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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L324	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Glauconycteris beatrix	Chalinolobus beatrix	Glauconycteris beatrix	Chalinolobus beatrix	Chalinolobus beatrix	Glauconycteris beatrix	Glauconycteris beatrix	Glauconycteris beatrix	Glauconycteris beatrix	Glauconycteris beatrix	Glauconycteris beatrix	Glauconycteris beatrix	Glauconycteris beatrix	Glauconycteris beatrix	Glauconycteris beatrix		[MSW2] Subgenus Glauconycteris. Includes humeralis; see Koopman (1971:7, 8).; [MSW3] Does not include humeralis; see Hill and Harrison (1987) and Heller et al. (1994), but also see Eger and Schlitter (2001).; [HMW] Glauconycteris beatrix Thomas, 1901 , “Benito River, French Congo [= Republic of the Congo ], fifteen miles [= 24 km ] from mouth.” Form humeralis has been previously treated under G. beatrix , but they are tentatively treated as separate species here. A. Hassanin and colleagues in 2018 retrieved G. beatrix as sister to G. curryae . Monotypic.; [batnames2022] Does not include humeralis ; see Hill and Harrison (1987) and Heller et al. (1994), but also see Eger and Schlitter (2001).; [IUCN] Synonym = Chalinolobus beatrix .; [batnames2023] Does not include humeralis ; see Hill and Harrison (1987) and Heller et al. (1994), but also see Eger and Schlitter (2001).; [batnames2025_1.7] Does not include humeralis; see Hill and Harrison (1987) and Heller et al. (1994), but also see Eger and Schlitter (2001).				humeralis		humeralis.	beatrix, humeralis							beatrix	Synonym = Chalinolobus beatrix .			beatrix	beatrix			beatrix O. Thomas, 1901		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Beatrix bat	Cameroun – Uganda	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Chalinolobus beatrix	Equatorial Guinea, Rio Muni, Benito River, 15 mi. (24 km) from mouth.	Thomas	1901	Ann. Mag. Nat. Hist., ser. 7, 8:256.	Distribution: Ranging in forested areas from Ivory Coast to northern An- gola and east to western Kenya.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Beatrix bat	Ivory Coast – Kenya	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1901	Ann. Mag. Nat. Hist., ser. 7, 8:256.	Subgenus Glauconycteris. Includes humeralis; see Koopman (1971:7, 8).	Ivory Coast, Kenya, Congo Republic, perhaps Guinea (Bissau).	Equatorial Guinea, Rio Muni, Benito River, 15 mi. (24 km) from mouth.		THOMAS	1901	Plagiopatagium and uropatagium dark in color, without a reticulated pattern. Fur dark brown, often with a white shoulder spot. Ear pinna short and rounded. Size relatively small (forearm length, 33-41 mm). Inner upper incisor strongly bicuspid with subequal cusps.	Distribution: Ranging in forested areas from Ivory Coast to northern Angola and east to western Kenya.	Two subspecies are here recognized:	C. b. beatrix (Ivory Coast to Angola), C. b. humeralis (Zaire to Kenya).	124	species	C. beatrix	THOMAS	1901	Glauconycteris	subgenus	Chalinolobus beatrix				Plagiopatagium and uropatagium dark in color, without a reticulated pattern. Fur dark brown, often with a white shoulder spot. Ear pinna short and rounded. Size relatively small (forearm length, 33-41 mm). Inner upper incisor strongly bicuspid with subequal cusps.	Two subspecies are here recognized:		7. C. beatrix (THOMAS 1901).	7	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Vespertilionini	Glauconycteris beatrix	Glauconycteris		beatrix	Thomas		1901		Ann. Mag. Nat. Hist., ser. 7	8		256		Beatrix Butterfly Bat	Equatorial Guinea, Rio Muni, Benito River, 15 mi. (24 km) from mouth.	Equatorial Guinea, Côte d’Ivoire, Ghana, Nigeria, Cameroon, Gabon, Angola. A report of this species from Guinea-Bissau is in error (J. Fahr, pers. comm.)	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt) as Chalinolobus beatrix.		Does not include humeralis; see Hill and Harrison (1987) and Heller et al. (1994), but also see Eger and Schlitter (2001).	4C3D87E8FFB06A0FFF8E9AC91957BD10	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	833	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFB06A0FFF8E9AC91957BD10.xml	Glauconycteris beatrix	Vespertilionidae	Glauconycteris	beatrix	Thomas	1901	Glauconyctére beatrice @fr | Kleine Schmetterlingsfledermaus @de | Glauconicteriopequefio @es | Beatrix's Bat @en	Glauconycteris beatrix Thomas, 1901 , “Benito River, French Congo [= Republic of the Congo ], fifteen miles [= 24 km ] from mouth.” Form humeralis has been previously treated under G. beatrix , but they are tentatively treated as separate species here. A. Hassanin and colleagues in 2018 retrieved G. beatrix as sister to G. curryae . Monotypic.	W & C Africa, from Ivory Coast E through Ghana and Nigeria to Cameroon and Equatorial Guinea (including Bioko I), with scattered records from S Gabon , SW Central African Republic , NW Angola , and EC DR Congo .	Head-body ¢.36-50 mm, tail 35-51 mm, ear 7-12 mm, hindfoot 6-8-7-5 mm, forearm 35-42 mm; weight 4-9 g. Pelage is dense, with 8-9 mm middorsal hairs. Dorsal and ventral fur is dark sepia-brown. White shoulder spotis present in holotype but apparently usually absent. Ears are very rounded, and separate, short for a vespertilionid and Glauconycteris . Inner margin has rounded lobe at base, and outer margin is connected with well-developed,fleshy lobe on lower lip near corner of mouth. Wings and uropatagium are dark brown and notreticulated. Tibia is long (18-21 mm) for Glauconycteris . Baculum is triangular, with basal part deeply lobed and distal part short and blunt. Head is high-domed, and muzzle is short, broad, and flattish. Skull is small, and profile of forehead is weakly concave compared with other Glauconycteris . I? is bicuspid, with secondary cusp equal in size or smaller than inner cusp. Lower incisors are tricuspid or with four cusps and not crowded. Diploid number is 2n = 22.	Generally lowland tropical moist forests but also forests outside rainforest zones. A few Beatrix’s Butterfly Bats were recorded by a waterhole in secondary forest and in cocoa plantations and swamp forests.	No information.	No information.	No information.	No information.	Classified as Least Concern on The IUCN Red List.	Happold, M. (2013av) | Hassanin et al. (2018) | Hill & Harrison (1987) | Koopman (1989, 1994) | Peterson & Smith (1973) | Porteret al. (2010)	https://zenodo.org/record/6398089/files/figure.png	152. Beatrix’s Butterfly Bat Glauconycteris beatrix French: Glauconyctére beatrice / German: Kleine Schmetterlingsfledermaus / Spanish: Glauconicterio pequefio Other common names: Beatrix's Bat Taxonomy. Glauconycteris beatrix Thomas, 1901 , “Benito River, French Congo [= Republic of the Congo ], fifteen miles [= 24 km ] from mouth.” Form humeralis has been previously treated under G. beatrix , but they are tentatively treated as separate species here. A. Hassanin and colleagues in 2018 retrieved G. beatrix as sister to G. curryae . Monotypic. Distribution. W & C Africa, from Ivory Coast E through Ghana and Nigeria to Cameroon and Equatorial Guinea (including Bioko I), with scattered records from S Gabon , SW Central African Republic , NW Angola , and EC DR Congo . Descriptive notes. Head-body ¢.36-50 mm, tail 35-51 mm, ear 7-12 mm, hindfoot 6-8-7-5 mm, forearm 35-42 mm; weight 4-9 g. Pelage is dense, with 8-9 mm middorsal hairs. Dorsal and ventral fur is dark sepia-brown. White shoulder spotis present in holotype but apparently usually absent. Ears are very rounded, and separate, short for a vespertilionid and Glauconycteris . Inner margin has rounded lobe at base, and outer margin is connected with well-developed,fleshy lobe on lower lip near corner of mouth. Wings and uropatagium are dark brown and notreticulated. Tibia is long (18-21 mm) for Glauconycteris . Baculum is triangular, with basal part deeply lobed and distal part short and blunt. Head is high-domed, and muzzle is short, broad, and flattish. Skull is small, and profile of forehead is weakly concave compared with other Glauconycteris . I? is bicuspid, with secondary cusp equal in size or smaller than inner cusp. Lower incisors are tricuspid or with four cusps and not crowded. Diploid number is 2n = 22. Habitat. Generally lowland tropical moist forests but also forests outside rainforest zones. A few Beatrix’s Butterfly Bats were recorded by a waterhole in secondary forest and in cocoa plantations and swamp forests. Food and Feeding. No information. Breeding. No information. Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCN Red List. Bibliography. Happold, M. (2013av), Hassanin et al. (2018), Hill & Harrison (1987), Koopman (1989, 1994), Peterson & Smith (1973), Porteret al. (2010).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Glauconycteris beatrix	Glauconycteris		beatrix	Thomas	1901	0	Ann. Mag. Nat. Hist.	ser. 7, 8: 256	Beatrix Butterfly Bat	None.	Equatorial Guinea, Rio Muni, Benito River, 15 mi. (24 km) from mouth.	Equatorial Guinea, Côte d’Ivoire, Ghana, Nigeria, Cameroon, Gabon, Angola. A report of this species from Guinea-Bissau is in error (J. Fahr, pers. comm.)	Not listed.	Least Concern	Does not include humeralis ; see Hill and Harrison (1987) and Heller et al. (1994), but also see Eger and Schlitter (2001).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Glauconycteris beatrix	23	Beatrix's Butterfly Bat	Beatrix's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	EPTESICINI	Glauconycteris	NA	beatrix	O. Thomas	1901	0	Glauconycteris_beatrix	Thomas, O. (1901). New Species of Macroscelides and Glauconycteris. Annals and Magazine of Natural History, ser. 7, 8, 256.	https://www.biodiversitylibrary.org/item/94923#page/284/mode/1up	BM 1898.5.4.19		"Benito River, French Congo [= Republic of the Congo], fifteen miles [= 24 km] from mouth."			beatrix O. Thomas, 1901	NA	NA	Côte d'Ivoire|Ghana|Nigeria|Cameroon|Equatorial Guinea|Gabon|Central African Republic|Angola|Republic of the Congo|Democratic Republic of the Congo	Africa	Afrotropic	LC	0	0	0	Glauconycteris_beatrix	0	sciname match	Glauconycteris_beatrix	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	44791	Glauconycteris beatrix	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Glauconycteris	beatrix	(Thomas, 1901)	Synonym = Chalinolobus beatrix .	20000000	Glauconycteris beatrix	Least Concern		2017	2016-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population (especially in Central Africa), and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	This species is generally associated with lowland tropical moist forest (Happold, 1987). There appears to be little information available on the roosting habits of this species, but presumably it can be found within hollow trees and dense vegetation. It is not known if the species can persist in areas of disturbed forest.	This species is threatened by general habitat loss, often resulting from logging activities and the conversion of land to agricultural use.	This appears to be a rare, or at least a rarely recorded, species.	Unknown	This species is distributed in West and Central Africa. It ranges from Côte d'Ivoire in the west, through Ghana and Nigeria to Cameroon and Equatorial Guinea (Rio Muni and Bioko), with scattered records from Gabon, Central African Republic, Angola, Congo, and the Democratic Republic of the Congo.		Terrestrial	The species has been recorded from a number of protected areas. Further studies are needed into the distribution, natural history and threats to this little known bat.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Glauconycteris		beatrix	Thomas	1901	0	Ann. Mag. Nat. Hist.	ser. 7, 8: 256	Beatrix Butterfly Bat	None.	Equatorial Guinea, Rio Muni, Benito River, 15 mi. (24 km) from mouth.	Equatorial Guinea, Côte d’Ivoire, Ghana, Nigeria, Cameroon, Gabon, Angola. A report of this species from Guinea-Bissau is in error (J. Fahr, pers. comm.)	Not listed.	Least Concern	Does not include humeralis ; see Hill and Harrison (1987) and Heller et al. (1994), but also see Eger and Schlitter (2001).	Glauconycteris beatrix	1005538	23	Beatrix's Butterfly Bat	Beatrix's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	EPTESICINI	Glauconycteris	NA	beatrix	O. Thomas	1901	0	Glauconycteris_beatrix	Thomas, O. (1901). New Species of Macroscelides and Glauconycteris. Annals and Magazine of Natural History, ser. 7, 8, 256.	https://www.biodiversitylibrary.org/item/94923#page/284/mode/1up	BM 1898.5.4.19		"Benito River, French Congo [= Republic of the Congo], fifteen miles [= 24 km] from mouth."			beatrix O. Thomas, 1901	NA	NA				Côte d'Ivoire|Ghana|Nigeria|Cameroon|Equatorial Guinea|Gabon|Central African Republic|Angola|Republic of the Congo|Democratic Republic of the Congo	Africa	Afrotropic	LC	0	0	0	Glauconycteris_beatrix	0	sciname match	Glauconycteris_beatrix	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Glauconycteris_beatrix	1005538	23	Beatrix's Butterfly Bat	Beatrix's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Nycticeiini	Glauconycteris	NA	beatrix	O. Thomas	0	Glauconycteris beatrix	Thomas, O. 1901-09-01. New species of _Macroscelides_ and _Glauconycteris_. Annals and Magazine of Natural History (7)8(45):255-257.	https://www.biodiversitylibrary.org/page/29980304	BMNH:Mamm:1898.5.4.19	holotype	https://data.nhm.ac.uk/object/718485b7-71bb-4d7c-9ba7-6f90a5e9ad79	"Benito River, French Congo [= Republic of the Congo], fifteen miles [= 24 km] from mouth."			NA	NA				Cote d'Ivoire|Ghana|Nigeria|Cameroon|Equatorial Guinea|Gabon|Central African Republic|Angola|Republic of the Congo|Democratic Republic of the Congo	Africa	Afrotropic	LC	0	0	0	Glauconycteris_beatrix	0	sciname match	Glauconycteris_beatrix	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Glauconycteris		beatrix	Thomas	1901	0	Ann. Mag. Nat. Hist.	ser. 7, 8: 256	Beatrix Butterfly Bat	None.	Equatorial Guinea, Rio Muni, Benito River, 15 mi. (24 km) from mouth.	Equatorial Guinea, Côte d’Ivoire, Ghana, Nigeria, Cameroon, Gabon, Angola. A report of this species from Guinea-Bissau is in error (J. Fahr, pers. comm.)	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/44791/22068514/' target='_blank'>Least Concern</a>	Does not include humeralis; see Hill and Harrison (1987) and Heller et al. (1994), but also see Eger and Schlitter (2001).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Glauconycteris beatrix; Glauconycteris beatrix; Glauconycteris beatrix; Glauconycteris beatrix; Glauconycteris beatrix; Glauconycteris beatrix; beatrix; Glauconyctére beatrice; Kleine Schmetterlingsfledermaus; Glauconicteriopequefio; Beatrix's Bat; Beatrix's Butterfly Bat; Beatrix's Bat; Beatrix Butterfly Bat; Beatrix Butterfly Bat; G. beatrix