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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L29	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Pipistrellus coromandra	Pipistrellus coromandra	Pipistrellus coromandra	Pipistrellus coromandra	Pipistrellus coromandra	Pipistrellus coromandra	Pipistrellus coromandra	Pipistrellus coromandra	Pipistrellus coromandra	Pipistrellus coromandra	Pipistrellus coromandra	Pipistrellus coromandra	Pipistrellus coromandra	Pipistrellus coromandra	Alionoctula coromandra		[MSW2] Subgenus Pipistrellus. Does not include aladdin; see Corbet (1978c:53). See comment under P. pipistrellus.; [MSW3] Subgenus Pipistrellus. Does not include aladdin; see Corbet (1978c). Does not include portensis and tramatus; see Corbet and Hill (1992). See comment under pipistrellus. Reviewed by Bates and Harrison (1997).; [HMW] Scotophilus coromandra J. E. Gray, 1838 , Pondicherry , Coromandel Coast, India . Pipistrellus coromandra is externally identical to P. tenuis (being only larger in size), but the two species are considered distinct based on mtDNA. Populations in the north-western part of the distribution ( Afghanistan and Pakistan ) are grayer and distinguishable from the rest of the population and might be a separate subspecies (for which unavailable name afghanus has been applied), but additionalstudies are needed to resolve this. Monotypic.; [batnames2022] Subgenus Pipistrellus . Does not include aladdin ; see Corbet (1978 c ). Does not include portensis and tramatus ; see Corbet and Hill (1992). See comment under pipistrellus . Reviewed by Bates and Harrison (1997).; [IUCN] This taxon was traditionally placed to the â€œcoromandra â€ subgroup of the â€œpipistrellus â€ species group (e.g.: Corbet and Hill, 1992, Koopman, 1994). According to available molecular genetic data (Roehrs et al. , 2010, Benda et al. , 2016, orig.), P. coromandra is a part of genetic cluster of Oriental pipistrelles. This cluster is highly divergent from all the West Palearctic pipistrelles and may be referred to as â€œjavanicus â€ or â€œcoromandra â€ species group (both species were described by Gray on the same page of the same publication). Some earlier authors (e.g. Ellerman and Morrison-Scott 1951) included aladdin Thomas, 1905 to this species, though now it is treated as a partial synonym of P. pipistrellus . Gaisler (1970) proposed the nomen afghanus to represent forms from Pakistan and Afghanistan (Corbet and Hill 1992, Bates and Harrison 1997), however which is not accepted here vide Benda and Gaisler (2015). As with most Pipistrellus , the problem of identification coupled with taxonomic uncertainties means that the abundance and the distribution of the species cannot be easily defined. Previous records referable to this and other Pipistrellus species have often been confused (G. Csorba, pers. comm.). Nonetheless, according to DNA barcoding data, all available P. coromandra from different parts of Indochina and southern China form at least monophyletic cluster (S. Kruskop, pers. comm.).; [batnames2023] Subgenus Pipistrellus . Does not include aladdin ; see Corbet (1978 c ). Does not include portensis and tramatus ; see Corbet and Hill (1992). See comment under pipistrellus . Reviewed by Bates and Harrison (1997).; [batnames2025_1.7] Subgenus Pipistrellus. Does not include aladdin; see Corbet (1978c). Does not include portensis and tramatus; see Corbet and Hill (1992). See comment under pipistrellus. Reviewed by Bates and Harrison (1997).; [MDD2025_2.2] moved from Pipistrellus to Alionoctula						afghanus, blythii, coromandelicus, micropus, parvipes, portensis, tramatus.	coromandra, tramatus, portensis		afghanus, blythii, coromandelianus, coromandelicus, micropus, nicobaricus, parvipes			coromandra 	coromandra - afghanus, blythii, coromandelianus, coromandelicus, micropus, nicobaricus, parvipes	coromandra, coromandelicus, parvipes, blythii, nicobaricus, coromandelianus, micropus, afghanus	This taxon was traditionally placed to the â€œcoromandra â€ subgroup of the â€œpipistrellus â€ species group (e.g.: Corbet and Hill, 1992, Koopman, 1994). According to available molecular genetic data (Roehrs et al. , 2010, Benda et al. , 2016, orig.), P. coromandra is a part of genetic cluster of Oriental pipistrelles. This cluster is highly divergent from all the West Palearctic pipistrelles and may be referred to as â€œjavanicus â€ or â€œcoromandra â€ species group (both species were described by Gray on the same page of the same publication). Some earlier authors (e.g. Ellerman and Morrison-Scott 1951) included aladdin Thomas, 1905 to this species, though now it is treated as a partial synonym of P. pipistrellus . Gaisler (1970) proposed the nomen afghanus to represent forms from Pakistan and Afghanistan (Corbet and Hill 1992, Bates and Harrison 1997), however which is not accepted here vide Benda and Gaisler (2015). As with most Pipistrellus , the problem of identification coupled with taxonomic uncertainties means that the abundance and the distribution of the species cannot be easily defined. Previous records referable to this and other Pipistrellus species have often been confused (G. Csorba, pers. comm.). Nonetheless, according to DNA barcoding data, all available P. coromandra from different parts of Indochina and southern China form at least monophyletic cluster (S. Kruskop, pers. comm.).	coromandra 	coromandra - afghanus, blythii, coromandelianus, coromandelicus, micropus, nicobaricus, parvipes	coromandra, coromandelicus, parvipes, blythii, nicobaricus, coromandelianus, micropus, afghanus	coromandel, coromandelicus, coromandelicus, coromandra, sykesii, coromandelicus, sykesi, coromandelianus, parvipes, blythi, nicobaricus, blythii, sykesii, micropus, coromandrus, blythii, afghanus	coromandra 	coromandra - afghanus, blythii, coromandelianus, coromandelicus, micropus, nicobaricus, parvipes	coromandel (F. Cuvier, 1832) [not intended as a scientific name]|coromandelicus (G. Cuvier, 1833) [nomen nudum]|coromandelica (Lesson, 1836)|coromandra (J. E. Gray, 1838)|sykesii (J. E. Gray, 1843) [nomen nudum]|coromandelica (E. Blyth, 1851) [unjustified emendation | preoccupied]|sykesi (E. Blyth, 1851) [incorrect subsequent spelling]|coromandelianus (E. Blyth, 1852) [incorrect subsequent spelling]|parvipes (E. Blyth, 1853)|blythi (J. A. Wagner, 1855)|nicobaricus (Fitzinger, 1861) [nomen nudum]|blythii (Fitzinger, 1870) [incorrect subsequent spelling]|sykesii (Fitzinger, 1870)|micropus (W. C. H. Peters, 1872)|coromandra (O. Thomas, 1928) [unjustified emendation]|afghanus (Gaisler, 1970) [conditional]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Indian pipistrelle	E Afghanistan – S China, Vietnam, Sri Lanka	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Pipistrellus coromandra	India, Coromandel Coast, Pondicherry.	Gray	1838	Mag. Zool. Bot., 2:498.	Distribution: Ranging from Afghanistan to southern China and south to Ceylon and Thailand.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Indian pipistrelle	E Afghanistan – S China, Sri Lanka, Nicobar Is	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Gray	1838	Mag. Zool. Bot., 2:498.	Subgenus Pipistrellus. Does not include aladdin; see Corbet (1978c:53). See comment under P. pipistrellus.	Afghanistan, Pakistan, India (including Nicobar Isis), Sri Lanka, Nepal, Bhutan, Burma, S China, Thailand, Vietnam.	India, Coromandel Coast, Pondicherry.		GRAY	1838	Size relatively small (forearm length, 29-32 mm). Inner upper incisor bicuspid or unicuspid. Anterior upper premolar more or less displaced medially. Rostrum fairly broad. Forehead slightly concave to almost flat.	Distribution: Ranging from Afghanistan to southern China and south to Ceylon and Thailand.	Three subspecies are recognized:	P. c. coromandra (Afghanistan to Ceylon), P. c. tramatus (southern mainland China to Thailand), P. c. portensis (Hainan island).	112	species	P. coromandra	GRAY	1838	Pipistrellus	subgenus	Pipistrellus coromandra				Size relatively small (forearm length, 29-32 mm). Inner upper incisor bicuspid or unicuspid. Anterior upper premolar more or less displaced medially. Rostrum fairly broad. Forehead slightly concave to almost flat.	Three subspecies are recognized:		10. P. coromandra (GRAY 1838) [pipistrellus group],	10	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Pipistrellini	Pipistrellus coromandra	Pipistrellus	Pipistrellus	coromandra	Gray	y	1838		Mag. Zool. Bot.	2		498		Indian Pipistrelle	India, Coromandel Coast, Pondicherry.	Afghanistan, Bangladesh, India (including Nicobar Isls), Sri Lanka, Pakistan, Nepal, Bhutan, Burma, Cambodia, Thailand, S China.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	afghanus Gaisler, 1970; blythii Wagner, 1855; coromandelianus Blyth, 1863; coromandelicus Blyth, 1851; micropus Peters, 1872; nicobaricus Fitzinger, 1861; parvipes Blyth, 1853.	Subgenus Pipistrellus. Does not include aladdin; see Corbet (1978c). Does not include portensis and tramatus; see Corbet and Hill (1992). See comment under pipistrellus. Reviewed by Bates and Harrison (1997).	4C3D87E8FFEE6A52FA97928D1A44B831	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	779	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFEE6A52FA97928D1A44B831.xml	Pipistrellus coromandra	Vespertilionidae	Pipistrellus	coromandra		1838	Pipistrelle de Coromandel @fr | German @en | ndien-Zwergfledermaus @en | Pipistrela de Coromandel @es | Coromandel Pipistrelle @en | Little @en | ndian Bat @en	Scotophilus coromandra J. E. Gray, 1838 , Pondicherry , Coromandel Coast, India . Pipistrellus coromandra is externally identical to P. tenuis (being only larger in size), but the two species are considered distinct based on mtDNA. Populations in the north-western part of the distribution ( Afghanistan and Pakistan ) are grayer and distinguishable from the rest of the population and might be a separate subspecies (for which unavailable name afghanus has been applied), but additionalstudies are needed to resolve this. Monotypic.	NE Afghanistan ( Nangarhar Province ), N Pakistan ( Khyber Pakhtunkhwa and Punjab provinces), India , Sri Lanka , Andaman and NicobarIs, Nepal , Bhutan , Bangladesh , S China (S Tibet [= Xizang ]), Myanmar , Thailand , Laos , Vietnam , and Cambodia .	Head-body 34-49 mm, tail 22-39 mm, ear 7-14 mm, hindfoot 3-8 mm, forearm 25-35 mm; weight 3-4-8-3 g. The Indian Pipistrelle is extremely similar to the Least Pipistrelle ( P. tenuis ), but the Indian Pipistrelle averages larger for all external and cranial measurements (it is morphologically impossible to distinguish between a small Indian Pipistrelle and a large Least Pipistrelle). Dorsal pelage ranges from chestnut to medium to dark brown (being grayer brown in Afghanistan and Pakistan ); ventral pelage is paler, with beige-brown or cinnamon-brown tips and darker bases to hairs. Ears, face, and membranes are medium to dark brown and nearly naked. Ears are narrow and broadly rounded; tragus is more or less the same thickness throughout and narrow, with broadly rounded tip. Uropatagium has few hairs next to body and tail, and extends from calcar to nearly tail tip (only extreme tip is free). Penis is short (less than 8 mm ). Baculum (3-8 mm long, one specimen) has straight or slightly sinuous shaft, bifurcated tip, and ventrally deflected basal lobes. Skull is larger than that of the Least Pipistrelle but smaller than that of the Javan Pipistrelle (PF. javanicus ); zygomatic arches are thin and lack processes; I? is bicuspid, and I’ is unicuspid and higher than height of second cusp of I?; C! usually has secondary cusp; P* is subequal in crown area to I? and is usually (not always) intruded from tooth row; and lower molars are nyctalodont. Chromosomal complement has 2n = 30 and FN = 56 ( India ).	Primary and secondary montane and lowland forests, some disturbed forests, and highly disturbed agricultural and urban areas at elevations of 100-2000 m.	Indian Pipistrelles forage ¢.8-12 m aboveground or over water; along forest edges, creeks, and ravines; over cornfields; and in clearings. They are slow and erratic fliers and forage around cluttered vegetation in India . Stomach contents of some individuals in Bihar , northern India , contained small ants and mouthparts offlies.	Female Indian Pipistrelles seem to give birth to 1-2 young (rarely one) at a time and at least three times a year, with records in April, July-August, and October— November in Bihar . Females with two suckling young were captured in April, July, and August in Bihar . Pregnant females were captured in July in the same region, and a volantjuvenile with milk in its gut was captured in May.	Indian Pipistrelles emerge a few minutes before to 16 minutes after sunset and return 5-10 minutes before sunrise. In India , they were recorded feeding for 90 minutes before returning to roosts for short rests of 20-30 minutes before leaving roosts again to feed. They roost in dense vegetation, crevices in walls of mud houses,ceilings, and roofs, and bamboo thatch roofs. One individual in Myanmar was captured outside a cave. Search call is high-intensity FM call and was recorded with maximum energy at ¢.40-42 kHz in Vietnam . In southern India , calls were recorded with maximum frequencies of 115-7-127-6 kHz (average of 122-8 kHz), minimum frequencies of 35-9-43-1 kHz (38-8 kHz), frequencies of maximum energy of 47-3— 56-5 kHz (50-2 kHz), and durations of 1-2-2-1 milliseconds (1-6 milliseconds).	Indian Pipistrelles roost alone or in groups of 2-25 individuals in India . They occasionally roost with Least Pipistrelles but remain isolated from each other in the same roost.	Classified as Least Concern on The IUCNRed List. The Indian Pipistrelle is widespread and common throughoutits distribution and is commonly found in disturbed and urban habitats.	Abramov et al. (2010) | Bates & Harrison (1997) | Bates et al. (2005) | Benda & Gaisler (2015) | Cor bet & Hill (1992) | Csorba, Bates, Furey, Bumrungsri et al. (2008a) | Francis (2008) | Hill & Harrison (1987) | Kruskop (2013a) | Myers, P et al. (2000) | Raghuram et al. (2014) | Sinha (1984, 1986) | Smith & Xie Yan (2008) | Sreepada et al. (1996) | Srinivasulu et al. (2017) | Struebig et al. (2005) | Thapa, Subedi et al. (2012)	https://zenodo.org/record/6397848/files/figure.png	37. Indian Pipistrelle Pipistrellus coromandra French: Pipistrelle de Coromandel / German: Indien-Zwergfledermaus / Spanish: Pipistrela de Coromandel Other common names: Coromandel Pipistrelle , Little Indian Bat Taxonomy. Scotophilus coromandra J. E. Gray, 1838 , Pondicherry , Coromandel Coast, India . Pipistrellus coromandra is externally identical to P. tenuis (being only larger in size), but the two species are considered distinct based on mtDNA. Populations in the north-western part of the distribution ( Afghanistan and Pakistan ) are grayer and distinguishable from the rest of the population and might be a separate subspecies (for which unavailable name afghanus has been applied), but additionalstudies are needed to resolve this. Monotypic. Distribution. NE Afghanistan ( Nangarhar Province ), N Pakistan ( Khyber Pakhtunkhwa and Punjab provinces), India , Sri Lanka , Andaman and NicobarIs, Nepal , Bhutan , Bangladesh , S China (S Tibet [= Xizang ]), Myanmar , Thailand , Laos , Vietnam , and Cambodia . Descriptive notes. Head-body 34-49 mm, tail 22-39 mm, ear 7-14 mm, hindfoot 3-8 mm, forearm 25-35 mm; weight 3-4-8-3 g. The Indian Pipistrelle is extremely similar to the Least Pipistrelle ( P. tenuis ), but the Indian Pipistrelle averages larger for all external and cranial measurements (it is morphologically impossible to distinguish between a small Indian Pipistrelle and a large Least Pipistrelle). Dorsal pelage ranges from chestnut to medium to dark brown (being grayer brown in Afghanistan and Pakistan ); ventral pelage is paler, with beige-brown or cinnamon-brown tips and darker bases to hairs. Ears, face, and membranes are medium to dark brown and nearly naked. Ears are narrow and broadly rounded; tragus is more or less the same thickness throughout and narrow, with broadly rounded tip. Uropatagium has few hairs next to body and tail, and extends from calcar to nearly tail tip (only extreme tip is free). Penis is short (less than 8 mm ). Baculum (3-8 mm long, one specimen) has straight or slightly sinuous shaft, bifurcated tip, and ventrally deflected basal lobes. Skull is larger than that of the Least Pipistrelle but smaller than that of the Javan Pipistrelle (PF. javanicus ); zygomatic arches are thin and lack processes; I? is bicuspid, and I’ is unicuspid and higher than height of second cusp of I?; C! usually has secondary cusp; P* is subequal in crown area to I? and is usually (not always) intruded from tooth row; and lower molars are nyctalodont. Chromosomal complement has 2n = 30 and FN = 56 ( India ). Habitat. Primary and secondary montane and lowland forests, some disturbed forests, and highly disturbed agricultural and urban areas at elevations of 100-2000 m. Food and Feeding. Indian Pipistrelles forage ¢.8-12 m aboveground or over water; along forest edges, creeks, and ravines; over cornfields; and in clearings. They are slow and erratic fliers and forage around cluttered vegetation in India . Stomach contents of some individuals in Bihar , northern India , contained small ants and mouthparts offlies. Breeding. Female Indian Pipistrelles seem to give birth to 1-2 young (rarely one) at a time and at least three times a year, with records in April, July-August, and October— November in Bihar . Females with two suckling young were captured in April, July, and August in Bihar . Pregnant females were captured in July in the same region, and a volantjuvenile with milk in its gut was captured in May. Activity patterns. Indian Pipistrelles emerge a few minutes before to 16 minutes after sunset and return 5-10 minutes before sunrise. In India , they were recorded feeding for 90 minutes before returning to roosts for short rests of 20-30 minutes before leaving roosts again to feed. They roost in dense vegetation, crevices in walls of mud houses,ceilings, and roofs, and bamboo thatch roofs. One individual in Myanmar was captured outside a cave. Search call is high-intensity FM call and was recorded with maximum energy at ¢.40-42 kHz in Vietnam . In southern India , calls were recorded with maximum frequencies of 115-7-127-6 kHz (average of 122-8 kHz), minimum frequencies of 35-9-43-1 kHz (38-8 kHz), frequencies of maximum energy of 47-3— 56-5 kHz (50-2 kHz), and durations of 1-2-2-1 milliseconds (1-6 milliseconds). Movements, Home range and Social organization. Indian Pipistrelles roost alone or in groups of 2-25 individuals in India . They occasionally roost with Least Pipistrelles but remain isolated from each other in the same roost. Status and Conservation. Classified as Least Concern on The IUCNRed List. The Indian Pipistrelle is widespread and common throughoutits distribution and is commonly found in disturbed and urban habitats. Bibliography. Abramov et al. (2010), Bates & Harrison (1997), Bates et al. (2005), Benda & Gaisler (2015), Cor bet & Hill (1992), Csorba, Bates, Furey, Bumrungsri et al. (2008a), Francis (2008), Hill & Harrison (1987), Kruskop (2013a), Myers, P et al. (2000), Raghuram et al. (2014), Sinha (1984, 1986), Smith & Xie Yan (2008), Sreepada et al. (1996), Srinivasulu et al. (2017), Struebig et al. (2005), Thapa, Subedi et al. (2012).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Pipistrellus coromandra	Pipistrellus	Pipistrellus	coromandra	Gray	1838	1	Mag. Zool. Bot.	0.4292	Indian Pipistrelle	 afghanus Gaisler, 1970; blythii Wagner, 1855; coromandelianus Blyth, 1863; coromandelicus Blyth, 1851; micropus Peters, 1872; nicobaricus Fitzinger, 1861; parvipes Blyth, 1853.	India, Coromandel Coast, Pondicherry.	Afghanistan, Bangladesh, India (including Nicobar Isls), Sri Lanka, Pakistan, Nepal, Bhutan, Burma, Cambodia, Thailand, S China.	Not listed.	Least Concern	Subgenus Pipistrellus . Does not include aladdin ; see Corbet (1978 c ). Does not include portensis and tramatus ; see Corbet and Hill (1992). See comment under pipistrellus . Reviewed by Bates and Harrison (1997).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Pipistrellus coromandra	23	Indian Pipistrelle	Coromandel Pipistrelle|Little Indian Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PIPISTRELLINI	Pipistrellus	NA	coromandra	J. E. Gray	1838	1						Pondicherry, Coromandel Coast, India.			coromandra (J. E. Gray, 1838)|coromandelicus (Blyth, 1851)|parvipes (Blyth, 1853)|blythii (J. A. Wagner, 1855)|nicobaricus (Fitzinger, 1861)|coromandelianus (Blyth, 1863)|micropus (W. Peters, 1872)|afghanus Gaisler, 1970	NA	NA	Afghanistan|Pakistan|India|Sri Lanka|Andaman Islands|Nicobar Islands|Nepal|Bhutan|Bangladesh|China|Myanmar|Thailand|Laos|Vietnam|Cambodia	Asia	Indomalaya|Palearctic	LC	0	0	0	Pipistrellus_coromandra	0	sciname match	Pipistrellus_coromandra	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	17335	Pipistrellus coromandra	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Pipistrellus	coromandra	(Gray, 1838)	This taxon was traditionally placed to the â€œcoromandra â€ subgroup of the â€œpipistrellus â€ species group (e.g.: Corbet and Hill, 1992, Koopman, 1994). According to available molecular genetic data (Roehrs et al. , 2010, Benda et al. , 2016, orig.), P. coromandra is a part of genetic cluster of Oriental pipistrelles. This cluster is highly divergent from all the West Palearctic pipistrelles and may be referred to as â€œjavanicus â€ or â€œcoromandra â€ species group (both species were described by Gray on the same page of the same publication). Some earlier authors (e.g. Ellerman and Morrison-Scott 1951) included aladdin Thomas, 1905 to this species, though now it is treated as a partial synonym of P. pipistrellus . Gaisler (1970) proposed the nomen afghanus to represent forms from Pakistan and Afghanistan (Corbet and Hill 1992, Bates and Harrison 1997), however which is not accepted here vide Benda and Gaisler (2015). As with most Pipistrellus , the problem of identification coupled with taxonomic uncertainties means that the abundance and the distribution of the species cannot be easily defined. Previous records referable to this and other Pipistrellus species have often been confused (G. Csorba, pers. comm.). Nonetheless, according to DNA barcoding data, all available P. coromandra from different parts of Indochina and southern China form at least monophyletic cluster (S. Kruskop, pers. comm.).	20000000	Pipistrellus coromandra	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, it occurs in a number of protected areas, has a tolerance of a degree of habitat modification, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	This species is found in varied habitat types from forested regions, agricultural landscapes to urban areas. It roosts in trees, crevices and cracks in walls and ceilings of houses, tiles of huts, old buildings, temples, under bark and in holes of large trees, signboards, tree hollows in small groups of few individuals. In Vietnam this bat prefers forested areas with either pristine mountainous or secondary forest formations. It is an early flyer, emerging soon after sunset, when it still relatively light. Flight similar to other pipistrelles: moderately fast and maneuverable, somewhat fluttering, animals were usually seen foraging mainly along the linear landscape elements: forest edges and streams, at about 8-15 m above the ground. There is not reliable information about its ration, but probably it consists of small moths and other small flying insects. In Lao Cai province of Vietnam during spring breeding season adult males were observed mainly in forest formations at 1,800-2,000 m asl, while females and young animals â€“ in disturbed and rural plant formations at about 1,300 m asl (Kruskop 2013). There are up to three breeding seasons (probably vary along the distribution range) and one to two young ones are born (Bates and Harrison 1997).	There are no major threats to this species as a whole. Local populations may be affected by continuing deforestation; it especially concerns populations inhabiting isolated mountain massifs.	In South Asia, it is a widely distributed and common species and the population seems to be doing well in its range (Molur et al. 2002). It is fairly common in Vietnam, even in cities and similar urban habitats, it is one of the most common bats on e.g. Da Lat Plateau and in Hoang Lien Son mountains (Abramov et al. 2009, Kruskop and Shchinov 2010, Tu et al. 2016).	Unknown	This widely distributed species is found throughout most of South Asia, parts of southern China and much of mainland Southeast Asia. In South Asia this species is presently known from Afghanistan (Konar, Laghman and Nangarhar Province) (Benda and Gaisler 2015), Bangladesh (no exact location), Bhutan (no exact location), India (Andaman and Nicobar Islands, Andhra Pradesh, Arunachal Pradesh, Assam, Bihar, Goa, Gujarat, Jammu and Kashmir, Jharkhand, Karnataka, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Nagaland, Orissa, Sikkim, Tamil Nadu, Tripura, Uttarakhand, Uttar Pradesh and West Bengal), Nepal (Central), Pakistan (North West Frontier Province and Punjab) and Sri Lanka (Central, North Central, North Western Northern, Southern and Uva provinces) (Das 2003, Khan 2001, Korad et al. 2007, Molur et al. 2002, Simmons 2005, Srinivasulu and Srinivasulu 2005, Vanitharani 2006, Srinivasulu and Srinivasulu 2012). In South Asia, it has been recorded from 100 to 2,769 m asl (Molur et al. 2002). According to distribution scheme provided by Corbett and Hill (1992), Afghan and Pakistan population (P.c. afghanus ) is probably isolated from the Indian part of distribution range. In China it has been recorded from Xizang (Smith and Xie 2008). In Southeast Asia, it is present in Myanmar, Thailand, Lao PDR, Vietnam, Cambodia and Peninsular Malaysia. In Vietnam this bat occurs from southern provinces to most northern provinces, it has been recorded from ca. 900 to 2,000 m asl (Kruskop, 2013, and Kruskop, S.V. pers. comm.).	There are no evidences that this species was used in commercial trade, in local cuisine or in local medicine. Like other insectivorous bats, P. coromandra plays certain role as natural regulator of some forestry and harvest pests.	Terrestrial	In South Asia, although there are no direct conservation measures in place, the species has been recorded from several protected areas including Kanha National Park in Madhya Pradesh and Nagarjunasagar-Srisailam Tiger Reserve in Andhra Pradesh (C. Srinivasulu pers. comm. 6 March 2008). In Southeast Asia it has been recorded from some protected areas, in particular in Vietnam it occurs in e.g. Bi Doup â€“ Nui Ba, Chu Yang Sin and Hoang Lien Son national parks.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Pipistrellus	Pipistrellus	coromandra	Gray	1838	1	Mag. Zool. Bot.	0.429167	Indian Pipistrelle	 afghanus Gaisler, 1970; blythii Wagner, 1855; coromandelianus Blyth, 1863; coromandelicus Blyth, 1851; micropus Peters, 1872; nicobaricus Fitzinger, 1861; parvipes Blyth, 1853.	India, Coromandel Coast, Pondicherry.	Afghanistan, Bangladesh, India (including Nicobar Isls), Sri Lanka, Pakistan, Nepal, Bhutan, Burma, Cambodia, Thailand, S China.	Not listed.	Least Concern	Subgenus Pipistrellus . Does not include aladdin ; see Corbet (1978 c ). Does not include portensis and tramatus ; see Corbet and Hill (1992). See comment under pipistrellus . Reviewed by Bates and Harrison (1997).	Pipistrellus coromandra	1005615	23	Indian Pipistrelle	Coromandel Pipistrelle|Little Indian Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PIPISTRELLINI	Pipistrellus	NA	coromandra	J. E. Gray	1838	1						Pondicherry, Coromandel Coast, India.			coromandra (J. E. Gray, 1838)|coromandelicus (Blyth, 1851)|parvipes (Blyth, 1853)|blythii (J. A. Wagner, 1855)|nicobaricus (Fitzinger, 1861)|coromandelianus (Blyth, 1863)|micropus (W. Peters, 1872)|afghanus Gaisler, 1970	NA	NA				Afghanistan|Pakistan|India|Sri Lanka|Andaman Islands|Nicobar Islands|Nepal|Bhutan|Bangladesh|China|Myanmar|Thailand|Laos|Vietnam|Cambodia	Asia	Indomalaya|Palearctic	LC	0	0	0	Pipistrellus_coromandra	0	sciname match	Pipistrellus_coromandra	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Alionoctula_coromandra	1005615	23	Indian Pipistrelle	Coromandel Pipistrelle|Little Indian Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Pipistrellini	Alionoctula	NA	coromandra	J. E. Gray	1	Scotophilus Coromandra	Gray, J.E. 1838-02-01. A revision of the genera of bats (Vespertilionidae), and the description of some new genera and species. Magazine of Zoology and Botany 2(12):483-505.	https://www.biodiversitylibrary.org/page/40026401				Pondicherry, Coromandel Coast, India.			moved from Pipistrellus to Alionoctula	Zhukova, S. S., Yuzefovich, A. P., Lebedev, V. S., & Kruskop, S. V. (2025). Reassessment of the Taxonomic Borders Within Pipistrellus (Chiroptera, Vespertilionidae, Pipistrellini). Diversity, 17(5), 317.				Afghanistan|Pakistan|India|Sri Lanka|Andaman and Nicobar Islands|Nicobar Islands|Nepal|Bhutan|Bangladesh|China|Myanmar|Thailand|Laos|Vietnam|Cambodia	Asia	Indomalaya|Palearctic	LC	0	0	0	Pipistrellus_coromandra	0	sciname match	Pipistrellus_coromandra	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Pipistrellus	Pipistrellus	coromandra	Gray	1838	1	Mag. Zool. Bot.	0.429167	Indian Pipistrelle	afghanus Gaisler, 1970; blythii Wagner, 1855; coromandelianus Blyth, 1863; coromandelicus Blyth, 1851; micropus Peters, 1872; nicobaricus Fitzinger, 1861; parvipes Blyth, 1853.	India, Coromandel Coast, Pondicherry.	Afghanistan, Bangladesh, India (including Nicobar Isls), Sri Lanka, Pakistan, Nepal, Bhutan, Burma, Cambodia, Thailand, S China.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/17335/22130140/' target='_blank'>Least Concern</a>	Subgenus Pipistrellus. Does not include aladdin; see Corbet (1978c). Does not include portensis and tramatus; see Corbet and Hill (1992). See comment under pipistrellus. Reviewed by Bates and Harrison (1997).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Pipistrellus coromandra; Pipistrellus coromandra; Pipistrellus coromandra; Pipistrellus coromandra; Pipistrellus coromandra; Pipistrellus coromandra; afghanus; blythii; coromandelianus; coromandelicus; micropus; nicobaricus; parvipes; afghanus; blythii; coromandelianus; coromandelicus; micropus; nicobaricus; parvipes; coromandra; coromandelicus; parvipes; blythii; nicobaricus; coromandelianus; micropus; afghanus; Pipistrelle de Coromandel; German; ndien-Zwergfledermaus; Pipistrela de Coromandel; Coromandel Pipistrelle; Little; ndian Bat; Indian Pipistrelle; Coromandel Pipistrelle; Little Indian Bat; Indian Pipistrelle; Indian Pipistrelle; P. coromandra
