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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L280 application/vnd.openxmlformats-officedocument.spreadsheetml.sheet Micropteropus intermedius Micropteropus intermedius Micropteropus intermedius Micropteropus intermedius Micropteropus intermedius Micropteropus intermedius Epomophorus intermedius Epomophorus intermedius Epomophorus intermedius Micropteropus intermedius Epomophorus intermedius Epomophorus intermedius Epomophorus intermedius Epomophorus intermedius Epomophorus intermedius [HMW] Micropteropus intermedius Hayman, 1963 , Dundo, Lunda, Angola . Epomophorus intermedius is only known from one adult male and four adult females. Monotypic.; [batnames2022] Previously considered a species of Micropteropus , but moved to Epomophorus by Almeida et al. (2020).; [MDD2022] moved from Micropteropus to Epomophorus; [batnames2023] Previously considered a species of Micropteropus , but moved to Epomophorus by Almeida et al. (2020).; [MDD2023] moved from Micropteropus to Epomophorus; [MDD2025_2.0] moved from Micropteropus to Epomophorus; [batnames2025_1.7] Previously considered a species of Micropteropus, but moved to Epomophorus by Almeida et al. (2020).; [MDD2025_2.2] moved from Micropteropus to Epomophorus intermedius intermedius intermedius intermedius (Hayman, 1963) Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp. Hayman's epauletted fruit NE Angola, S Zaire bat Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp. Micropteropus intermedius Angola, Lunda, Dundo. Hayman 1963 Publ. Cult. Comp. Diamantes Angola, 66:100. Distribution: Known by a few specimens from southwestern Zaire and northern Angola. Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5 Hayman's epauletted fruit NE Angola, SE Zaire bat Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp. Hayman 1963 Publ. Cult. Comp. Diamantes Angola, 66:100. N Angola, SE Zaire. Angola, Lunda, Dundo. HAYMAN 1963 Five palatal ridges, second to fifth divided medially, converg ing posteriorly in clearly separate pairs. Size me dium (forearm length, 58-64 mm; total length of skull, 32-34 mm). Distribution: Known by a few specimens from southwestern Zaire and northern Angola. No subspecies. 32 species M. intermedius HAYMAN 1963 Micropteropus genus Micropteropus intermedius Five palatal ridges, second to fifth divided medially, converg ing posteriorly in clearly separate pairs. Size me dium (forearm length, 58-64 mm; total length of skull, 32-34 mm). No subspecies. 2. M. intermedius HAYMAN 1963. 2 NA Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu). CHIROPTERA Pteropodidae Micropteropus intermedius Micropteropus intermedius Hayman 1963 Publ. Cult. Comp. Diamantes Angola 66 100 Hayman's Lesser Epauletted Fruit Bat Angola, Lunda, Dundo. N Angola, SE Dem. Rep. Congo. IUCN/SSC Action Plan (1992) – Rare. IUCN 2003 – Data Deficient. 03AD87FAFFEFF6018C6735CDF9D4F7E3 Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions 978-84-16728-19-0 hbmw_9_Pteropodidae_16.pdf.imf hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b 101 zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFEFF6018C6735CDF9D4F7E3.xml Epomophorus intermedius Pteropodidae Epomophorus intermedius 1963 Epomophore de Hayman @fr | Hayman-Epaulettenflughund @de | Epoméforo de Hayman @es | Hayman’'s Dwarf Epauletted Fruit Bat @en | Hayman’s Lesser Epauletted Fruit Bat @en Micropteropus intermedius Hayman, 1963 , Dundo, Lunda, Angola . Epomophorus intermedius is only known from one adult male and four adult females. Monotypic. N Angola and SE & S DR Congo . Tail 3 mm (single male) and 5 mm (single female), ear 15 mm (single male) and 14- 3-17 mm (females), hindfoot 16 mm (single male) and 17- 18 mm (females), forearm 58 mm (single male) and 57-67 mm (females); weight 35-40 g . Head is round, and muzzle is rather short and blunt. Eyes are very large; ears are rounded and naked, with brown pigmentation and anterior and posterior light ear patches. Adult males have white epaulettes; adult females have curl of hairs on shoulders of same color and length as dorsal hairs. Dorsum is pale fawn to medium brown, paler on shoulders; hairs are medium sepia-brown at bases; and pelage is soft and moderately long. Venter is pale brown, darker on flanks. Wings have claw on second digits, and membranes are light to dull brown and attach to second toes; uropatagium is narrow; and calcars are present. Skull is short; rostrum is of medium relative length; braincase is more globose than in other species of Epomophorus , zygomatic arches are weak; and post-dental palate is almost flat, with prominent rim. There are six thick palatal ridges, one hastate and five deeply divided by wide median groove and two are post-dental; ridges 2-6 are divided by medial gap; and ridges 2-3 are underdeveloped, sometimes fused. Southern Rainforest-Savanna Mosaic biotic zone in open secondary grasslands and possibly woodland savannas of Zambezian Woodland biotic zone at elevations of ¢. 600 m . Hayman'’s Epauletted Fruit Bat is frugivorous and nectarivorous. One pregnant Hayman’s Epauletted Fruit Bat was recorded in June-July. Hayman'’s Epauletted Fruit Bats are nocturnal. One Hayman'’s Epauletted Fruit Bat was collected from a cave entrance. Classified as Data Deficient on The [UCN Red List (as Micropteropus intermedius ). Habitat loss due to deforestation might be a threat to Hayman’s Epauletted Fruit Bat, but additional research is needed to clarify its population and conservation status. It occurs in Mangroves National Park in DR Congo . Bergmans (1989) | Happold, M. (2013) | Mickleburgh, Hutson, Bergmans & Fahr (2008) | Monadjem, Taylor et al. (2010) 72. Hayman’s Epauletted Fruit Bat Epomophorus intermedius French: Epomophore de Hayman / German: Hayman-Epaulettenflughund / Spanish: Epoméforo de Hayman Other common names: Hayman’'s Dwarf Epauletted Fruit Bat , Hayman’s Lesser Epauletted Fruit Bat Taxonomy. Micropteropus intermedius Hayman, 1963 , Dundo, Lunda, Angola . Epomophorus intermedius is only known from one adult male and four adult females. Monotypic. Distribution. N Angola and SE & S DR Congo . Descriptive notes. Tail 3 mm (single male) and 5 mm (single female), ear 15 mm (single male) and 14- 3-17 mm (females), hindfoot 16 mm (single male) and 17- 18 mm (females), forearm 58 mm (single male) and 57-67 mm (females); weight 35-40 g . Head is round, and muzzle is rather short and blunt. Eyes are very large; ears are rounded and naked, with brown pigmentation and anterior and posterior light ear patches. Adult males have white epaulettes; adult females have curl of hairs on shoulders of same color and length as dorsal hairs. Dorsum is pale fawn to medium brown, paler on shoulders; hairs are medium sepia-brown at bases; and pelage is soft and moderately long. Venter is pale brown, darker on flanks. Wings have claw on second digits, and membranes are light to dull brown and attach to second toes; uropatagium is narrow; and calcars are present. Skull is short; rostrum is of medium relative length; braincase is more globose than in other species of Epomophorus , zygomatic arches are weak; and post-dental palate is almost flat, with prominent rim. There are six thick palatal ridges, one hastate and five deeply divided by wide median groove and two are post-dental; ridges 2-6 are divided by medial gap; and ridges 2-3 are underdeveloped, sometimes fused. Habitat. Southern Rainforest-Savanna Mosaic biotic zone in open secondary grasslands and possibly woodland savannas of Zambezian Woodland biotic zone at elevations of ¢. 600 m . Food and Feeding. Hayman'’s Epauletted Fruit Bat is frugivorous and nectarivorous. Breeding. One pregnant Hayman’s Epauletted Fruit Bat was recorded in June-July. Activity patterns. Hayman'’s Epauletted Fruit Bats are nocturnal. Movements, Home range and Social organization. One Hayman'’s Epauletted Fruit Bat was collected from a cave entrance. Status and Conservation. Classified as Data Deficient on The [UCN Red List (as Micropteropus intermedius ). Habitat loss due to deforestation might be a threat to Hayman’s Epauletted Fruit Bat, but additional research is needed to clarify its population and conservation status. It occurs in Mangroves National Park in DR Congo . Bibliography. Bergmans (1989), Happold, M. (2013), Mickleburgh, Hutson, Bergmans & Fahr (2008), Monadjem, Taylor et al. (2010). Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022. Pteropodidae Epomophorus intermedius Epomophorus intermedius Hayman 1963 1 Publ. Cult. Comp. Diamantes Angola 67:40:00 Hayman's Lesser Epauletted Fruit Bat None. Angola, Lunda, Dundo. N Angola, SE Dem. Rep. Congo. Not listed. Data Deficient as Micropteropus intermedius Previously considered a species of Micropteropus , but moved to Epomophorus by Almeida et al. (2020). Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023 Epomophorus intermedius 23 Hayman's Epauletted Fruit Bat Hayman's Dwarf Epauletted Fruit Bat|Hayman's Lesser Epauletted Fruit Bat Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA PTEROPODIFORMES NA NA PTEROPODOIDEA PTEROPODIDAE ROUSETTINAE EPOMOPHORINI Epomophorus NA intermedius Hayman 1963 1 Micropteropus_intermedius Hayman, R. W. (1963). Mammals from Angola, mainly from the Lunda District. Publicações Culturais da Companhia de Diamantes de Angola, 66, 100. BM 1962.2073 Dundo, Lunda, Angola. intermedius (Hayman, 1963) moved from Micropteropus to Epomophorus Almeida, F. C., Simmons, N. B., & Giannini, N. P. (2020). A Species-level Phylogeny of Old World Fruit Bats with a New Higher-level Classification of the Family Pteropodidae. American Museum Novitates, 3950, 1-24. Angola|Democratic Republic of the Congo Africa Afrotropic DD 0 0 0 Epomophorus_intermedius 0 sciname match Micropteropus_intermedius 0 IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022]. 13401 Micropteropus intermedius ANIMALIA CHORDATA MAMMALIA CHIROPTERA PTEROPODIDAE Micropteropus intermedius Hayman, 1963 20000000 Micropteropus intermedius Data Deficient 2020 2020-06-23 00:00:00 UTC 3.1 English Listed as Data Deficient in view of the absence of recent information on its extent of occurrence, ecological requirements, threats and conservation status. This species is generally associated with moist savanna and forest mosaic habitats. Populations most probably remain south of the Central African rain forest block. Whether or not it does occur in woodlands to the south, which seems more likely, remains to be discovered (Bergmans 1989). The specimen from Thysville was found roosting under a rock near the entrance to a cave (Bergmans 1989). The threats to this species are not well known, but may include general habitat loss through ongoing deforestation. It is currently only known from four specimens. Some of these records are old, and the species does not appear to have been recorded since the 1950s. Unknown This little known species has only been recorded from four localities in northern Angola (the type locality of Dundo) and southern Democratic Republic of the Congo (Luluabourg, Banana/Netonna, and Thysville). Terrestrial There appear to be no direct conservation measures in place. It is not known if the species is present in any protected areas. Further surveys are needed to locate populations of this species, and to record additional details on natural history and possible threats. Afrotropical FALSE FALSE Global Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 Pteropodidae Epomophorus intermedius Hayman 1963 1 Publ. Cult. Comp. Diamantes Angola 67:40:00 Hayman's Lesser Epauletted Fruit Bat None. Angola, Lunda, Dundo. N Angola, SE Dem. Rep. Congo. Not listed. Data Deficient as Micropteropus intermedius Previously considered a species of Micropteropus , but moved to Epomophorus by Almeida et al. (2020). Epomophorus intermedius 1004522 23 Hayman's Epauletted Fruit Bat Hayman's Dwarf Epauletted Fruit Bat|Hayman's Lesser Epauletted Fruit Bat Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA PTEROPODIFORMES NA NA PTEROPODOIDEA Pteropodidae ROUSETTINAE EPOMOPHORINI Epomophorus NA intermedius Hayman 1963 1 Micropteropus_intermedius Hayman, R. W. (1963). Mammals from Angola, mainly from the Lunda District. Publicações Culturais da Companhia de Diamantes de Angola, 66, 100. BM 1962.2073 Dundo, Lunda, Angola. intermedius (Hayman, 1963) moved from Micropteropus to Epomophorus Almeida, F. C., Simmons, N. B., & Giannini, N. P. (2020). A Species-level Phylogeny of Old World Fruit Bats with a New Higher-level Classification of the Family Pteropodidae. American Museum Novitates, 3950, 1-24. Angola|Democratic Republic of the Congo Africa Afrotropic DD 0 0 0 Epomophorus_intermedius 0 sciname match Micropteropus_intermedius 0 Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393 Epomophorus_intermedius 1004522 23 Hayman's Epauletted Fruit Bat Hayman's Dwarf Epauletted Fruit Bat|Hayman's Lesser Epauletted Fruit Bat Theria Placentalia Boreoeutheria Laurasiatheria Chiroptera Yinpterochiroptera NA NA Pteropodoidea Pteropodidae Epomophorinae Epomophorini Epomophorus NA intermedius Hayman 1 Micropteropus intermedius Hayman, R.W. 1963-10-21. Mammals from Angola, mainly from the Lunda District. Publicações culturais, Companhia de Diamantes de Angola 66:85-139. BMNH:Mamm:1962.2073 holotype https://data.nhm.ac.uk/object/e1b2c834-e033-4dc6-ae95-80143083044d Dundo, Lunda, Angola. moved from Micropteropus to Epomophorus Almeida, F. C., Simmons, N. B., & Giannini, N. P. (2020). A Species-level Phylogeny of Old World Fruit Bats with a New Higher-level Classification of the Family Pteropodidae. American Museum Novitates, 3950, 1-24. Angola|Democratic Republic of the Congo Africa Afrotropic DD 0 0 0 Epomophorus_intermedius 0 sciname match Micropteropus_intermedius 0 Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586 Pteropodidae Epomophorus intermedius Hayman 1963 1 Publ. Cult. Comp. Diamantes Angola 67:40:00 Hayman's Lesser Epauletted Fruit Bat None. Angola, Lunda, Dundo. N Angola, SE Dem. Rep. Congo. Not Listed Data Deficient Previously considered a species of Micropteropus, but moved to Epomophorus by Almeida et al. (2020). Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505 NA Micropteropus intermedius; Epomophorus intermedius; Epomophorus intermedius; Epomophorus intermedius; Micropteropus intermedius; Epomophorus intermedius; intermedius; Epomophore de Hayman; Hayman-Epaulettenflughund; Epoméforo de Hayman; Hayman’'s Dwarf Epauletted Fruit Bat; Hayman’s Lesser Epauletted Fruit Bat; Hayman's Epauletted Fruit Bat; Hayman's Dwarf Epauletted Fruit Bat; Hayman's Lesser Epauletted Fruit Bat; Hayman's Lesser Epauletted Fruit Bat; Hayman's Lesser Epauletted Fruit Bat; E. intermedius