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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L281	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Epomophorus labiatus	Epomophorus labiatus	Epomophorus labiatus	Epomophorus labiatus	Epomophorus labiatus	Epomophorus labiatus	Epomophorus labiatus	Epomophorus labiatus	Epomophorus labiatus	Epomophorus labiatus	Epomophorus labiatus	Epomophorus labiatus	Epomophorus labiatus	Epomophorus labiatus	Epomophorus labiatus		[MSW2] Includes anurus and minor; see Claessen and Vree (1991), but see also Bergmans (1988).; [MSW3] gambianus species group. Includes anurus; see Kock (1969a), Bergmans (1988, 1997), and Claessen and De Vree (1991). Koopman (1994) recognized two subspecies (labiatus and anurus), but this arrangement does not appear justified given the morphometric data presented by Claessen and De Vree (1991), who did not recognize subspecies. Apparently does not include minor contra Claessen and De Vree (1991), see discussion in Bergmans (1988, 1997). Middle Eastern forms reviewed by Horácek et al. (2000).; [HMW] Pteropus labiatus Temminck, 1837 , Sudan , “ Abyssinia .” Restricted by W. Bergmans in 1988 to Sennar, Sudan . Epomophorus labiatus is in the gambianus species group and includes E. anurus. Systematics of small-sized forms of Epomophorus remains unclear. Two concepts of E. labiatus exist: one of a widely varying species with pronounced geographical clinal trends, including large specimens of E. minor , and another one not inclusive of E. minor and with more restricted morphometric variation (only larger specimens of the labiatus form). Additional research is needed to improve systematic resolution. Monotypic.; [batnames2022]  gambianus species group. Includes anurus; see Kock (1969a), Bergmans (1988, 1997), and Claessen and De Vree (1991). Koopman (1994) recognizedtwo subspecies (labiatus and anurus), but this arrangement does not appear justified given the morphometric data presented byClaessen and De Vree (1991), who did not recognize subspecies. Apparently does not include minor contra Claessen and De Vree (1991),see discussion in Bergmans (1988, 1997). Middle Eastern forms reviewed by Horácek et al. (2000).; [IUCN] We follow Bergmans (1988) by treating Epomophorus minor as a separate species from E . labiatus .; [batnames2023]  gambianus species group. Includes anurus; see Kock (1969a), Bergmans (1988, 1997), and Claessen and De Vree (1991). Koopman (1994) recognizedtwo subspecies (labiatus and anurus), but this arrangement does not appear justified given the morphometric data presented byClaessen and De Vree (1991), who did not recognize subspecies. Apparently does not include minor contra Claessen and De Vree (1991),see discussion in Bergmans (1988, 1997). Middle Eastern forms reviewed by Horácek et al. (2000).; [batnames2025_1.7] gambianusspecies group. Includes anurus; see Kock (1969a), Bergmans (1988, 1997), and Claessen and De Vree (1991). Koopman (1994) recognizedtwo subspecies (labiatus and anurus), but this arrangement does not appear justified given the morphometric data presented byClaessen and De Vree (1991), who did not recognize subspecies. Apparently does not include minor contra Claessen and De Vree (1991),see discussion in Bergmans (1988, 1997). Middle Eastern forms reviewed by Horácek et al. (2000).				anurus	(anurus)	anurus, doriae, minor, schoensis.	labiatus		anurus, doriae, schoensis, schovanus			labiatus	labiatus - anurus, doriae, schoensis, schovanus	labiatus , schoensis, anurus, doriae	We follow Bergmans (1988) by treating Epomophorus minor as a separate species from E . labiatus .	labiatus	labiatus - anurus, doriae, schoensis, schovanus	labiatus, schoensis, anurus, doriae	labiatus, schoensis, anurus, schovanus, doriae	labiatus	labiatus - anurus, doriae, schoensis, schovanus	labiatus (Temminck, 1837)|schoensis (Rüppell, 1842)|anurus von Heuglin, 1864|schovanus (von Heuglin, 1877) [unjustified emendation]|doriae Matschie, 1899		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Little epauletted fruit bat	N Ethiopia, S Sudan, ? Congo Rep.; ref. 4.33	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Epomophorus labiatus	Sudan, Blue Nile Prov., Sennar.	Temminck	1837	Monogr. Mamm., 2:83.	Distribution: Chiefly northern tropical Africa from Senegal to Ethiopia and south to southern Congo Republic, northern Zaire, Burundi, and northern Tanzania.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Little epauletted fruit bat	Nigeria – Ethiopia – Malawi, Tanzania, ? Senegal	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Temminck	1837	Monogr. Mamm., 2:83.	Includes anurus and minor; see Claessen and Vree (1991), but see also Bergmans (1988).	Nigeria to Ethiopia and south to Congo Republic and Malawi. Senegal records are probably erroneous (see Bergmans, 1988).	Sudan, Blue Nile Prov., Sennar.		HEUGLIN	1864	Fourth palatal ridge midway between third and fifth. Size medium (forearm length, 62-81 mm; total length of skull, 40-49 mm).	Distribution: Chiefly northern tropical Africa from Senegal to Ethiopia and south to southern Congo Republic, northern Zaire, Burundi, and northern Tanzania.	Two subspecies (KOOPMAN 1975):	E. I. labiatus (Ethiopia and northeastern Sudan), E. !. anurus (remaining range).	31	species	E. labiatus	HEUGLIN	1864	Epomophorus	genus	Epomophorus labiatus				Fourth palatal ridge midway between third and fifth. Size medium (forearm length, 62-81 mm; total length of skull, 40-49 mm).	Two subspecies (KOOPMAN 1975):		7. E. labiatus HEUGLIN 1864 [gambianus group].	7	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Epomophorus labiatus	Epomophorus		labiatus	Temminck	y	1837		Monogr. Mamm.	2		83		Little Epauletted Fruit Bat	Sudan, Blue Nile Prov., Sennar.	Saudi Arabia; Nigeria to Ethiopia and Djibouti, south to Republic of Congo and Malawi. Senegal records are probably erroneous (see Bergmans, 1988).	IUCN/SSC Action Plan (1992) – Not Threatened. IUCN 2003 – Lower Risk (lc).	anurus Heuglin, 1864; doriae Matscheie, 1899; schoensis Rüppell, 1842; schovanus Heuglin, 1877.	gambianus species group. Includes anurus; see Kock (1969a), Bergmans (1988, 1997), and Claessen and De Vree (1991). Koopman (1994) recognized two subspecies (labiatus and anurus), but this arrangement does not appear justified given the morphometric data presented by Claessen and De Vree (1991), who did not recognize subspecies. Apparently does not include minor contra Claessen and De Vree (1991), see discussion in Bergmans (1988, 1997). Middle Eastern forms reviewed by Horácek et al. (2000).	03AD87FAFFECF60289AF32FFF975F959	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	98	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFECF60289AF32FFF975F959.xml	Epomophorus labiatus	Pteropodidae	Epomophorus	labiatus	Temminck	1837	Epomophore labiaire @fr | Athiopien-Epaulettenflughund @de | Epoméforo de Etiopia @es | Ethiopian Epauletted Fruit Bat @en	Pteropus labiatus Temminck, 1837 , Sudan , “ Abyssinia .” Restricted by W. Bergmans in 1988 to Sennar, Sudan . Epomophorus labiatus is in the gambianus species group and includes E. anurus. Systematics of small-sized forms of Epomophorus remains unclear. Two concepts of E. labiatus exist: one of a widely varying species with pronounced geographical clinal trends, including large specimens of E. minor , and another one not inclusive of E. minor and with more restricted morphometric variation (only larger specimens of the labiatus form). Additional research is needed to improve systematic resolution. Monotypic.	Arabian Peninsula (SW Saudi Arabia and Yemen ) and patchily distributed in Africa from E Sudan , Djibouti , and Ethiopia S to E DR Congo , E Zambia , Malawi , and NW Mozambique , also on Unguja and Mafia Is ( Zanzibar Archipelago) and isolated localities in NE Nigeria , Chad , and on the coast of the Republic of Congo .	Head-body 105-113 mm (males) and 90-98 mm (females), tail 0-5 mm , ear 17-21 mm , hindfoot 12-18 mm , forearm 66-80 mm (males) and 58- 76 mm (females); weight 48-64 g (males) and 34-51 g (females). Males frequently are darker and on average slightly larger than females, with broader muzzle and folded upper lip. Muzzle is rather long but relatively medium-short and broad for Epomophorus . Tongue is long, with small backward-pointing papillae near tip, very extendable, and extensively used for grooming. Eyes are large;irises are brown or chestnut-brown. Ears are relatively short, rounded, naked, and brown, with darker rims and anterior and posterior light ear tufts at bases. White epaulettes occur on adult males. Dorsum is light sandy brown on males, hairs are dark brown at bases, mid-dorsal hairs are ¢. 10 mm ; dorsum is light brown (fawn) on females, hairs are beige with pale brown tips; pelage of both sexes is soft and fluffy, extending along forearm dorsally and ventrally. Venter of males is pale brown on chest to pure white on abdomen; some males have dark brown shoulders and throat; chest of females is pale fawn and becomes darker on abdomen. Wings have claw on second digits; membranes emerge from sides of body, attach to second toes, and are light brown and sparsely and very thinly covered in hair. Skull is medium to short; rostrum is relatively short, broad, and in profile flat as is interorbital region; parietal region is rounded and deflected downward; sagittal crest is absent; nuchal crest is weak; and post-dental palate is moderately concave. There are six thick palatal ridges; fourth is midway between third and fifth, and fifth is partially or fully postdental; and ridges 2-4 are not divided. Chromosomal complement has 2n = 36 and FNa = 72 (females), with twelve pairs of metacentric and six pairs of submetacentric autosomes.	Sudan and Guinea Savanna (undifferentiated woodland, wetter miombo woodland, evergreen and semi-evergreen bushlands, Acacia [ Fabaceae ] and Commiphora [ Burseraceae ] bushland and thicket and Acacia wooded grasslands), Rainforest-Savanna Mosaic, Coastal Forest Mosaic, Zambezian Woodland and Afromontane-Afroalpine ( Ethiopia ) biotic zones from sea level up to elevations of ¢. 2200 m .	The Little Epauletted Fruit Bat eats indigenous and cultivated soft fruits; e.g. Ficus ( Moraceae ), Terminalia ( Combretaceae ), Mangifera ( Anacardiaceae ), Irvingia ( Irvingiaceae ), and Salvadora ( Salvadoraceae ). Fruit is packed into expandable cheeks and carried to a perch whereitis chewed. Nectar and pollen are consumed from flowers of Kigelia pinnata and K. aethiopica ( Bignoniaceae ), among other plants. Captive individuals did not drink water.	Litter size of the Little Epauletted Fruit Bat is one. In Uganda , reproductive cycle is continuous bimodal polyestry, with postpartum estrus, and seasonal polyestry to monoestry in drier localities. Gestation lasts 5-6 months. Females are in reproductive synchrony, and males are probably also in synchrony with females. Parturition and postpartum mating occur just before or at beginning of wet seasons (March to early April and late September to early October). In Malawi and Zambia where there is only one wet season (November to early April), there is no reproductive synchrony, and births occur in October-February.	During the day, Little Epauletted Fruit Bats hang (often alone) freely from branches of trees and shrubs (sometime under dense foliage), banana leaves, or exposed tree roots. Adult males probably perform calling displays, but due to the small size of larynx, calls probably have limited reach.	The Little Epauletted Fruit Bat generally roosts alone, but groups of up to 40 individuals have been observed. In Malawi , young were captured in March-May but no adults in March—June or September, suggesting it might be nomadic.	Classified as Least Concern on The IUCN Red List. The Little Epauletted Fruit Bat has a wide distribution and large population. It is probably not declining fast enough to be assigned to a higher category. It faces no major threats, but overharvesting for subsistence food (and medicine in certain parts of Cameroon ) might be a problem.	Benda, Al-Jumaily et al. (2011) | Bergmans (1988, 1997) | Bobo & Ntumwel (2010) | Claessen & De Vree (1990) | Duli¢ & Mutere (1973) | Gaucher (1992) | Happold, M. (2013e) | Koopman (1975) | Monadjem, Taylor et al. (2010) | Taylor (2016b)		65. Little Epauletted Fruit Bat Epomophorus labiatus French: Epomophore labiaire / German: Athiopien-Epaulettenflughund / Spanish: Epoméforo de Etiopia Other common names: Ethiopian Epauletted Fruit Bat Taxonomy. Pteropus labiatus Temminck, 1837 , Sudan , “ Abyssinia .” Restricted by W. Bergmans in 1988 to Sennar, Sudan . Epomophorus labiatus is in the gambianus species group and includes E. anurus. Systematics of small-sized forms of Epomophorus remains unclear. Two concepts of E. labiatus exist: one of a widely varying species with pronounced geographical clinal trends, including large specimens of E. minor , and another one not inclusive of E. minor and with more restricted morphometric variation (only larger specimens of the labiatus form). Additional research is needed to improve systematic resolution. Monotypic. Distribution. Arabian Peninsula (SW Saudi Arabia and Yemen ) and patchily distributed in Africa from E Sudan , Djibouti , and Ethiopia S to E DR Congo , E Zambia , Malawi , and NW Mozambique , also on Unguja and Mafia Is ( Zanzibar Archipelago) and isolated localities in NE Nigeria , Chad , and on the coast of the Republic of Congo . Descriptive notes. Head-body 105-113 mm (males) and 90-98 mm (females), tail 0-5 mm , ear 17-21 mm , hindfoot 12-18 mm , forearm 66-80 mm (males) and 58- 76 mm (females); weight 48-64 g (males) and 34-51 g (females). Males frequently are darker and on average slightly larger than females, with broader muzzle and folded upper lip. Muzzle is rather long but relatively medium-short and broad for Epomophorus . Tongue is long, with small backward-pointing papillae near tip, very extendable, and extensively used for grooming. Eyes are large;irises are brown or chestnut-brown. Ears are relatively short, rounded, naked, and brown, with darker rims and anterior and posterior light ear tufts at bases. White epaulettes occur on adult males. Dorsum is light sandy brown on males, hairs are dark brown at bases, mid-dorsal hairs are ¢. 10 mm ; dorsum is light brown (fawn) on females, hairs are beige with pale brown tips; pelage of both sexes is soft and fluffy, extending along forearm dorsally and ventrally. Venter of males is pale brown on chest to pure white on abdomen; some males have dark brown shoulders and throat; chest of females is pale fawn and becomes darker on abdomen. Wings have claw on second digits; membranes emerge from sides of body, attach to second toes, and are light brown and sparsely and very thinly covered in hair. Skull is medium to short; rostrum is relatively short, broad, and in profile flat as is interorbital region; parietal region is rounded and deflected downward; sagittal crest is absent; nuchal crest is weak; and post-dental palate is moderately concave. There are six thick palatal ridges; fourth is midway between third and fifth, and fifth is partially or fully postdental; and ridges 2-4 are not divided. Chromosomal complement has 2n = 36 and FNa = 72 (females), with twelve pairs of metacentric and six pairs of submetacentric autosomes. Habitat. Sudan and Guinea Savanna (undifferentiated woodland, wetter miombo woodland, evergreen and semi-evergreen bushlands, Acacia [ Fabaceae ] and Commiphora [ Burseraceae ] bushland and thicket and Acacia wooded grasslands), Rainforest-Savanna Mosaic, Coastal Forest Mosaic, Zambezian Woodland and Afromontane-Afroalpine ( Ethiopia ) biotic zones from sea level up to elevations of ¢. 2200 m . Food and Feeding. The Little Epauletted Fruit Bat eats indigenous and cultivated soft fruits; e.g. Ficus ( Moraceae ), Terminalia ( Combretaceae ), Mangifera ( Anacardiaceae ), Irvingia ( Irvingiaceae ), and Salvadora ( Salvadoraceae ). Fruit is packed into expandable cheeks and carried to a perch whereitis chewed. Nectar and pollen are consumed from flowers of Kigelia pinnata and K. aethiopica ( Bignoniaceae ), among other plants. Captive individuals did not drink water. Breeding. Litter size of the Little Epauletted Fruit Bat is one. In Uganda , reproductive cycle is continuous bimodal polyestry, with postpartum estrus, and seasonal polyestry to monoestry in drier localities. Gestation lasts 5-6 months. Females are in reproductive synchrony, and males are probably also in synchrony with females. Parturition and postpartum mating occur just before or at beginning of wet seasons (March to early April and late September to early October). In Malawi and Zambia where there is only one wet season (November to early April), there is no reproductive synchrony, and births occur in October-February. Activity patterns. During the day, Little Epauletted Fruit Bats hang (often alone) freely from branches of trees and shrubs (sometime under dense foliage), banana leaves, or exposed tree roots. Adult males probably perform calling displays, but due to the small size of larynx, calls probably have limited reach. Movements, Home range and Social organization. The Little Epauletted Fruit Bat generally roosts alone, but groups of up to 40 individuals have been observed. In Malawi , young were captured in March-May but no adults in March—June or September, suggesting it might be nomadic. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Little Epauletted Fruit Bat has a wide distribution and large population. It is probably not declining fast enough to be assigned to a higher category. It faces no major threats, but overharvesting for subsistence food (and medicine in certain parts of Cameroon ) might be a problem. Bibliography. Benda, Al-Jumaily et al. (2011), Bergmans (1988, 1997), Bobo & Ntumwel (2010), Claessen & De Vree (1990), Duli¢ & Mutere (1973), Gaucher (1992), Happold, M. (2013e), Koopman (1975), Monadjem, Taylor et al. (2010), Taylor (2016b).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Epomophorus labiatus	Epomophorus		labiatus	Temminck	1837	1	Monogr. Mamm.	0.141	Little Epauletted Fruit Bat	 anurus Heuglin, 1864; doriae Matscheie, 1899; schoensis R&uuml;ppell, 1842; schovanus Heuglin, 1877.	Sudan, Blue Nile Prov., Sennar.	Saudi Arabia; Nigeria to Ethiopia and Djibouti, south to Republic of Congo, Malawi, and Mozambique (see Monadjem et al. 2010). Senegal records are probably erroneous (see Bergmans, 1988).	Not listed.	Least Concern	 gambianus species group. Includes anurus; see Kock (1969a), Bergmans (1988, 1997), and Claessen and De Vree (1991). Koopman (1994) recognizedtwo subspecies (labiatus and anurus), but this arrangement does not appear justified given the morphometric data presented byClaessen and De Vree (1991), who did not recognize subspecies. Apparently does not include minor contra Claessen and De Vree (1991),see discussion in Bergmans (1988, 1997). Middle Eastern forms reviewed by Horácek et al. (2000).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Epomophorus labiatus	23	Little Epauletted Fruit Bat	Ethiopian Epauletted Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	ROUSETTINAE	EPOMOPHORINI	Epomophorus	NA	labiatus	Temminck	1837	1	Pteropus_labiatus	Temminck, C. J. (1837). Monographies de mammalogie, ou Description de quelques genres de mammifères, dont les espèces ont été observées dans les différens musées de l'Europe. Vol. 2. Leiden, 83.	https://www.google.com/books/edition/Monographies_de_mammalogie/yHQQAQAAMAAJ?hl=en&gbpv=1&dq=temminck+monographies+de+mammalogie&printsec=frontcover	RMNH 27342 [lectotype]		Sudan, "Abyssinia." Restricted by W. Bergmans in 1988 to Sennar, Sudan.			labiatus (Temminck, 1837)|schoensis (Rüppell, 1842)|anurus Heuglin, 1864|doriae Matschie, 1899	NA	NA	Saudi Arabia|Yemen|Sudan|Djibouti|Eritrea|Ethiopia|South Sudan|Nigeria|Chad|Republic of the Congo|Democratic Republic of the Congo|Uganda|Kenya|Rwanda|Burundi|Tanzania|Malawi|Mozambique	Africa|Asia	Afrotropic	LC	0	0	0	Epomophorus_labiatus	0	sciname match	Epomophorus_labiatus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Epomophorus labiatus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Epomophorus	labiatus	(Temminck, 1837)	We follow Bergmans (1988) by treating Epomophorus minor as a separate species from E . labiatus .	20000000	Epomophorus labiatus	Least Concern		2016	2015-12-29 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	This species is regularly recorded from moist forest-grassland mosaics (including rainforest and Afromontane forest), and mesic woodland and savanna (including Sudanean and Miombo woodlands) habitats. In East African coastal areas it has been recorded from mangroves. It has been found resting together in small groups of about a dozen animals.	There appear to be no major threats to this species as a whole. Some populations may be threatened by overharvesting for subsistence food.	This is a common species.	Stable	This species is largely distributed in Central Africa and East Africa and closely associated with highlands of the Rift Valley, occurring at higher altitudes than E. minor . It appears to range from northern Nigeria into Chad, southern and central South Sudan, Ethiopia and Eritrea, and from here south into eastern Democratic Republic of the Congo, Uganda, Rwanda and Burundi, Kenya and northwestern Tanzania, being recorded as far south as Malawi and soouthern Tanzania. There is one isolated record from coastal Congo. Based on collectors’ notes on labels of accurately identified specimens (Bergmans 1988), apart from the isolated coastal locality of Point Noire, Congo, it occurs at elevations from 610 m to 2,200 asl at Gondar, Ethiopia.	Is hunted for the bushmeat trade.	Terrestrial	This species has been recorded from many protected areas. No direct conservation measures are currently needed for the species as a whole. Additional research is needed into the impact of possible overharvesting of animals on some populations.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Epomophorus		labiatus	Temminck	1837	1	Monogr. Mamm.	0.140972	Little Epauletted Fruit Bat	 anurus Heuglin, 1864; doriae Matscheie, 1899; schoensis R&uuml;ppell, 1842; schovanus Heuglin, 1877.	Sudan, Blue Nile Prov., Sennar.	Saudi Arabia; Nigeria to Ethiopia and Djibouti, south to Republic of Congo, Malawi, and Mozambique (see Monadjem et al. 2010). Senegal records are probably erroneous (see Bergmans, 1988).	Not listed.	Least Concern	 gambianus species group. Includes anurus; see Kock (1969a), Bergmans (1988, 1997), and Claessen and De Vree (1991). Koopman (1994) recognizedtwo subspecies (labiatus and anurus), but this arrangement does not appear justified given the morphometric data presented byClaessen and De Vree (1991), who did not recognize subspecies. Apparently does not include minor contra Claessen and De Vree (1991),see discussion in Bergmans (1988, 1997). Middle Eastern forms reviewed by Horácek et al. (2000).	Epomophorus labiatus	1004523	23	Little Epauletted Fruit Bat	Ethiopian Epauletted Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	ROUSETTINAE	EPOMOPHORINI	Epomophorus	NA	labiatus	Temminck	1837	1	Pteropus_labiatus	Temminck, C. J. (1837). Monographies de mammalogie, ou Description de quelques genres de mammifères, dont les espèces ont été observées dans les différens musées de l'Europe. Vol. 2. Leiden, 83.	https://www.google.com/books/edition/Monographies_de_mammalogie/yHQQAQAAMAAJ?hl=en&gbpv=1&dq=temminck+monographies+de+mammalogie&printsec=frontcover	RMNH 27342 [lectotype]		Sudan, "Abyssinia." Restricted by W. Bergmans in 1988 to Sennar, Sudan.			labiatus (Temminck, 1837)|schoensis (Rüppell, 1842)|anurus Heuglin, 1864|doriae Matschie, 1899	NA	NA				Saudi Arabia|Yemen|Sudan|Djibouti|Eritrea|Ethiopia|South Sudan|Nigeria|Chad|Republic of the Congo|Democratic Republic of the Congo|Uganda|Kenya|Rwanda|Burundi|Tanzania|Malawi|Mozambique	Africa|Asia	Afrotropic	LC	0	0	0	Epomophorus_labiatus	0	sciname match	Epomophorus_labiatus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Epomophorus_labiatus	1004523	23	Little Epauletted Fruit Bat	Ethiopian Epauletted Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Epomophorinae	Epomophorini	Epomophorus	NA	labiatus	Temminck	1	Pteropus labiatus	Temminck, C.J. 1837. Livraison 2. Pp. 49–140 in Temminck, C.J. 1835-1841. Monographies de Mammalogie. Tome second. C. C. van der Hoek, Leiden, 392 pp.	https://archive.org/details/monographiedema00temmgoog	RMNH.MAM.27342	lectotype	https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.27342.a | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.27342.b	Sudan, "Abyssinia." Restricted by W. Bergmans in 1988 to Sennar, Sudan.			NA	NA				Saudi Arabia|Yemen|Sudan|Djibouti|Eritrea|Ethiopia|South Sudan|Nigeria|Chad|Republic of the Congo|Democratic Republic of the Congo|Uganda|Kenya|Rwanda|Burundi|Tanzania|Malawi|Mozambique	Africa|Asia	Afrotropic	LC	0	0	0	Epomophorus_labiatus	0	sciname match	Epomophorus_labiatus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Epomophorus		labiatus	Temminck	1837	1	Monogr. Mamm.	0.140972	Little Epauletted Fruit Bat	anurus Heuglin, 1864; doriae Matscheie, 1899; schoensis R&uuml;ppell, 1842; schovanus Heuglin, 1877.	Sudan, Blue Nile Prov., Sennar.	Saudi Arabia; Nigeria to Ethiopia and Djibouti, south to Republic of Congo, Malawi, and Mozambique (see Monadjem et al. 2010). Senegal records are probably erroneous (see Bergmans, 1988).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/84457881/22122505/' target='_blank'>Least Concern</a>	gambianusspecies group. Includes anurus; see Kock (1969a), Bergmans (1988, 1997), and Claessen and De Vree (1991). Koopman (1994) recognizedtwo subspecies (labiatus and anurus), but this arrangement does not appear justified given the morphometric data presented byClaessen and De Vree (1991), who did not recognize subspecies. Apparently does not include minor contra Claessen and De Vree (1991),see discussion in Bergmans (1988, 1997). Middle Eastern forms reviewed by Horácek et al. (2000).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Epomophorus labiatus; Epomophorus labiatus; Epomophorus labiatus; Epomophorus labiatus; Epomophorus labiatus; Epomophorus labiatus; anurus; doriae; schoensis; schovanus; anurus; doriae; schoensis; schovanus; labiatus; schoensis; anurus; doriae; Epomophore labiaire; Athiopien-Epaulettenflughund; Epoméforo de Etiopia; Ethiopian Epauletted Fruit Bat; Little Epauletted Fruit Bat; Ethiopian Epauletted Fruit Bat; Little Epauletted Fruit Bat; Little Epauletted Fruit Bat; E. labiatus