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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L309	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Eumops bonariensis [synonym of]	Eumops glaucinus nanus	Eumops bonariensis nanus	Eumops nanus	Eumops nanus	Eumops nanus	Eumops nanus	Eumops nanus	Eumops nanus	Eumops nanus	Eumops nanus	Eumops nanus		[HMW] Promops nanus G. S. Miller, 1900 , “Bogava [= Bugaba District], Chiriqui [Province], Panama (alt. 250 m .).” Eumops nanus was previously considered a subspecies of E. bonariensis based on morphological and molecular differences. Monotypic.; [batnames2022] Distinct from bonariensis; see Eger (2007); Gregorin (2009).; [MDD2022] split from bonariensis; [IUCN] It was previously treated as a subspecies of ;E. bonariensis ;(see Simmons 2005, Eger 2008).; [batnames2023] Distinct from bonariensis; see Eger (2007); Gregorin (2009).; [MDD2023] split from bonariensis; [MDD2025_2.0] split from bonariensis; [batnames2025_1.7] Distinct from bonariensis; see Eger (2007); Gregorin (2009).; [MDD2025_2.2] split from bonariensis														nanus	It was previously treated as a subspecies of ;E. bonariensis ;(see Simmons 2005, Eger 2008).			nanus	nanus			nanus (G. S. Miller, 1900)						N/A																																								NA																											194287C9FFB1BA1EB1B1FF76B02DFDD4	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Molossidae_598.pdf.imf	hash://md5/e57bffb1ffbcba10b412f760b226ffce	632	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/19/42/87/194287C9FFB1BA1EB1B1FF76B02DFDD4.xml	Eumops nanus	Molossidae	Eumops	nanus		1900	Eumope nain @fr | Kleine Bulldogfledermaus @de | Eumops nano @es | Dwarf Bonneted Bat @en	Promops nanus G. S. Miller, 1900 , “Bogava [= Bugaba District], Chiriqui [Province], Panama (alt. 250 m .).” Eumops nanus was previously considered a subspecies of E. bonariensis based on morphological and molecular differences. Monotypic.	Patchily from S Mexico ( Veracruz and Oaxaca ) SE through Central America to extreme N Colombia , NW Venezuela , S Guyana , and Peru ; also on Cozumel 1.	Head—body 49-68 mm, tail 28-47 mm, ear 12-19 mm, hindfoot 6-11 mm, forearm 39-48 mm; weight 7-13 g. See general characteristics of the genus under the Black Bonneted Bat ( E. auripendulus ). The Northern Dwarf Bonneted Bat is the smallest species of Eumopsin overall size. Dorsal furis grayish brown to dark brown, with c. 5mm long hairs being whitish at bases. Ventral fur is slightly paler and frosted on tips. Lips are wrinkled but less than in Wagner’s Bonneted Bat ( E. glaucinus ). Ears extendjust beyond nose tip when laid forward. Wing attaches near end of tibia. Upper incisors converge to each other and are not in contact with canines.	[Lowland tropical humid forests, thorn forests, and secondary forests.	Stomach contents of Northern Dwarf Bonneted Bats from Mexico contained 55% moths, 27% beetles, and 15% bugs (by volume).	In Mexico , Northern Dwarf Bonneted Bats were pregnant from late March to late June, with parturition occurring in midto late June. Lactating females were reported from early June to late July, with lactation lasting up to two months. Reproductively inactive females occurred in March-April, June-July, and September-—November. Subadults were found in October. Males had enlarged and secreting throat glands in May-June.	Primary activity of Northern Dwarf Bonneted Bats occurs during first two hours after sunset, with secondary activity before sunrise. They roost in tree holes and under roof tiles of houses in colonies of ¢.20 individuals in Mexico . These bats fly near water bodies, including human-made ponds and swimming pools.	No information.	Classified as [Least Concern on The IUCN Red Lust.	Bowles et al. (1990) | Eger (2008) | Handley (1976) | Hutterer et al. (1995) | Miller (1900a) | Reid (1997) | Torres-Morales et al. (2014)	https://zenodo.org/record/6418335/files/figure.png	36. Northern Dwarf Bonneted Bat Eumops nanus French: Eumope nain / German: Kleine Bulldogfledermaus / Spanish: Eumops nano Other common names: Dwarf Bonneted Bat Taxonomy. Promops nanus G. S. Miller, 1900 , “Bogava [= Bugaba District], Chiriqui [Province], Panama (alt. 250 m .).” Eumops nanus was previously considered a subspecies of E. bonariensis based on morphological and molecular differences. Monotypic. Distribution. Patchily from S Mexico ( Veracruz and Oaxaca ) SE through Central America to extreme N Colombia , NW Venezuela , S Guyana , and Peru ; also on Cozumel 1. Descriptive notes. Head—body 49-68 mm, tail 28-47 mm, ear 12-19 mm, hindfoot 6-11 mm, forearm 39-48 mm; weight 7-13 g. See general characteristics of the genus under the Black Bonneted Bat ( E. auripendulus ). The Northern Dwarf Bonneted Bat is the smallest species of Eumopsin overall size. Dorsal furis grayish brown to dark brown, with c. 5mm long hairs being whitish at bases. Ventral fur is slightly paler and frosted on tips. Lips are wrinkled but less than in Wagner’s Bonneted Bat ( E. glaucinus ). Ears extendjust beyond nose tip when laid forward. Wing attaches near end of tibia. Upper incisors converge to each other and are not in contact with canines. Habitat. [Lowland tropical humid forests, thorn forests, and secondary forests. Food and Feeding. Stomach contents of Northern Dwarf Bonneted Bats from Mexico contained 55% moths, 27% beetles, and 15% bugs (by volume). Breeding. In Mexico , Northern Dwarf Bonneted Bats were pregnant from late March to late June, with parturition occurring in midto late June. Lactating females were reported from early June to late July, with lactation lasting up to two months. Reproductively inactive females occurred in March-April, June-July, and September-—November. Subadults were found in October. Males had enlarged and secreting throat glands in May-June. Activity patterns. Primary activity of Northern Dwarf Bonneted Bats occurs during first two hours after sunset, with secondary activity before sunrise. They roost in tree holes and under roof tiles of houses in colonies of ¢.20 individuals in Mexico . These bats fly near water bodies, including human-made ponds and swimming pools. Movements, Home range and Social organization. No information. Status and Conservation. Classified as [Least Concern on The IUCN Red Lust. Bibliography. Bowles et al. (1990), Eger (2008), Handley (1976), Hutterer et al. (1995), Miller (1900a), Reid (1997), Torres-Morales et al. (2014).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Molossidae	Eumops nanus	Eumops		nanus	Miller	1900	1	Ann. Mag. Nat. Hist.	ser. 7, 6: 470	Dwarf Bonneted Bat	None.	Panama, Chiriquí, Bogava.	Panama	Not listed.	Least Concern	Distinct from bonariensis; see Eger (2007); Gregorin (2009).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Eumops nanus	23	Northern Dwarf Bonneted Bat	Dwarf Bonneted Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	MOLOSSIDAE	MOLOSSINAE	NA	Eumops	NA	nanus	G. S. Miller	1900	1	Promops_nanus	Miller, G. S., Jr. (1900). A new Free-tailed bat from Central America. Annals and Magazine of Natural History, ser. 7, 6, 470.	https://www.biodiversitylibrary.org/item/53364#page/488/mode/1up	BM 1900.7.11.99		"Bogava [= Bugaba District], Chiriqui [Province], Panama (alt. 250 m.)."			nanus (G. S. Miller, 1900)	split from bonariensis	Gregorin, R. (2009). Phylogeny of Eumops Miller, 1906 (Chiroptera: Molossidae) using morphological data. Acta Chiropterologica, 11(2), 247-258.	Mexico|Honduras|Nicaragua|Panama|Colombia|Venezuela|Guyana|Peru	North America|South America	Nearctic|Neotropic	LC	0	0	0	Eumops_nanus	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Eumops nanus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	MOLOSSIDAE	Eumops	nanus	(Miller, 1900)	It was previously treated as a subspecies of ;E. bonariensis ;(see Simmons 2005, Eger 2008).	90000000	Eumops nanus	Least Concern		2017	2016-10-26 00:00:00 UTC	3.1	English	This species is listed as Least Concern in view of its wide distribution, which overlaps several protected areas, and although its populations has not been fully assessed it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	Its diet is composed of moths, beetles, and other insects; it utilizes natural and artificial bodies of water to hunt its preys (Bowles et al. 1990, Arita 1999, Reid 2009). In México, ;E. nanus ;has been recorded in tropical dry forest, tropical thorn forest, deciduous forest, and secondary forests, besides of urban and suburban areas but always below 40 m ;(Torres-Morales et al. 2014). In Yucatan. a birth peak occurs in late June, and lactation lasts 6-8 weeks (Reid 2009).	Threats to this species, if any, are unknown at the present time.	Uncommon or rare; found in dry forests and thorn scrub. Roosts in tree holes and roofs of houses. A long-term study in Yucatan, Mexico, found this species to be fairly common (Bowles et al. 1990). A group of about 20 occupied roof tiles of a house near Merica, with 1-3 bats under each individual tile (Bowles et al. 1990).	Stable	Eumops nanus ; occurs in southern Mexico, patchily recorded through Central America, ;northern ;Colombia and Venezuela, Guyana, and Peru (Eger 2008, Reid 2009). In lowlands only. It was previously treated as a subspecies of ;E. bonariensis ;(see Simmons 2005).		Terrestrial	Throughout its distribution, this species is protected under several types of ;protected and natural areas, including Central America and northern South America. In some parts of Mexico, it is considered rare (Arita 1999).	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Molossidae	Eumops		nanus	Miller	1900	1	Ann. Mag. Nat. Hist.	ser. 7, 6: 470	Dwarf Bonneted Bat	None.	Panama, Chiriquí, Bogava.	Panama	Not listed.	Least Concern	Distinct from bonariensis; see Eger (2007); Gregorin (2009).	Eumops nanus	1005185	23	Northern Dwarf Bonneted Bat	Dwarf Bonneted Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Molossidae	MOLOSSINAE	NA	Eumops	NA	nanus	G. S. Miller	1900	1	Promops_nanus	Miller, G. S., Jr. (1900). A new Free-tailed bat from Central America. Annals and Magazine of Natural History, ser. 7, 6, 470.	https://www.biodiversitylibrary.org/item/53364#page/488/mode/1up	BM 1900.7.11.99		"Bogava [= Bugaba District], Chiriqui [Province], Panama (alt. 250 m.)."			nanus (G. S. Miller, 1900)	split from bonariensis	Gregorin, R. (2009). Phylogeny of Eumops Miller, 1906 (Chiroptera: Molossidae) using morphological data. Acta Chiropterologica, 11(2), 247-258.				Mexico|Honduras|Nicaragua|Panama|Colombia|Venezuela|Guyana|Peru	North America|South America	Nearctic|Neotropic	LC	0	0	0	Eumops_nanus	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Eumops_nanus	1005185	23	Northern Dwarf Bonneted Bat	Dwarf Bonneted Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Molossidae	Molossinae	NA	Eumops	NA	nanus	G. S. Miller	1	Promops nanus	Miller, G.S., Jr. 1900-11-01. A new free-tailed bat from Central America. Annals and Magazine of Natural History (7)6(35):470-471.	https://www.biodiversitylibrary.org/page/19266262	BMNH:Mamm:1900.7.11.99	holotype	https://data.nhm.ac.uk/object/5a2a7aae-f45f-457c-bcc7-6ae2bcd47d09	"Bogava [= Bugaba District], Chiriqui [Province], Panama (alt. 250 m.)."			split from bonariensis	Gregorin, R. (2009). Phylogeny of Eumops Miller, 1906 (Chiroptera: Molossidae) using morphological data. Acta Chiropterologica, 11(2), 247-258.				Mexico|Honduras|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Guyana|Peru	North America|South America	Nearctic|Neotropic	LC	0	0	0	Eumops_nanus	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Molossidae	Eumops		nanus	Miller	1900	1	Ann. Mag. Nat. Hist.	ser. 7, 6: 470	Dwarf Bonneted Bat	None.	Panama, Chiriquí, Bogava.	Panama	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/87994060/87994063/' target='_blank'>Least Concern</a>	Distinct from bonariensis; see Eger (2007); Gregorin (2009).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Eumops nanus; Eumops nanus; Eumops nanus; Eumops nanus; Eumops nanus; nanus; Eumope nain; Kleine Bulldogfledermaus; Eumops nano; Dwarf Bonneted Bat; Northern Dwarf Bonneted Bat; Dwarf Bonneted Bat; Dwarf Bonneted Bat; E. nanus