Butterfly Bat Glauconycteris variegata French: Glauconyctere réticulé / German: Genetzte Schmetterlingsfledermaus / Spanish: Glauconicterio reticulado Other common names: Butterfly Bat , Leaf-winged Bat Taxonomy. Scotophilus variegatus Tomes, 1861 , “Otjoro,” Namibia . Analysis of mitochondrial genes by A. Hassanin and colleagues in 2018 retrieved G. variegata as sister to all other species of Glauconycteris . Two subspecies recognized. Subspecies and Distribution. G. v. variegata Tomes, 1861 — widely but patchily distributed throughout much of sub-Saharan Africa, from Atlantic coast of Senegal E to Ethiopia and S to NE Namibia , N Botswana , and NE South Africa . G. v. phalaena Thomas, 1915 — SC Sudan and Somalia . Descriptive notes. Head-body ¢.48-62 mm, tail 40-53 mm, ear 11-14 mm, hindfoot 7-10 mm, forearm 38-45 mm; weight 5-14 g. Females are apparently larger than males in body dimensions but not body weight. Pelage is dense and soft, with 7-10 mm middorsal hairs. Dorsal fur lacks markings and ranges from pale creamy buff to yellowish fawn, paler on head and neck in some individuals. Hairs are pale creamy buff or yellowish fawn with paler bases or creamy white with brown bases and yellowish fawn tips. Ventral fur ranges from yellowish cream to pure white, suffused with pale gray particularly on throat in some individuals. Ears are light brown, separated, and shortfor a vespertilionid, with outer margins connecting with fleshy lobe near corner of mouth and inner margins with moderately long backward-pointing lobe at base. Tragus has straight anterior margin and convex posterior margin, with inconspicuous lobe at base. Eyes are very small. Nostrils are subcylindrical and open laterally. Well-developed glandular protuberances separate nostrils by deep groove,giving swollen appearance to muzzle. Flight membranes are conspicuously reticulated. Wings and uropatagium are pale yellowish orange, with dark brown pigment outlining bones and venation. Tibia is long (18-21 mm). Baculum is very small and triangular. Skull is moderately large and has high broad braincase, short obtuse rostrum, and domed cranium without sagittal crest. Profile of forehead region is weakly concave compared with other species of Glauconycteris . Supraoccipital crest is poorly developed, and frontal region of skull is not hollowed. Basisphenoid pits are shallow and indistinct. I* is longer than I’, and they vary from unicuspid to tricuspid. I° is almost rudimentary. I is trifid or rudimentary quadrifid, with moderately developed cingulum. Incisors are pressed tightly between canines. I, and I, are much smaller than I, and coneshaped with broad, evenly developed cingula. P* is large, well developed, and carnassial. P is absent. P, and P, are small and conical, with prominent cingula that form anteroposterior cusps. Dental formula for all species of Glauconycterisis 12/3, C1/1,P 1/2, M 3/3 (<2) = 32. Chromosomal complement has 2n = 18 and FN = 32, with two pairs of metacentric, five pairs of submetacentric, and one pair of subtelocentric autosomes. X-chromosomeis medium-sized subtelocentric, and Y-chromosomeis small metacentric. Habitat. Mainly savanna habitats, including open woodland savannas and open bush country known from elevations up to ¢. 1000 m . One female Variegated Butterfly Bat was caught in riverine forest near hot, humid, marshy grassland in Ethiopia , and another individual was caughtin dense riparian forest in Pafuri, north-eastern South Africa . Food and Feeding. The Variegated Butterfly Bat forages by moderately fast hawking, high aboveground, mainly in uncluttered habitats, but also between trees, around lights, close to the ground, and over water. It eats airborne prey, primarily soft-bodied insects. Breeding. The Variegated Butterfly Bat seems to be monoestrous, but seasonal reproductive cycle might vary acrossits distribution. Sexually active males were reported in July and Octoberin southern Uganda . In north-eastern DR Congo (3-4° N), three of ten females were pregnant in March, with parturition predicted in March-April. In Nairobi, Kenya ( 1°17° S ), a lactating female was recorded in April. In Zimbabwe ( 16— 20° S ), two pregnant females each carrying one fetus were recorded in August, and six females each with a newborn were recorded in early November. Breeding season varies across latitudes, ranging from an early boreal cycle (parturition in March-April) in the north to a late austral cycle (parturition around November) for southern populations. In northern latitudes, gestation lasts ¢.3 months, parturition occurs in April, and young are weaned through May. Litter size is 1-2, but usually one. Activity patterns. During the day, the Variegated Butterfly Bat roosts mainly in dense clusters ofleaves in trees. It also roosts in thatch of abandoned huts. It has been found in lychee trees (Lutchi, Sapindaceae ), in a Natal mahogany tree ( Trichilia emetica , Meliaceae ) 7-5 m aboveground, and on a low branch of a mango tree ( Mangifera , Anacardiaceae ). The species remains motionless and is reluctant to fly when disturbed. It has been observed leaving roosts in daylight (two hours before sunset) or in the beginning of the night. Aspect ratio is low, wing loading is low to very low, and wingtips are long and pointed, suggesting efficient low speed flight, but they can fly moderately fast, with moderate maneuverability. Minimum and maximum power speeds are 3-3 m/s and 4-4 m/s, respectively, and mean foraging flight speed is 4 m /s. Variegated Butterfly Bats can take off from ground. They emit audible squeaks (social calls) during flight. Echolocation vocalizationsare highintensity FM calls sweeping from 70 kHz to 30 kHz, sometimes including a second harmonic and rarely a third. Two foraging bats from Zimbabwe had smooth steep-shallow FM search-phase call shape, with mean start frequency of 53-4 kHz, mean end frequency of 33-7 kHz, mean peak frequency of37-7 kHz, mean bandwidth of 19-7 kHz, and mean call duration of 5-3 milliseconds. Approach-phase calls are shorter and broader in bandwidth. Movements, Home range and Social organization. Variegated Butterfly Bats roost alone, in pairs, or in groups of 3-12 individuals. Status and Conservation. Classified as Least Concern on The IUCN Red List. Bibliography. Happold, M. (2013cb), Fenton (2001), Hassanin et al. (2018), Hayman & Hill (1971), Koopman (1971a, 1993), Rambaldini (2010), Simmons (2005), Thomas (1905a).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Glauconycteris variegata	Glauconycteris		variegata	Tomes	1861	1	Proc. Zool. Soc. Lond.	######	Variegated Butterfly Bat	 papilio Thomas, 1915; <b> phalaena </b> Thomas, 1915.	Namibia, Otjoro.	Senegal to Somalia, south to South Africa.	Not listed.	Least Concern	Does not include machadoi , see Hayman and Hill (1971), but also see Koopman (1971 a ).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Glauconycteris variegata	23	Variegated Butterfly Bat	Butterfly Bat|Leaf-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	EPTESICINI	Glauconycteris	NA	variegata	Tomes	1861	1	Scotophilus_variegatus	Tomes, R. F. (1861). Notes on a collection of bats made by Mr. Andersson in the Damara Country, South-western Africa, with notices of some other African species. Proceedings of the Zoological Society London, 1861, 36.	https://www.biodiversitylibrary.org/item/91152#page/64/mode/1up	BM 1907.1.1.437, BM 1907.1.1.438, ZMB 3033 [syntypes]		"Otjoro," Namibia.			variegata (Tomes, 1861)|papilio O. Thomas, 1905|phalaena O. Thomas, 1915	NA	NA	Senegal|Gambia|Guinea|Mali|Côte d'Ivoire|Ghana|Togo|Benin|Nigeria|Cameroon|Chad|Central African Republic|Sudan|South Sudan|Ethiopia|Somalia|Kenya|Uganda|Rwanda|Burundi|Gabon|Republic of the Congo|Democratic Republic of the Congo|Tanzania|Angola|Zambia|Malawi|Mozambique|Zimbabwe|Namibia|Botswana|South Africa|Eswatini	Africa	Afrotropic	LC	0	0	0	Glauconycteris_variegata	0	sciname match	Glauconycteris_variegata	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	44800	Glauconycteris variegata	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Glauconycteris	variegata	(Tomes, 1861)	Two subspecies are currently recognized, but their validity remains uncertain (Happold 2013): Glauconycteris variegata phalaena from Sudan and Somalia, and G . v . variegata , which is distributed through the rest of the species range; including the assessment region (Skinner &; Chimimba 2005). Glauconycteris machadoi is sometimes considered a melanistic subspecies of G . variegata (Monadjem et al. 2010), but is considered a distinct species by Happold (2013).	20000000	Glauconycteris variegata	Least Concern		2017	2016-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category	This species is associated with open savanna woodland, bushland and riverine woodland (Skinner and Chimimba 2005). Animals are generally considered to be absent from areas of closed forest, although Rosevear (1965) suggests that the species is present in rainforest. Roosting colonies, containing a few pairs of animals, have been found in thatched roofs of abandoned huts and among dense vegetation (Allen et al. 1917; Ansell and Dowsett 1988).Taylor, 1998	In view of the species wide range and availability of extensive suitable habitat, there are no major threats to this species. This species is threatened in parts of its range by habitat loss resulting from logging activities and the conversion of land to agricultural use.	Colonies of this species are small, and there are believed to be less than 10,000 individuals in total.	Unknown	This species is widely, but patchily, recorded throughout much of sub-Saharan Africa. It ranges from Senegal and the Gambia in West Africa, through West and Central Africa to Ethiopia and Somalia in the east, from here it ranges south through East Africa and southern Africa, being recorded as far south as northeastern South Africa.		Terrestrial	There appear to be no direct conservation measures in place. In view of the species wide distribution it is presumably present in some protected areas.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Glauconycteris		variegata	Tomes	1861	1	Proc. Zool. Soc. Lond.	########	Variegated Butterfly Bat	 papilio Thomas, 1915; <b> phalaena </b> Thomas, 1915.	Namibia, Otjoro.	Senegal to Somalia, south to South Africa.	Not listed.	Least Concern	Does not include machadoi , see Hayman and Hill (1971), but also see Koopman (1971 a ).	Glauconycteris variegata	1005547	23	Variegated Butterfly Bat	Butterfly Bat|Leaf-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	EPTESICINI	Glauconycteris	NA	variegata	Tomes	1861	1	Scotophilus_variegatus	Tomes, R. F. (1861). Notes on a collection of bats made by Mr. Andersson in the Damara Country, South-western Africa, with notices of some other African species. Proceedings of the Zoological Society London, 1861, 36.	https://www.biodiversitylibrary.org/item/91152#page/64/mode/1up	BM 1907.1.1.437, BM 1907.1.1.438, ZMB 3033 [syntypes]		"Otjoro," Namibia.			variegata (Tomes, 1861)|papilio O. Thomas, 1905|phalaena O. Thomas, 1915	NA	NA				Senegal|Gambia|Guinea|Mali|Côte d'Ivoire|Ghana|Togo|Benin|Nigeria|Cameroon|Chad|Central African Republic|Sudan|South Sudan|Ethiopia|Somalia|Kenya|Uganda|Rwanda|Burundi|Gabon|Republic of the Congo|Democratic Republic of the Congo|Tanzania|Angola|Zambia|Malawi|Mozambique|Zimbabwe|Namibia|Botswana|South Africa|Eswatini	Africa	Afrotropic	LC	0	0	0	Glauconycteris_variegata	0	sciname match	Glauconycteris_variegata	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Glauconycteris_variegata	1005547	23	Variegated Butterfly Bat	Butterfly Bat|Leaf-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Nycticeiini	Glauconycteris	NA	variegata	Tomes	1	Scotophilus variegatus	Tomes, R.F. 1861-05. Notes on a collection of bats made by Mr. Andersson in the Damara Country, south-western Africa, with notices of some other African species. Proceedings of the Zoological Society of London 1861(1):31-40.	https://www.biodiversitylibrary.org/page/28672476	BMNH:Mamm:1907.1.1.437, BMNH:Mamm:1907.1.1.438, ZMB 3033	syntypes	https://data.nhm.ac.uk/object/e249dcdd-277b-46cb-83f3-76b526758595 | https://data.nhm.ac.uk/object/e97ab1d3-8817-4024-b87e-9db8d263e981	"Otjoro," Namibia.			NA	NA				Senegal|Gambia|Guinea|Mali|Cote d'Ivoire|Ghana|Togo|Benin|Nigeria|Cameroon|Chad|Central African Republic|Sudan|South Sudan|Ethiopia|Somalia|Kenya|Uganda|Rwanda|Burundi|Gabon|Republic of the Congo|Democratic Republic of the Congo|Tanzania|Angola|Zambia|Malawi|Mozambique|Zimbabwe|Namibia|Botswana|South Africa|Eswatini	Africa	Afrotropic	LC	0	0	0	Glauconycteris_variegata	0	sciname match	Glauconycteris_variegata	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Glauconycteris		variegata	Tomes	1861	1	Proc. Zool. Soc. Lond.	########	Variegated Butterfly Bat	papilio Thomas, 1915; phalaena Thomas, 1915.	Namibia, Otjoro.	Senegal to Somalia, south to South Africa.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/44800/22069727/' target='_blank'>Least Concern</a>	Does not include machadoi, see Hayman and Hill (1971), but also see Koopman (1971a).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Glauconycteris variegata; Glauconycteris variegata; Glauconycteris variegata; Glauconycteris variegata; Glauconycteris variegata; Glauconycteris variegata; variegata; phalaena; papilio; variegata; phalaena; phalaena; papilio; variegata; papilio; phalaena; Glauconyctere réticulé; Genetzte Schmetterlingsfledermaus; Glauconicterioreticulado; Butterfly Bat; Leaf-winged Bat; Variegated Butterfly Bat; Butterfly Bat; Leaf-winged Bat; Variegated Butterfly Bat; Variegated Butterfly Bat; G. variegata