http://www.w3.org/ns/prov#wasDerivedFrom	http://purl.org/dc/elements/1.1/format	name_CH1_1980	name_MSW1_1982	name_CH3_1991	name_MSW2_1993	name_Koopman_1994	name_MSW3_2005	name_HMW_2019	name_BatNames_2022	name_MDD_2022	name_IUCN_2022	name_BatNames_2023	name_MDD_2023	name_MDD_2025_2.0	name_batnames_2025_1.7	name_MDD_2025_2.2	column151	taxonomic_notes_concatenated	column171	synonyms_CH1	subspecies__MSW2	synonyms__MSW1	synonyms_CH3	synonyms_MSW2	subspecies_Koopman94_interpreted	subspecies_MSW3_interpreted	synonym_MSW3_interpreted	subspecies_HMW_interpreted	synonym_HMW_interpreted	subspecies_batnames_interpreted	synonym_batnames_interpreted	synonym_MDD_interpreted	synonym_IUCN_interpreted	subspecies_batnames2023_interpreted	synonym_batnames2023_interpreted	synonym_MDD2023_interpreted	synonym_MDD2025_interpreted	subspecies_batnames2025_interpreted	synonyms_batnames2025_interpreted	nominalNames	column391	docOrigin_CH1	commonName_CH1	distribution_CH1	docOrigin_MSW1	column451	typeLocality_MSW1	authority_MSW1	year_MSW1	citation_MSW1	distribution	comment_MSW1	docOrigin_CH3	commonName_CH3	distribution_CH3	docOrigin_MSW2	authority_MSW2	year_MSW2	citation_MSW2	comments_MSW2	distribution_MSW2	typeLocality_MSW2	docOrigin_Koopman94	authority_Koopman94	year_Koopman94	description_Koopman94	distribution_Koopman94	diversity_Koopman94	subspecies_Koopman94	page	rank	name	authority	year	parent	parent_rank	corrected_name	actual_species_count	claimed_species_count	dental_formula	description	diversity	full_subspecies_text	name_line	species_index	subspecies	synonym	text	docOrigin_MSW3	order_MSW3	family_MSW3	subfamily_MSW3	tribe_MSW3	name_MSW3	genus_MSW3	subgenus_MSW3	species_MSW3	authoritySpeciesAuthor_MSW3	(parentheses (1=author & date in parentheses)_MSW3	authoritySpeciesYear_MSW3	actualDate_MSW3	citation_MSW3	volume_MSW3	issue_MSW3	pages_MSW3	type_species_MSW3	commonName_MSW3	typeLocality_MSW3	distribution_MSW3	status_MSW3	synonym_MSW3	comments_MSW3	docId_HMW	docOrigin_HMW	docISBN_HMW	docName_HMW	docMasterId_HMW	docPageNumber_HMW	derivedFrom_HMW	name_HMW	family_HMW	genus_HMW	species_HMW	authoritySpeciesAuthor_HMW	authoritySpeciesYear	commonNames_HMW	taxonomy_HMW	subspeciesAndDistribution_HMW	descriptiveNotes_HMW	habitat_HMW	foodAndFeeding_HMW	breeding_HMW	activityPatterns_HMW	movementsHomeRangeAndSocialOrganization_HMW	statusAndConservation_HMW	bibliography_HMW	distributionImageURL_HMW	verbatimText_HMW	docOrigin_batnames	family_batnames	name_batnames	genus_batnames	subgenus_batnames	species_batnames	authoritySpeciesAuthor_batnames	date_batnames	parentheses_batnames (1=author & date in parentheses)	citation_batnames	docPageNumber_batnames	common Name_batnames	synonyms_batnames	type_locality_batnames	Distribution_batnames	CITES_batnames	IUCN_batnames	comments_batnames	docOrigin_MDD	name_MDD	phylosort_MDD	mainCommonName_MDD	otherCommonNames_MDD	subclass_MDD	infraclass_MDD	magnorder_MDD	superorder_MDD	order_MDD	suborder_MDD	infraorder_MDD	parvorder_MDD	superfamily_MDD	family_MDD	subfamily_MDD	tribe_MDD	genus_MDD	subgenus_MDD	specificEpithet_MDD	authoritySpeciesAuthor_MDD	authoritySpeciesYear_MDD	authorityParentheses_MDD	originalNameCombination_MDD	authoritySpeciesCitation_MDD	authoritySpeciesLink_MDD	holotypeVoucher_MDD	holotypeVoucherURIs_MDD	typeLocality_MDD	typeLocalityLatitude_MDD	typeLocalityLongitude_MDD	nominalNames_MDD	taxonomyNotes_MDD	taxonomyNotesCitation_MDD	countryDistribution_MDD	continentDistribution_MDD	biogeographicRealm_MDD	iucnStatus_MDD	extinct_MDD	domestic_MDD	flagged_MDD	CMW_sciName_MDD	diffSinceCMW_MDD	MSW3_matchtype_MDD	MSW3_sciName_MDD	diffSinceMSW3_MDD	docOrigin_IUCN	internalTaxonId_IUCN	NAME_IUCN	kingdomName_IUCN	phylumName_IUCN	className_IUCN	orderName_IUCN	familyName_IUCN	genusName_IUCN	speciesName_IUCN	authoritySpeciesAuthorYear_IUCN	taxonomicNotes_IUCN	assessmentId_IUCN	scientificName_IUCN	redlistCategory_IUCN	redlistCriteria_IUCN	yearPublished_IUCN	assessmentDate_IUCN	criteriaVersion_IUCN	language_IUCN	rationale_IUCN	habitat_IUCN	threats_IUCN	population_IUCN	populationTrend_IUCN	range_IUCN	useTrade_IUCN	systems_IUCN	conservationActions_IUCN	realm_IUCN	yearLastSeen_IUCN	possiblyExtinct_IUCN	possiblyExtinctInTheWild_IUCN	scopes_IUCN	docOrigin_batnames2023	FAMILY_batnames2023	GENUS_batnames2023	SUBGENUS_batnames2023	SPECIES_batnames2023	authoritySpeciesAuthor_batnames2023	authoritySpeciesYearbatnames2023	PARENTHESES_batnames2023 (1=AUTHOR & DATE IN PARENTHESES)	CITATION_batnames2023	PAGES_batnames2023	COMMON NAME_batnames2023	SYNONYMS_batnames2023	TYPE LOCALITY_batnames2023	DISTRIBUTION_batnames2023	CITES_batnames2023	IUCN_batnames2023	COMMENTS_batnames2023	name MDD2023	id_MDD2023	phylosort_MDD2023	mainCommonName_MDD2023	otherCommonNames_MDD2023	subclass_MDD2023	infraclass_MDD2023	magnorder_MDD2023	superorder_MDD2023	order_MDD2023	suborder_MDD2023	infraorder_MDD2023	parvorder_MDD2023	superfamily_MDD2023	Family_mdd2023	subfamily_MDD2023	tribe_MDD2023	genus_MDD2023	subgenus_MDD2023	specificEpithet_MDD2023	authoritySpeciesAuthor_MDD2023	authoritySpeciesYear_MDD2023	authorityParentheses_MDD2023	originalNameCombination_MDD2023	authoritySpeciesCitation_MDD2023	authoritySpeciesLink_MDD2023	holotypeVoucher_MDD2023	holotypeVoucherURIs_MDD2023	typeLocality_MDD2023	typeLocalityLatitude_MDD2023	typeLocalityLongitude_MDD2023	nominalNames_MDD2023	taxonomyNotes_MDD2023	taxonomyNotesCitation_MDD2023	distributionNotes_MDD2023	distributionNotesCitation_MDD2023	subregionDistribution_MDD2023	countryDistribution_MDD2023	continentDistribution_MDD2023	biogeographicRealm_MDD2023	iucnStatus_MDD2023	extinct_MDD2023	domestic_MDD2023	flagged_MDD2023	CMW_sciName_MDD2023	diffSinceCMW_MDD2023	MSW3_matchtype_MDD2023	MSW3_sciName_MDD2023	diffSinceMSW3_MDD2023	docOrigin_MDD2025	sciName	id	phylosort	mainCommonName	otherCommonNames	subclass	infraclass	magnorder	superorder	order	suborder	infraorder	parvorder	superfamily	family	subfamily	tribe	genus	subgenus	specificEpithet	authoritySpeciesAuthor	authorityParentheses	originalNameCombination	authoritySpeciesCitation	authoritySpeciesLink	typeVoucher	typeKind	typeVoucherURIs	typeLocality	typeLocalityLatitude	typeLocalityLongitude	taxonomyNotes	taxonomyNotesCitation	distributionNotes	distributionNotesCitation	subregionDistribution	countryDistribution	continentDistribution	biogeographicRealm	iucnStatus	extinct	domestic	flagged	CMW_sciName	diffSinceCMW	MSW3_matchtype	MSW3_sciName	diffSinceMSW3	docOrigin_batnames2025	Family	Genus	Subgenus	Species	Author	Date	Parentheses (1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L253	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Dyacopterus rickarti	Dyacopterus rickarti	Dyacopterus rickarti	Dyacopterus rickarti	Dyacopterus rickarti	Dyacopterus rickarti	Dyacopterus rickarti	Dyacopterus rickarti	Dyacopterus rickarti		[HMW] Dyacopterus rickarti Helgen et al, 2007 , “San Isidro ( 17°27'N , 120°37’E ), Luzon ( Abra Province ), Philippines .” This species is monotypic.; [MDD2022] recently described; [MDD2023] recently described; [MDD2025_2.0] recently described; [MDD2025_2.2] recently described														rickarti				rickarti	rickarti			rickarti K. M. Helgen, Kock, Gomez, & Ingle in K. M. Helgen, Kock, Gomez, Ingle, & M. H. Sinaga, 2007						N/A																																								NA																											03AD87FAFFCAF62489B83515FF5EF3B4	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	72	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFCAF62489B83515FF5EF3B4.xml	Dyacopterus rickarti	Pteropodidae	Dyacopterus	rickarti	Helgen et al.	2007	Cynoptere de Rickart @fr | Philippinen-Dayak-Flughund @de | Dyacoéptero de Rickart @es | Rickart's Dayak Fruit Bat @en	Dyacopterus rickarti Helgen et al, 2007 , “San Isidro ( 17°27'N , 120°37’E ), Luzon ( Abra Province ), Philippines .” This species is monotypic.	Philippines , suspected to be widespread in the archipelago, but so far recorded in only one locality on Luzon I and four localities on Mindanao I.	Head-body 130- 144 mm , tail 18-29 mm , ear 21-25 mm , hindfoot 25 mm , forearm 91-96- 4 mm ; weight 138-148 g . Muzzle of the Philippine Large-headed Bat is stout, almost hairless, with grayish brown skin; nostrils are shortly tubular and divergent, with thickened rims and obvious philtrum, reaching upper lip that has faint, inconspicuous white mustache of very sparse hairs; lower lip has very large but not prominent paired pads and other small papillae. Eyes are moderately large; iris is dark brown. Ears are short and grayish brown, with pointed tips. Head is massive, very wide, and divided by obvious temporal muscles. Head pelage is very short and grayish brown and continues on nape, becoming warm brown along dorsum. Tail is long, surpassing wide uropatagium; calcar is short. Throat and chest are sparsely haired and light brown that becomes darker on flanks; belly is pale brown to cream. Small tuft of longer, warm brown hairs occurs on shoulder. Wing membranes are dark gray from sides of body and attach to second toe, index claw is present, and all claws are unpigmented and whitish. Skull has no basicranial deflection, is generally very robust, and long posteriorly. Laterally, premaxillae are fused; narial opening is deeply concave; rostrum is very deep, ascending and continued into flat forehead; juga of canine and cheekteeth roots are obvious; orbit is large, with well-marked rim; zygomatic root is above upper alveolar line; zygoma is strongly arched; and braincase is low and elongated. Dorsally, rostrum widens posteriorly, paranasal recesses are inflated, postorbital foramen is absent, postorbital constriction is obvious, postorbital processes are thick and divergent, forehead looks like a shield, temporal lines are joined in sharp sagittal crest, and braincase is oval and elongated, ending in marked nuchal crest. Ventrally, palate is flat; tooth rows diverge; post-dental short and converging, without palatine spine; sphenoidal crest is low; and ectotympanic is very wide, especially anteriorly, and internally edged by ribbon-like entotympanic. Mandible is stout; symphysis slopes, continuing in a convex ventral border; coronoid slopes steeply, almost vertical, and tip is rounded; condyle is strong, above lower alveolar line; and angle is stout, with marked rim. There are 15-16 palatal ridges, with posterior and some anterior ridges divided. Upper incisors are small and crowded; I' is longest; C' is straight and proclivous, has well-developed vertical groove on its anteromedial surface and obvious cingulum and posterolateral basal cusp; P' is absent; next premolar (P?) is massive, stout, high-crowned, very wide, and square in outline; last premolar has anteromedial cingular ledge; and M' is more elongated and low crowned. Lower incisors are very small and spaced; C, is short, stout, and strongly decurved; P, is relatively large and wide; posterior cheekteeth are massive, changing from high-crowned triangular outline to very wide, square outline in last premolar, to more elongated, low-crowned M; and M,is peg-like.	Primary and regenerating montane and mossy rainforest at elevations of 260- 1680 m . The Philippine Large-headed Batis suspected to use subcanopy and canopy strata.	The Philippine Large-headed Bat is primarily frugivorous, probably relying on figs ( Ficus spp. , Moraceae ) and hard fruits,judging from its robust skull and dentition.	A pregnant Philippine Large-headed Bat with one young was captured in May.	The few captures of Philippine Large-Headed Bats suggest that it uses open areas as flyways.	No information.	Not assessed on The [UCN Red List. The Philippine Largeheaded Bat was included under the Common Dayak Fruit Bat ( D. spadiceus ), which was classified as Near Threatened. Additional research is needed to evaluate conservation status of the Philippine Large-headed Bat, which is rare and infrequently captured. Present in some protected areas; e.g. Mount Kitanglad Range Nature Park (Mindanao).	Heaney, Balete & Rickart (2016) | Heaney, Tabaranza et al. (2006) | Helgen et al. (2007) | Kock (1969a)		17. Philippine Large-headed Fruit Bat Dyacopterus rickarti French: Cynoptere de Rickart / German: Philippinen-Dayak-Flughund / Spanish: Dyacoéptero de Rickart Other common names: Rickart's Dayak Fruit Bat Taxonomy. Dyacopterus rickarti Helgen et al, 2007 , “San Isidro ( 17°27'N , 120°37’E ), Luzon ( Abra Province ), Philippines .” This species is monotypic. Distribution. Philippines , suspected to be widespread in the archipelago, but so far recorded in only one locality on Luzon I and four localities on Mindanao I. Descriptive notes. Head-body 130- 144 mm , tail 18-29 mm , ear 21-25 mm , hindfoot 25 mm , forearm 91-96- 4 mm ; weight 138-148 g . Muzzle of the Philippine Large-headed Bat is stout, almost hairless, with grayish brown skin; nostrils are shortly tubular and divergent, with thickened rims and obvious philtrum, reaching upper lip that has faint, inconspicuous white mustache of very sparse hairs; lower lip has very large but not prominent paired pads and other small papillae. Eyes are moderately large; iris is dark brown. Ears are short and grayish brown, with pointed tips. Head is massive, very wide, and divided by obvious temporal muscles. Head pelage is very short and grayish brown and continues on nape, becoming warm brown along dorsum. Tail is long, surpassing wide uropatagium; calcar is short. Throat and chest are sparsely haired and light brown that becomes darker on flanks; belly is pale brown to cream. Small tuft of longer, warm brown hairs occurs on shoulder. Wing membranes are dark gray from sides of body and attach to second toe, index claw is present, and all claws are unpigmented and whitish. Skull has no basicranial deflection, is generally very robust, and long posteriorly. Laterally, premaxillae are fused; narial opening is deeply concave; rostrum is very deep, ascending and continued into flat forehead; juga of canine and cheekteeth roots are obvious; orbit is large, with well-marked rim; zygomatic root is above upper alveolar line; zygoma is strongly arched; and braincase is low and elongated. Dorsally, rostrum widens posteriorly, paranasal recesses are inflated, postorbital foramen is absent, postorbital constriction is obvious, postorbital processes are thick and divergent, forehead looks like a shield, temporal lines are joined in sharp sagittal crest, and braincase is oval and elongated, ending in marked nuchal crest. Ventrally, palate is flat; tooth rows diverge; post-dental short and converging, without palatine spine; sphenoidal crest is low; and ectotympanic is very wide, especially anteriorly, and internally edged by ribbon-like entotympanic. Mandible is stout; symphysis slopes, continuing in a convex ventral border; coronoid slopes steeply, almost vertical, and tip is rounded; condyle is strong, above lower alveolar line; and angle is stout, with marked rim. There are 15-16 palatal ridges, with posterior and some anterior ridges divided. Upper incisors are small and crowded; I' is longest; C' is straight and proclivous, has well-developed vertical groove on its anteromedial surface and obvious cingulum and posterolateral basal cusp; P' is absent; next premolar (P?) is massive, stout, high-crowned, very wide, and square in outline; last premolar has anteromedial cingular ledge; and M' is more elongated and low crowned. Lower incisors are very small and spaced; C, is short, stout, and strongly decurved; P, is relatively large and wide; posterior cheekteeth are massive, changing from high-crowned triangular outline to very wide, square outline in last premolar, to more elongated, low-crowned M; and M,is peg-like. Habitat. Primary and regenerating montane and mossy rainforest at elevations of 260- 1680 m . The Philippine Large-headed Batis suspected to use subcanopy and canopy strata. Food and Feeding. The Philippine Large-headed Bat is primarily frugivorous, probably relying on figs ( Ficus spp. , Moraceae ) and hard fruits,judging from its robust skull and dentition. Breeding. A pregnant Philippine Large-headed Bat with one young was captured in May. Activity patterns. The few captures of Philippine Large-Headed Bats suggest that it uses open areas as flyways. Movements, Home range and Social organization. No information. Status and Conservation. Not assessed on The [UCN Red List. The Philippine Largeheaded Bat was included under the Common Dayak Fruit Bat ( D. spadiceus ), which was classified as Near Threatened. Additional research is needed to evaluate conservation status of the Philippine Large-headed Bat, which is rare and infrequently captured. Present in some protected areas; e.g. Mount Kitanglad Range Nature Park (Mindanao). Bibliography. Heaney, Balete & Rickart (2016), Heaney, Tabaranza et al. (2006), Helgen et al. (2007), Kock (1969a).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Dyacopterus rickarti	Dyacopterus		rickarti	Helgen, Kock, Gomes & Ingle	2007	0	Journal of Mammalogy	88(2): 311	Phillippine Large-Headed Fruit Bat	None.	San Isidro, Luzon, Phillippines	Luzon and Mindanao (Phillippines)	Not listed.	Endangered		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Dyacopterus rickarti	23	Philippine Large-headed Fruit Bat	Rickart's Dayak Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	CYNOPTERINAE	BALIONYCTERINI	Dyacopterus	NA	rickarti	K. M. Helgen, Kock, Gomez, Ingle, & Sinaga	2007	0						"San Isidro (17°27'N, 120°37'E), Luzon (Abra Province), Philippines."	17.45	120.62	rickarti K. M. Helgen, Kock, Gomez, Ingle, & Sinaga, 2007	recently described	Helgen, K. M., Kock, D., Gomez, R. K. S. C., Ingle, N. R., & Sinaga, M. H. (2007). Taxonomy and natural history of the Southeast Asian fruit-bat genus Dyacopterus. Journal of Mammalogy, 88(2), 302-318.	Philippines	Asia	Indomalaya	EN	0	0	0	Dyacopterus_rickarti	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Dyacopterus rickarti	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Dyacopterus	rickarti	Helgen, Kock, Gomez, Ingle &; Sinaga, 2007		100000000	Dyacopterus rickarti	Endangered	B2ab(iii)	2020	2019-05-27 00:00:00 UTC	3.1	English	Dyacopterus rickarti is listed as Endangered under criterion B2ab(iii) as the species is restricted to a few mountain ranges on Mindanao Island and Northern Luzon, Philippines. Its population is highly fragmented and it is only known from seven captures in six locations (four in Mindanao and two in Luzon). Its area of occupancy is 36 km². There is also continuing loss in area, extent and quality of habitat in the mountains where it has been captured.	Before 2007, this species was initially categorized as Dyacopterus spadiceus , which is also found in neighboring Southeast Asian countries. Data from Helgen et al. (2007) showed differences from this species to that of other Dyacopterus species. Limited records for this species are from montane forest habitats generally above 1,000 m asl to 1,680 m asl, suggesting it is restricted relatively high elevations on montane to mossy forest. However, the Mt. Pangal capture in Northern Sierra Madre is the lowest elevation record for this species at 550 m asl and is characterized as a lower montane forest. Records from New Bataan and Mt. Mahuson and Mt. Apo National Park were from degraded habitats, which may indicate that this species can tolerate some level of disturbance. In Mt. Apo National Park, for example, this species was captured in a net placed in a cultivated area. More research needs to be done on distribution extent and ecological requirements of this species to determine specific conservation needs of this endemic fruit bat. If it is similar to Dyacopterus spadiceus , this species may require cave habitats.	The primary threats to the species is from past and ongoing forest habitat degradation and loss through the intrusion of agriculture, recreation/ecotourism, and rural communities into its known range. If the species is documented to use caves, like the similar Dyacopterus spadiceus , then we expect cave disturbance to be a significant threat as it is prevalent throughout the Philippines. While there are currently no records of hunting of this species, given its size, it is likely taken when other species are hunted in an area. More data are needed to ascertain the degree of threat from hunting.	Very little information is available about the population status for this species due to the limited data available. It is rarely captured, mostly at high elevations, and may be mistaken for similar species. More data are needed to determine population size and trends, and threats to this species.	Decreasing	Dyacopterus rickarti is a Philippine endemic species that has only been documented from seven (7) sites in six (6) locations, including Abra Province (550 m asl), Mt. Malibato, Opol, Misamis Oriental (altitude unrecorded), Mt. Kitanglad National Park, Sumilao, Bukidnon (1,450 m asl), Mt. Apo National Park, Davao City (1,185 m asl), Mt. Puting Bato, New Bataan, Compostela Valley (1,680 m asl), and Mt. Mahuson, Arakan Valley Conservation Area, Arakan, North Cotabato (1,260 m asl). Its area of occupancy (AOO) is 36 km². ;Additional information is needed to refine the understanding of the species distribution across the Philippines.		Terrestrial	The current distribution of this species is at relatively high elevations, 550 m asl and above, which are mostly forest areas protected under Philippine law. But with continued intrusion of agriculture, recreation/ecotourism, and communities, habitats of this species must be protected. More research is needed to determine distribution, population, abundance, ecological requirements, threats, conservation and management of this species.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Dyacopterus		rickarti	Helgen, Kock, Gomes & Ingle	2007	0	Journal of Mammalogy	88(2): 311	Phillippine Large-Headed Fruit Bat	None.	San Isidro, Luzon, Phillippines	Luzon and Mindanao (Phillippines)	Not listed.	Endangered		Dyacopterus rickarti	1004365	23	Philippine Large-headed Fruit Bat	Rickart's Dayak Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	CYNOPTERINAE	BALIONYCTERINI	Dyacopterus	NA	rickarti	K. M. Helgen, Kock, Gomez, & Ingle in K. M. Helgen, Kock, Gomez, Ingle, & Sinaga	2007	0						"San Isidro (17°27'N, 120°37'E), Luzon (Abra Province), Philippines."	17.45	120.6167	rickarti K. M. Helgen, Kock, Gomez, & Ingle in K. M. Helgen, Kock, Gomez, Ingle, & Sinaga, 2007	recently described	Helgen, K. M., Kock, D., Gomez, R. K. S. C., Ingle, N. R., & Sinaga, M. H. (2007). Taxonomy and natural history of the Southeast Asian fruit-bat genus Dyacopterus. Journal of Mammalogy, 88(2), 302-318.				Philippines	Asia	Indomalaya	EN	0	0	0	Dyacopterus_rickarti	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Dyacopterus_rickarti	1004365	23	Philippine Large-headed Fruit Bat	Rickart's Dayak Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Cynopterinae	Balionycterini	Dyacopterus	NA	rickarti	K. M. Helgen, Kock, Gomez, & Ingle in K. M. Helgen, Kock, Gomez, Ingle, & M. H. Sinaga	0	Dyacopterus rickarti	Helgen, K.M., Kock, D., Gomez, R.K.S.C., Ingle, N.R. and Sinaga, M.H. 2007-04-20. Taxonomy and natural history of the Southeast Asian fruit-bat genus _Dyacopterus_. Journal of Mammalogy 88(2):302-318.	https://doi.org/10.1644/06-MAMM-A-276R.1	SMF:MAMM:33333	holotype		"San Isidro (17°27'N, 120°37'E), Luzon (Abra Province), Philippines."	17.45	120.6167	recently described	Helgen, K. M., Kock, D., Gomez, R. K. S. C., Ingle, N. R., & Sinaga, M. H. (2007). Taxonomy and natural history of the Southeast Asian fruit-bat genus Dyacopterus. Journal of Mammalogy, 88(2), 302-318.				Philippines	Asia	Indomalaya	EN	0	0	0	Dyacopterus_rickarti	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Dyacopterus		rickarti	Helgen, Kock, Gomez & Ingle	2007	0	Journal of Mammalogy	88(2): 311	Phillippine Large-Headed Fruit Bat	None.	San Isidro, Luzon, Phillippines	Luzon and Mindanao (Phillippines)	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/84457541/95642280/' target='_blank'>Endangered</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Dyacopterus rickarti; Dyacopterus rickarti; Dyacopterus rickarti; Dyacopterus rickarti; Dyacopterus rickarti; rickarti; Cynoptere de Rickart; Philippinen-Dayak-Flughund; Dyacoéptero de Rickart; Rickart's Dayak Fruit Bat; Philippine Large-headed Fruit Bat; Rickart's Dayak Fruit Bat; Phillippine Large-Headed Fruit Bat; D. rickarti