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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L328	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	humeralis	N/A	Chalinolobus beatrix [synonym of]	Chalinolobus beatrix humeralis	Glauconycteris humeralis	Glauconycteris humeralis	Glauconycteris humeralis	Glauconycteris humeralis	Glauconycteris humeralis	Glauconycteris humeralis	Glauconycteris humeralis	Glauconycteris humeralis	Glauconycteris humeralis	Glauconycteris humeralis		[MSW3] Apparently distinct from beatrix; see Hill and Harrison (1987) and Heller et al. (1994), but also see Eger and Schlitter (2001).; [HMW] Glauconycteris humeralis]. A. Allen, 1917 , “Medje [Haut-Uele Province], Belgian Congo [= DR Congo ].” Form hwmeralis has been treated as a species and a subspecies of G. beatrix . Here it is treated as distinct from G. beatrix , pending revision ofits taxonomic status. A. Hassanin and colleagues in 2018 retrieved G. humeralis as possibly polyphyletic, with partsister to a clade including G. atra, G. cf. humeralis , and G. cf. beatrix . Monotypic.; [batnames2022] Apparently distinct from beatrix ; see Hill and Harrison (1987) and Heller et al. (1994), but also see Eger and Schlitter (2001).; [IUCN] Distinct from beatrix .; [batnames2023] Apparently distinct from beatrix ; see Hill and Harrison (1987) and Heller et al. (1994), but also see Eger and Schlitter (2001).; [batnames2025_1.7] Apparently distinct from beatrix; see Hill and Harrison (1987) and Heller et al. (1994), but also see Eger and Schlitter (2001).														humeralis	Distinct from beatrix .			humeralis	humeralis			humeralis J. A. Allen in J. A. Allen, Lang, & Chapin, 1917						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Vespertilionini	Glauconycteris humeralis	Glauconycteris		humeralis	J. A. Allen		1917		Bull. Am. Mus. Nat. Hist.	37		448		Spotted Butterfly Bat	Dem. Rep. Congo, Oriental, Medje.	Dem. Rep. Congo, Uganda, Kenya.	IUCN 2003 – Not evaluated; not considered in IUCN/SSC Action Plan (2001).		Apparently distinct from beatrix; see Hill and Harrison (1987) and Heller et al. (1994), but also see Eger and Schlitter (2001).	4C3D87E8FFB06A08FA919A111D95BEC7	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	833	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFB06A08FA919A111D95BEC7.xml	Glauconycteris humeralis	Vespertilionidae	Glauconycteris	humeralis	J. A. Allen	1917	Glauconyctere tacheté @fr | Gefleckte Schmetterlingsfledermaus @de | Glauconicteriomanchado @es | Allen's Spotted Bat @en	Glauconycteris humeralis]. A. Allen, 1917 , “Medje [Haut-Uele Province], Belgian Congo [= DR Congo ].” Form hwmeralis has been treated as a species and a subspecies of G. beatrix . Here it is treated as distinct from G. beatrix , pending revision ofits taxonomic status. A. Hassanin and colleagues in 2018 retrieved G. humeralis as possibly polyphyletic, with partsister to a clade including G. atra, G. cf. humeralis , and G. cf. beatrix . Monotypic.	Confirmed records from DR Congo , Uganda , and Tanzania ; it might occur in Kenya .	Head-body ¢.39-45 mm, tail 37-59 mm, ear 6-12 mm, forearm 35-41 mm; weight 4-7 g. Pelage ofthe Spotted Butterfly Bat is dense. Dorsal and ventral fur is dark sepia-brown, with white shoulder spot. Ears are yellowish brown, very rounded, separate, and short for a vespertilionid but medium-short for a Glauconycteris . Inner margin has rounded lobe at base, and outer margin is connected with well-developed fleshy lobe on lowerlip near corner of mouth. Tragusis short and broad, with straight anterior margin and convex posterior margin. Wings and uropatagium are dark brown and not reticulated. Wing bones are pale brown on both sides. Tibia is of medium length (16-20 mm) for Glauconycteris . Basal part of baculum is not expanded and unilobed, and distal part is lengthened to a short shaft. Head is high-domed, and muzzle is short, broad, and flattish. Skull is small, and profile of forehead is moderately concave compared with other Glauconycteris . I* is strongly bicuspid, with secondary cusp equal in size or slightly smaller. Lower incisors are tricuspid or with four cusps and not crowded.	Rainforests.	No information.	A pregnant Spotted Butterfly Bat was recorded in February in Avakubi, north-eastern DR Congo .	No information.	No information.	Classified as Data Deficient on The IUCN Red List. Possible threats include deforestation for logging, mining activities, and agriculture.	Eger & Schlitter (2001) | Happold, M. (2013ay) | Hassanin et al. (2018) | Hayman & Hill (1971) | Hayman et al. (1966) | Koopman (1971a, 1993) | Peterson & Smith (1973)	https://zenodo.org/record/6398093/files/figure.png	154. Spotted Butterfly Bat Glauconycteris humeralis French: Glauconyctere tacheté / German: Gefleckte Schmetterlingsfledermaus / Spanish: Glauconicterio manchado Other common names: Allen's Spotted Bat Taxonomy. Glauconycteris humeralis]. A. Allen, 1917 , “Medje [Haut-Uele Province], Belgian Congo [= DR Congo ].” Form hwmeralis has been treated as a species and a subspecies of G. beatrix . Here it is treated as distinct from G. beatrix , pending revision ofits taxonomic status. A. Hassanin and colleagues in 2018 retrieved G. humeralis as possibly polyphyletic, with partsister to a clade including G. atra, G. cf. humeralis , and G. cf. beatrix . Monotypic. Distribution. Confirmed records from DR Congo , Uganda , and Tanzania ; it might occur in Kenya . Descriptive notes. Head-body ¢.39-45 mm, tail 37-59 mm, ear 6-12 mm, forearm 35-41 mm; weight 4-7 g. Pelage ofthe Spotted Butterfly Bat is dense. Dorsal and ventral fur is dark sepia-brown, with white shoulder spot. Ears are yellowish brown, very rounded, separate, and short for a vespertilionid but medium-short for a Glauconycteris . Inner margin has rounded lobe at base, and outer margin is connected with well-developed fleshy lobe on lowerlip near corner of mouth. Tragusis short and broad, with straight anterior margin and convex posterior margin. Wings and uropatagium are dark brown and not reticulated. Wing bones are pale brown on both sides. Tibia is of medium length (16-20 mm) for Glauconycteris . Basal part of baculum is not expanded and unilobed, and distal part is lengthened to a short shaft. Head is high-domed, and muzzle is short, broad, and flattish. Skull is small, and profile of forehead is moderately concave compared with other Glauconycteris . I* is strongly bicuspid, with secondary cusp equal in size or slightly smaller. Lower incisors are tricuspid or with four cusps and not crowded. Habitat. Rainforests. Food and Feeding. No information. Breeding. A pregnant Spotted Butterfly Bat was recorded in February in Avakubi, north-eastern DR Congo . Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The IUCN Red List. Possible threats include deforestation for logging, mining activities, and agriculture. Bibliography. Eger & Schlitter (2001), Happold, M. (2013ay), Hassanin et al. (2018), Hayman & Hill (1971), Hayman et al. (1966), Koopman (1971a, 1993), Peterson & Smith (1973).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Glauconycteris humeralis	Glauconycteris		humeralis	J. A. Allen	1917	0	Bull. Am. Mus. Nat. Hist.	44:28:00	Spotted Butterfly Bat	None.	Dem. Rep. Congo, Oriental, Medje.	Dem. Rep. Congo, Uganda, Kenya.	Not listed.	Data Deficient	Apparently distinct from beatrix ; see Hill and Harrison (1987) and Heller et al. (1994), but also see Eger and Schlitter (2001).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Glauconycteris humeralis	23	Spotted Butterfly Bat	Allen's Spotted Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	EPTESICINI	Glauconycteris	NA	humeralis	J. A. Allen	1917	0	Glauconycteris_humeralis	Allen, J. A. (1917). I. Systematic List. In J. A. Allen, H. Lang & J. P. Chapin. The American Museum Congo Expedition collection of bats. Bulletin of the American Museum of Natural History, 37, 448.	http://digitallibrary.amnh.org/bitstream/handle/2246/1068/B37a18.pdf?sequence=1&isAllowed=y	AMNH 49013		"Medje [Haut-Uele Province], Belgian Congo [= DR Congo]."			humeralis J. A. Allen, 1917	NA	NA	Democratic Republic of the Congo|Uganda|Tanzania|Kenya?	Africa	Afrotropic	DD	0	0	0	Glauconycteris_humeralis	0	sciname match	Glauconycteris_humeralis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	44795	Glauconycteris humeralis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Glauconycteris	humeralis	J.A. Allen, 1917	Distinct from beatrix .	20000000	Glauconycteris humeralis	Data Deficient		2019	2018-08-31 00:00:00 UTC	3.1	English	Confirmed as Data Deficient in view of the absence of sufficient information on its extent of occurrence, natural history, threats and conservation status.	This species is associated with lowland tropical moist forest. Presumably it roosts within hollow trees and dense vegetation, although this needs confirmation.	The threats to this species are not known. It may be threatened within parts of its range by deforestation resulting from logging and mining activities, and the conversion of forest to farmland.	Little information is available on the population abundance or size of this species.	Unknown	This little known species has been recorded in eastern Democratic Republic of the Congo and Uganda. It has been reported from Kenya, but no specific locality information is available (Aggundey and Schlitter 1984).		Terrestrial	There appear to be no direct conservation measures in place. It is not known if the species is present in any protected areas. Further research is needed into the distribution, natural history and threats to this poorly known forest species.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Glauconycteris		humeralis	J. A. Allen	1917	0	Bull. Am. Mus. Nat. Hist.	44:28:00	Spotted Butterfly Bat	None.	Dem. Rep. Congo, Oriental, Medje.	Dem. Rep. Congo, Uganda, Kenya.	Not listed.	Data Deficient	Apparently distinct from beatrix ; see Hill and Harrison (1987) and Heller et al. (1994), but also see Eger and Schlitter (2001).	Glauconycteris humeralis	1005542	23	Spotted Butterfly Bat	Allen's Spotted Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	EPTESICINI	Glauconycteris	NA	humeralis	J. A. Allen	1917	0	Glauconycteris_humeralis	Allen, J. A. (1917). I. Systematic List. In J. A. Allen, H. Lang & J. P. Chapin. The American Museum Congo Expedition collection of bats. Bulletin of the American Museum of Natural History, 37, 448.	http://digitallibrary.amnh.org/bitstream/handle/2246/1068/B37a18.pdf?sequence=1&isAllowed=y	AMNH 49013		"Medje [Haut-Uele Province], Belgian Congo [= DR Congo]."			humeralis J. A. Allen, 1917	NA	NA				Democratic Republic of the Congo|Uganda|Tanzania|Kenya?	Africa	Afrotropic	DD	0	0	0	Glauconycteris_humeralis	0	sciname match	Glauconycteris_humeralis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Glauconycteris_humeralis	1005542	23	Spotted Butterfly Bat	Allen's Spotted Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Nycticeiini	Glauconycteris	NA	humeralis	J. A. Allen in J. A. Allen, Lang, & Chapin	0	Glauconycteris humeralis	Allen, J.A., Lang, H. and Chapin, J.P. 1917-09-29. The American Museum Congo Expedition collection of bats. Bulletin of the American Museum of Natural History 37(18):405-563.	https://hdl.handle.net/2246/1068	AMNH M-49013	holotype	http://portal.vertnet.org/o/amnh/mammals?id=urn-catalog-amnh-mammals-m-49013	"Medje [Haut-Uele Province], Belgian Congo [= DR Congo]."			NA	NA				Democratic Republic of the Congo|Uganda|Tanzania|Kenya?	Africa	Afrotropic	DD	0	0	0	Glauconycteris_humeralis	0	sciname match	Glauconycteris_humeralis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Glauconycteris		humeralis	J. A. Allen	1917	0	Bull. Am. Mus. Nat. Hist.	44:28:00	Spotted Butterfly Bat	None.	Dem. Rep. Congo, Oriental, Medje.	Dem. Rep. Congo, Uganda, Kenya.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/44795/22070303/' target='_blank'>Data Deficient</a>	Apparently distinct from beatrix; see Hill and Harrison (1987) and Heller et al. (1994), but also see Eger and Schlitter (2001).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Glauconycteris humeralis; Glauconycteris humeralis; Glauconycteris humeralis; Glauconycteris humeralis; Glauconycteris humeralis; Glauconycteris humeralis; humeralis; Glauconyctere tacheté; Gefleckte Schmetterlingsfledermaus; Glauconicteriomanchado; Allen's Spotted Bat; Spotted Butterfly Bat; Allen's Spotted Bat; Spotted Butterfly Bat; Spotted Butterfly Bat; G. humeralis
