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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L28	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Pipistrellus angulatus [synonym of]	N/A	Pipistrellus collinus	Pipistrellus tenuis [synonym of]	Pipistrellus tenuis collinus	Pipistrellus collinus	Pipistrellus collinus	Pipistrellus collinus	Pipistrellus collinus	Pipistrellus collinus	Pipistrellus collinus	Pipistrellus collinus	Pipistrellus collinus	Pipistrellus collinus	Alionoctula collina		[MSW3] Subgenus Pipistrellus. Included in angulatus by Laurie and Hill (1954) and in tenuis by Koopman (1993, 1994), but see Kitchener et al. (1986). Also see Flannery (1995a) and Bonaccoroso (1998).; [HMW] Pipistrellus papuanus collinus Thomas, 1920 , “Bihagi, head of Mambare River , British Papua [= Papua New Guineal.” Pipistrellus collinus and all other Papuan and Australian Pipistrellus species have been included under ' P tenuis but are considered distinct species here based on morphology. Monotypic.; [batnames2022] Subgenus Pipistrellus . Included in angulatus by Laurie and Hill (1954) and in tenuis by Koopman (1993, 1994), butsee Kitchener et al. (1986). Also see Flannery (1995 a ) and Bonaccoroso (1998).; [IUCN] Koopman in Wilson and Reeder (1993) treated this as a synonym of P. tenuis whereas Simmons (2005) considers it a valid species.; [batnames2023] Subgenus Pipistrellus . Included in angulatus by Laurie and Hill (1954) and in tenuis by Koopman (1993, 1994), butsee Kitchener et al. (1986). Also see Flannery (1995 a ) and Bonaccoroso (1998).; [batnames2025_1.7] Subgenus Pipistrellus. Included in angulatus by Laurie and Hill (1954) and in tenuis by Koopman (1993, 1994), butsee Kitchener et al. (1986). Also see Flannery (1995a) and Bonaccoroso (1998).; [MDD2025_2.2] moved from Pipistrellus to Alionoctula														collinus	Koopman in Wilson and Reeder (1993) treated this as a synonym of P. tenuis whereas Simmons (2005) considers it a valid species.			collinus	collinus			collina (O. Thomas, 1920)						N/A							Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Papua New Guinea; ref. 4.128																															NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Pipistrellini	Pipistrellus collinus	Pipistrellus	Pipistrellus	collinus	Thomas		1920		Ann. Mag. Nat. Hist., ser. 9	6		533		Greater Papuan Pipistrelle	British Papua (= Papua New Guinea), head of Mambare River, Bihagi, 8°04'S, 148°01'E.	Highlands of Papua New Guinea.	IUCN 2003 – Not listed (lapsus); IUCN/SSC Action Plan (2001) – Lower Risk (lc).		Subgenus Pipistrellus. Included in angulatus by Laurie and Hill (1954) and in tenuis by Koopman (1993, 1994), but see Kitchener et al. (1986). Also see Flannery (1995a) and Bonaccoroso (1998).	4C3D87E8FFE36A5CFF4B951F1D2AB15B	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	782	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFE36A5CFF4B951F1D2AB15B.xml	Pipistrellus collinus	Vespertilionidae	Pipistrellus	collinus	Thomas	1920	Pipistrelle des montagnes @fr | Gro RRe Papua-Zwergfledermaus @de | Pipistrelagrande de Papua @es | Mountain Pipistrelle @en	Pipistrellus papuanus collinus Thomas, 1920 , “Bihagi, head of Mambare River , British Papua [= Papua New Guineal.” Pipistrellus collinus and all other Papuan and Australian Pipistrellus species have been included under ' P tenuis but are considered distinct species here based on morphology. Monotypic.	Central Cordillera of New Guinea and Goodenough I.	Head-body 34-8-53 mm,tail 30-45-2 mm, ear 7-5-13 mm, hindfoot 6-5—-10 mm, forearm 34-38-5 mm; weight 4-3-8-5 g. The Greater Papuan Pipistrelle is the largest Pipistrellus in New Guinea . It is apparently sexually dimorphic according to T. F. Flannery in 1995, with males being more drab brown and females more reddish or gold-tinged. Pelage is long. Dorsum varies from drab medium brown to reddish; venter is considerably lighter and buffy. Face, ears, and membranes are medium brown, and uropatagium has sparse buffy brown hairs along venation. Ears are subtriangular, with broadly rounded tips and convex anterior edges; tragus is elongated, with curved tip. Uropatagium stretches nearly to end of tail (only extreme tip is free), and postcalcarial lobe is elongated. Baculum is long with pronounced ventral notch, has narrow shaft, and is flattened dorsally and bifurcated at tip (making up 10% of length). Skull is large and similar to that of the New Guinea Pipistrelle ( P. angulatus ) in overall shape and tooth morphology butis larger in all other characteristics; sagittal crest is absent; lambdoidal crest is moderately developed laterally but absent at apex of skull; I? is high, and I’ is higher than second cusp of I*; I? is bicuspid, and I” is unicuspid; P* is one-third the height of P*; and lower molars are nyctalodont.	Tropical hill forests, mid-montane forests, and rural gardens at 700-2900 m.	The Greater Papuan Pipistrelle is insectivorous.	Two lactating Greater Papuan Pipistrelles were collected in June and nonreproductive females in March.	Greater Papuan Pipistrelles are nocturnal. It has been reported roosting in hollow trees ( Pandanus , Pandanaceae ). Search-call shape is FM/QCEF, with characteristic frequency of 40 kHz; upsweep of call is between 10 kHz and 30 kHz.	Most recorded roosts of Greater Papuan Pipistrelles consist of a single individual (solitary males), and one roost had an adult male and eleven adult females, suggesting harem-like social and reproductive systems.	Classified as Least Concern on The IUCN Red List. Very little is currently known about ecology of the Greater Papuan Pipistrelle. It is reportedly locally common. It does seem to face any major threats, and much of its habitat is intact.	Bonaccorso (1998) | Flannery (1995b) | Hill & Harrison (1987) | Hutson, Schlitter, Csorba & Bonaccorso (2008) | Kitchener et al. (1986) | Robson et al. (2012)	https://zenodo.org/record/6397860/files/figure.png	43. Greater Papuan Pipistrelle Pipistrellus collinus French: Pipistrelle des montagnes / German: GroRRe Papua-Zwergfledermaus / Spanish: Pipistrela grande de Papua Other common names: Mountain Pipistrelle Taxonomy. Pipistrellus papuanus collinus Thomas, 1920 , “Bihagi, head of Mambare River , British Papua [= Papua New Guineal.” Pipistrellus collinus and all other Papuan and Australian Pipistrellus species have been included under ' P tenuis but are considered distinct species here based on morphology. Monotypic. Distribution. Central Cordillera of New Guinea and Goodenough I. Descriptive notes. Head-body 34-8-53 mm,tail 30-45-2 mm, ear 7-5-13 mm, hindfoot 6-5—-10 mm, forearm 34-38-5 mm; weight 4-3-8-5 g. The Greater Papuan Pipistrelle is the largest Pipistrellus in New Guinea . It is apparently sexually dimorphic according to T. F. Flannery in 1995, with males being more drab brown and females more reddish or gold-tinged. Pelage is long. Dorsum varies from drab medium brown to reddish; venter is considerably lighter and buffy. Face, ears, and membranes are medium brown, and uropatagium has sparse buffy brown hairs along venation. Ears are subtriangular, with broadly rounded tips and convex anterior edges; tragus is elongated, with curved tip. Uropatagium stretches nearly to end of tail (only extreme tip is free), and postcalcarial lobe is elongated. Baculum is long with pronounced ventral notch, has narrow shaft, and is flattened dorsally and bifurcated at tip (making up 10% of length). Skull is large and similar to that of the New Guinea Pipistrelle ( P. angulatus ) in overall shape and tooth morphology butis larger in all other characteristics; sagittal crest is absent; lambdoidal crest is moderately developed laterally but absent at apex of skull; I? is high, and I’ is higher than second cusp of I*; I? is bicuspid, and I” is unicuspid; P* is one-third the height of P*; and lower molars are nyctalodont. Habitat. Tropical hill forests, mid-montane forests, and rural gardens at 700-2900 m. Food and Feeding. The Greater Papuan Pipistrelle is insectivorous. Breeding. Two lactating Greater Papuan Pipistrelles were collected in June and nonreproductive females in March. Activity patterns. Greater Papuan Pipistrelles are nocturnal. It has been reported roosting in hollow trees ( Pandanus , Pandanaceae ). Search-call shape is FM/QCEF, with characteristic frequency of 40 kHz; upsweep of call is between 10 kHz and 30 kHz. Movements, Home range and Social organization. Most recorded roosts of Greater Papuan Pipistrelles consist of a single individual (solitary males), and one roost had an adult male and eleven adult females, suggesting harem-like social and reproductive systems. Status and Conservation. Classified as Least Concern on The IUCN Red List. Very little is currently known about ecology of the Greater Papuan Pipistrelle. It is reportedly locally common. It does seem to face any major threats, and much of its habitat is intact. Bibliography. Bonaccorso (1998), Flannery (1995b), Hill & Harrison (1987), Hutson, Schlitter, Csorba & Bonaccorso (2008), Kitchener et al. (1986), Robson et al. (2012).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Pipistrellus collinus	Pipistrellus	Pipistrellus	collinus	Thomas	1920	0	Ann. Mag. Nat. Hist., ser. 9	ser. 9, 6: 533	Greater Papuan Pipistrelle	None.	British Papua (= Papua New Guinea), head of Mambare River, Bihagi, 8°04'S, 148°01'E.	Highlands of Papua New Guinea.	Not listed.	Least Concern	Subgenus Pipistrellus . Included in angulatus by Laurie and Hill (1954) and in tenuis by Koopman (1993, 1994), butsee Kitchener et al. (1986). Also see Flannery (1995 a ) and Bonaccoroso (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Pipistrellus collinus	23	Greater Papuan Pipistrelle	Mountain Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PIPISTRELLINI	Pipistrellus	NA	collinus	O. Thomas	1920	0	Pipistrellus_papuanus_collinus	Thomas, O. (1920). New small mammals from New Guinea. Annals and Magazine of Natural History, Ser. 9, 6, 533.	https://www.biodiversitylibrary.org/item/63417#page/545/mode/1up	BM 1913.11.7.4		"Bihagi, head of Mambare River, British Papua [= Papua New Guinea]."			collinus O. Thomas, 1920	NA	NA	Indonesia|Papua New Guinea	Oceania	Australasia/Oceania	LC	0	0	0	Pipistrellus_collinus	0	sciname match	Pipistrellus_collinus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	17334	Pipistrellus collinus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Pipistrellus	collinus	Thomas, 1920	Koopman in Wilson and Reeder (1993) treated this as a synonym of P. tenuis whereas Simmons (2005) considers it a valid species.	20000000	Pipistrellus collinus	Least Concern		2020	2019-07-20 00:00:00 UTC	3.1	English	This species is listed as Least Concern. Although there is little information on population trends, there are no obvious threats to the species (especially given its preference for hill and montane forests), it is widely distributed, and not thought to be declining.	The Mountain Pipistrelle has been reported from tropical hill forests, montane forests, and from subsistence gardens. Animals have been recorded roosting in tree hollows (Cole et al. 1997) and the hollows of trunks of Pandanus (Flannery 1995). Males have been recorded roosting singularly (Flannery 1995) or in mixed sex groups (Cole et al. 1997). Flannery (1995) records a group of one male with eleven females and suggests that the species may possess a harem-like social system with seasonal reproduction. Two females collected in the month of June were lactating. Apart from this, nothing else is known about the species reproduction.  It is an aerial insectivore. On the Huon Peninsula, the species echolocation call is frequency modulated, with a characteristic frequency around 40 kHz (Robson et al. 2012 - based on 5 calls recorded on release).	There are no known major threats to this species. Much of the montane habitat used by this species are intact and are not generally considered to be threatened. The absence of large-scale industrial logging from the highlands of Papua New Guinea (PNG) explains the comparatively low rate of forest change (~ 1 % cleared between 2002 and 2014) compared to the lowlands of PNG (Bryan et al. 2015). However, forest fires particularly during El Niňo years, may threaten montane forests resulting in the encroachment of grassland and scrub into previously forested areas (Shearman and Bryan 2011).	Population abundance and trends are unknown but are likely stable. Flannery (1995) found it to be common in the Kwiyawagi area of Papua Province, Indonesia. It can be locally common.	Stable	This poorly-known species has been recorded from a number of scattered localities along the Central Cordillera of the island of New Guinea (Indonesia and Papua New Guinea), and from Goodenough Island, Milne Bay Province, Papua New Guinea (Flannery 1995, Bonaccorso 1998). It was also recently recorded in YUS Conservation Area (6.02756° S, 146.83376° E) on the Huon Peninsula at 2050 m (Robson et al. 2012) and may occur throughout the mountains of the Huon Peninsula. It is found between 700 and 3,000 m asl (Helgen 2007, Bonaccorso 1998, Flannery 1995).	Unlikely to be traded or used for food given its small size.	Terrestrial	This species occurs in YUS Conservation Area and is presumed to be present in other protected areas. Further studies are needed into the distribution, abundance, natural history, and threats to this species.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Pipistrellus	Pipistrellus	collinus	Thomas	1920	0	Ann. Mag. Nat. Hist., ser. 9	ser. 9, 6: 533	Greater Papuan Pipistrelle	None.	British Papua (= Papua New Guinea), head of Mambare River, Bihagi, 8°04'S, 148°01'E.	Highlands of Papua New Guinea.	Not listed.	Least Concern	Subgenus Pipistrellus . Included in angulatus by Laurie and Hill (1954) and in tenuis by Koopman (1993, 1994), butsee Kitchener et al. (1986). Also see Flannery (1995 a ) and Bonaccoroso (1998).	Pipistrellus collinus	1005614	23	Greater Papuan Pipistrelle	Mountain Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PIPISTRELLINI	Pipistrellus	NA	collinus	O. Thomas	1920	0	Pipistrellus_papuanus_collinus	Thomas, O. (1920). New small mammals from New Guinea. Annals and Magazine of Natural History, Ser. 9, 6, 533.	https://www.biodiversitylibrary.org/item/63417#page/545/mode/1up	BM 1913.11.7.4		"Bihagi, head of Mambare River, British Papua [= Papua New Guinea]."			collinus O. Thomas, 1920	NA	NA				Indonesia|Papua New Guinea	Oceania	Australasia/Oceania	LC	0	0	0	Pipistrellus_collinus	0	sciname match	Pipistrellus_collinus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Alionoctula_collina	1005614	23	Greater Papuan Pipistrelle	Mountain Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Pipistrellini	Alionoctula	NA	collina	O. Thomas	1	Pipistrellus papuanus collinus	Thomas, O. 1920-11-30. New small mammals from New Guinea. Annals and Magazine of Natural History (9)6(36):533-537.	https://www.biodiversitylibrary.org/page/15687828	BMNH:Mamm:1913.11.7.4	holotype	https://data.nhm.ac.uk/object/4c204721-4eff-4618-beea-52b2b7561c2d	"Bihagi, head of Mambare River, British Papua [= Papua New Guinea]."	-8.06667	148.0167	moved from Pipistrellus to Alionoctula	Zhukova, S. S., Yuzefovich, A. P., Lebedev, V. S., & Kruskop, S. V. (2025). Reassessment of the Taxonomic Borders Within Pipistrellus (Chiroptera, Vespertilionidae, Pipistrellini). Diversity, 17(5), 317.				Indonesia|Papua New Guinea	Oceania (Continent)	Australasia	LC	0	0	0	Pipistrellus_collinus	0	sciname match	Pipistrellus_collinus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Pipistrellus	Pipistrellus	collinus	Thomas	1920	0	Ann. Mag. Nat. Hist.	ser. 9, 6: 533	Greater Papuan Pipistrelle	None.	British Papua (= Papua New Guinea), head of Mambare River, Bihagi, 8°04'S, 148°01'E.	Highlands of Papua New Guinea.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/17334/22130362/' target='_blank'>Least Concern</a>	Subgenus Pipistrellus. Included in angulatus by Laurie and Hill (1954) and in tenuis by Koopman (1993, 1994), butsee Kitchener et al. (1986). Also see Flannery (1995a) and Bonaccoroso (1998).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Pipistrellus collinus; Pipistrellus collinus; Pipistrellus collinus; Pipistrellus collinus; Pipistrellus collinus; Pipistrellus collinus; collinus; Pipistrelle des montagnes; Gro RRe Papua-Zwergfledermaus; Pipistrelagrande de Papua; Mountain Pipistrelle; Greater Papuan Pipistrelle; Mountain Pipistrelle; Greater Papuan Pipistrelle; Greater Papuan Pipistrelle; P. collinus