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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L319	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Mimon crenulatum [synonym of]	Mimon crenulatum keenani	Mimon crenulatum keenani	Gardnerycteris keenani	Gardnerycteris keenani	Gardnerycteris keenani	N/A	Gardnerycteris keenani	Gardnerycteris keenani	Gardnerycteris keenani	Gardnerycteris keenani	Gardnerycteris keenani		[HMW] Mimon crenulatum keenani Handley, 1960 , “Fort Gulick, Panama Canal Zone,” Panama . Based on phylogenetic analysis, N. Hurtado and V. Pacheco in 2014 reassigned keenani to the new genus Gardnerycteris . Monotypic.; [batnames2022] Formerly included in crenulatum but distinct; see Hurtado and D'Elia (2018).; [MDD2022] split from G. crenulatum; moved from Mimon to Gardnerycteris; [batnames2023] Formerly included in crenulatum but distinct; see Hurtado and D'Elia (2018).; [MDD2023] split from G. crenulatum; moved from Mimon to Gardnerycteris; [MDD2025_2.0] split from G. crenulatum; moved from Mimon to Gardnerycteris; [batnames2025_1.7] Formerly included in crenulatum but distinct; see Hurtado and D'Elia (2018).; [MDD2025_2.2] split from G. crenulata; moved from Mimon to Gardnerycteris														keenani				keenani	keenani			keenani (Handley, 1960)						N/A																																								NA																											03A687BCFFA3FFA3168DFDCAF5F9FBCF	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	502	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFFA3FFA3168DFDCAF5F9FBCF.xml	Gardnerycteris keenani	Phyllostomidae	Gardnerycteris	keenani		1960	Gardneryctére de Keenan @fr | Keenan-Haarblattnase @de | Gardnericterio de Keenan @es	Mimon crenulatum keenani Handley, 1960 , “Fort Gulick, Panama Canal Zone,” Panama . Based on phylogenetic analysis, N. Hurtado and V. Pacheco in 2014 reassigned keenani to the new genus Gardnerycteris . Monotypic.	Central America from S Mexico (Lacandon region and S Yucatan Peninsula) and through Caribbean slope to Panama and in South American W Andes in Colombia , W Venezuela , W Ecuador , and extreme NW Peru .	Head-body 40-66 mm, tail 23-24 mm, hindfoot 10-11 mm, ear 23-25 mm,forearm 48-52 mm; weight 12-16 g. Keenan’s Hairy-nosed Bat differs from other species of Gardnerycteris by the following combination of characteristics: broad and yellowish cream dorsalstripe, large and prominent yellowish auricular patches, third metacarpal shorter than fifth, and uropatagium fringe absent. No other bat species has crenulated noseleaf with large hairs on tip and short ones on base and a broad yellowish stripe on back. Anterior border of nasal bones is sigmoidal at each side, rostrum has shallow median depression, root of zygomatic arch is flattened,sagittal crest is low, accessory median foramen is present, bullae are broad and high, basioccipital at cochlear level is broad, basisphenoid pits are deep, anterior crest of coronoid process is curved, coronoid process does not pass base of molar row, and median sulcus between hypocone and paracone is present in M! and M?.	Equatorial dry forest, Pacific tropical rainforests, tropical moist deciduous forest, and banana groves from sea level up to elevations of ¢. 600 m .	Keenan's Hairy-nosed Batis mainly insectivorous, eating Coleoptera ( Cerambycidae ), Hemiptera , Lepidoptera , Diptera , and small vertebrates (lizards).	Pregnant Keenan’s Hairy-nosed Bats are known from beginning of rainy season (April) in Costa Rica and from February in Campeche , Mexico . They have one young per year, and young are found during dry season (December) and early rainy season.	Keenan's Hairy-nosed Bat has been caught more often near the ground than at subcanopylevels. It probably gleans insects and small vertebrates over vegetation. It roosts in hollow tree trunks,rotting logs, and hollow tree stumps, preferring humid areas; in Venezuela ,it was caught near buildings.	Colonies of Keenan’s Hairy-nosed Bats are small, with less than ten individuals. They are frequently mist-netted in male— female pairs, suggesting that pairs forage together.	Not assessed on The IUCNRed List. Keenan’s Hairy-nosed Bat is considered endangered on the Mexican list of threatened species (as Mimon crenulatum ).	Humphrey et al. (1983) | Hurtado & D’Elia (2018) | Hurtado & Pacheco (2014) | Hurtado et al. (2014) | McCarthy (1987) | McCarthy et al. (1993) | Medellin (1983) | SEMARNAT (2010) | Timm etal. (1989)	https://zenodo.org/record/6458656/files/figure.png	27. Keenan's Hairy-nosed Bat Gardnerycteris keenani French: Gardneryctére de Keenan / German: Keenan-Haarblattnase / Spanish: Gardnericterio de Keenan Taxonomy. Mimon crenulatum keenani Handley, 1960 , “Fort Gulick, Panama Canal Zone,” Panama . Based on phylogenetic analysis, N. Hurtado and V. Pacheco in 2014 reassigned keenani to the new genus Gardnerycteris . Monotypic. Distribution. Central America from S Mexico (Lacandon region and S Yucatan Peninsula) and through Caribbean slope to Panama and in South American W Andes in Colombia , W Venezuela , W Ecuador , and extreme NW Peru . Descriptive notes. Head-body 40-66 mm, tail 23-24 mm, hindfoot 10-11 mm, ear 23-25 mm,forearm 48-52 mm; weight 12-16 g. Keenan’s Hairy-nosed Bat differs from other species of Gardnerycteris by the following combination of characteristics: broad and yellowish cream dorsalstripe, large and prominent yellowish auricular patches, third metacarpal shorter than fifth, and uropatagium fringe absent. No other bat species has crenulated noseleaf with large hairs on tip and short ones on base and a broad yellowish stripe on back. Anterior border of nasal bones is sigmoidal at each side, rostrum has shallow median depression, root of zygomatic arch is flattened,sagittal crest is low, accessory median foramen is present, bullae are broad and high, basioccipital at cochlear level is broad, basisphenoid pits are deep, anterior crest of coronoid process is curved, coronoid process does not pass base of molar row, and median sulcus between hypocone and paracone is present in M! and M?. Habitat. Equatorial dry forest, Pacific tropical rainforests, tropical moist deciduous forest, and banana groves from sea level up to elevations of ¢. 600 m . Food and Feeding. Keenan's Hairy-nosed Batis mainly insectivorous, eating Coleoptera ( Cerambycidae ), Hemiptera , Lepidoptera , Diptera , and small vertebrates (lizards). Breeding. Pregnant Keenan’s Hairy-nosed Bats are known from beginning of rainy season (April) in Costa Rica and from February in Campeche , Mexico . They have one young per year, and young are found during dry season (December) and early rainy season. Activity patterns. Keenan's Hairy-nosed Bat has been caught more often near the ground than at subcanopylevels. It probably gleans insects and small vertebrates over vegetation. It roosts in hollow tree trunks,rotting logs, and hollow tree stumps, preferring humid areas; in Venezuela ,it was caught near buildings. Movements, Home range and Social organization. Colonies of Keenan’s Hairy-nosed Bats are small, with less than ten individuals. They are frequently mist-netted in male— female pairs, suggesting that pairs forage together. Status and Conservation. Not assessed on The IUCNRed List. Keenan’s Hairy-nosed Bat is considered endangered on the Mexican list of threatened species (as Mimon crenulatum ). Bibliography. Humphrey et al. (1983), Hurtado & D’Elia (2018), Hurtado & Pacheco (2014), Hurtado et al. (2014), McCarthy (1987), McCarthy et al. (1993), Medellin (1983), SEMARNAT (2010), Timm etal. (1989).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Gardnerycteris keenani	Gardnerycteris		keenani	Handley	1960	1	Proc. U.S. Natl. Mus.	112(3442): 460	Keenan&apos;s Hairy-nosed Bat	None.	Panama, Panama Canal Zone, Fort Gulick	Mexico, Belize, Guatemala, Honduras, Nicaragua, Costa Rica, Pananma, W Ecuador, W Peru, W Venezuela	Not listed.	Least Concern under Mimon crenulatum 	Formerly included in crenulatum but distinct; see Hurtado and D'Elia (2018).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Gardnerycteris keenani	23	Keenan's Hairy-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	PHYLLOSTOMINAE	PHYLLOSTOMINI	Gardnerycteris	NA	keenani	Handley	1960	1	Mimon_crenulatum_keenani	Handley, C. O., Jr. (1960). Decriptions of New Bats from Panama. Proceedings of the United States National Museum, 112, 460.	https://repository.si.edu/bitstream/handle/10088/16682/USNMP-112_3442_1960.pdf?sequence=1&isAllowed=y	USNM 311951		"Fort Gulick, Panama Canal Zone," Panama.			keenani Handley, 1960	split from G. crenulatum; moved from Mimon to Gardnerycteris	Hurtado-Miranda, N. E., & Pacheco-Torres, V. (2014). Análisis filogenético del género Mimon Gray, 1847 (Mammalia, Chiroptera, Phyllostomidae) con la descripción de un nuevo género. Therya, 5(3), 751-791.|Hurtado, N., & D'Elía, G. (2018). Taxonomy of the genus Gardnerycteris (Chiroptera: Phyllostomidae). Acta Chiropterologica, 20(1), 99-115.	Mexico|Belize|Guatemala|Honduras|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Ecuador|Peru	North America|South America	Neotropic	NA	0	0	0	Gardnerycteris_keenani	0	unmatched	NA	1																																			Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Gardnerycteris		keenani	Handley	1960	1	Proc. U.S. Natl. Mus.	112(3442): 460	Keenan&apos;s Hairy-nosed Bat	None.	Panama, Panama Canal Zone, Fort Gulick	Mexico, Belize, Guatemala, Honduras, Nicaragua, Costa Rica, Pananma, W Ecuador, W Peru, W Venezuela	Not listed.	Least Concern under Mimon crenulatum 	Formerly included in crenulatum but distinct; see Hurtado and D'Elia (2018).	Gardnerycteris keenani	1004970	23	Keenan's Hairy-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	PHYLLOSTOMINAE	PHYLLOSTOMINI	Gardnerycteris	NA	keenani	Handley	1960	1	Mimon_crenulatum_keenani	Handley, C. O., Jr. (1960). Decriptions of New Bats from Panama. Proceedings of the United States National Museum, 112, 460.	https://repository.si.edu/bitstream/handle/10088/16682/USNMP-112_3442_1960.pdf?sequence=1&isAllowed=y	USNM 311951		"Fort Gulick, Panama Canal Zone," Panama.			keenani Handley, 1960	split from G. crenulatum; moved from Mimon to Gardnerycteris	Hurtado-Miranda, N. E., & Pacheco-Torres, V. (2014). Análisis filogenético del género Mimon Gray, 1847 (Mammalia, Chiroptera, Phyllostomidae) con la descripción de un nuevo género. Therya, 5(3), 751-791.|Hurtado, N., & D'Elía, G. (2018). Taxonomy of the genus Gardnerycteris (Chiroptera: Phyllostomidae). Acta Chiropterologica, 20(1), 99-115.				Mexico|Belize|Guatemala|Honduras|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Ecuador|Peru	North America|South America	Neotropic	NA	0	0	0	Gardnerycteris_keenani	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Gardnerycteris_keenani	1004970	23	Keenan's Hairy-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Phyllostominae	Phyllostomini	Gardnerycteris	NA	keenani	Handley	1	Mimon crenulatum keenani	Handley, C.O., Jr. 1960-10-06. Descriptions of new bats from Panama. Proceedings of the United States National Museum 112:459-479.	https://www.biodiversitylibrary.org/page/7717356	USNM:MAMM:311951	holotype	http://n2t.net/ark:/65665/3f25c6fce-ba7b-4a86-8285-be1da1b18136	"Fort Gulick, Panama Canal Zone," Panama.			split from G. crenulata; moved from Mimon to Gardnerycteris	Hurtado-Miranda, N. E., & Pacheco-Torres, V. (2014). Análisis filogenético del género Mimon Gray, 1847 (Mammalia, Chiroptera, Phyllostomidae) con la descripción de un nuevo género. Therya, 5(3), 751-791.|Hurtado, N., & D'Elía, G. (2018). Taxonomy of the genus Gardnerycteris (Chiroptera: Phyllostomidae). Acta Chiropterologica, 20(1), 99-115.				Mexico|Belize|Guatemala|Honduras|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Ecuador|Peru	North America|South America	Neotropic	NE	0	0	0	Gardnerycteris_keenani	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Gardnerycteris		keenani	Handley	1960	1	Proc. U.S. Natl. Mus.	112(3442): 460	Keenan&apos;s Hairy-nosed Bat	None.	Panama, Panama Canal Zone, Fort Gulick	Mexico, Belize, Guatemala, Honduras, Nicaragua, Costa Rica, Pananma, W Ecuador, W Peru, W Venezuela	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	Not Evaluated	Formerly included in crenulatum but distinct; see Hurtado and D'Elia (2018).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Gardnerycteris keenani; Gardnerycteris keenani; Gardnerycteris keenani; Gardnerycteris keenani; keenani; Gardneryctére de Keenan; Keenan-Haarblattnase; Gardnericterio de Keenan; Keenan's Hairy-nosed Bat; Keenan&apos;s Hairy-nosed Bat; G. keenani