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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L251	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Dryadonycteris capixaba	Dryadonycteris capixaba	Dryadonycteris capixaba	Dryadonycteris capixaba	Dryadonycteris capixaba	Dryadonycteris capixaba	Dryadonycteris capixaba	Dryadonycteris capixaba	Dryadonycteris capixaba		[HMW] Dryadonycteris capixaba Nogueira et al, 2012 , “ Brazil , state of Espirito Santo , municipality of Linhares, Reserva Natural Vale, 19°08'29” S and 40°04'08” W , approximately 60 m elevation.” This species is monotypic.; [MDD2022] recently described genus and species; [MDD2023] recently described genus and species; [MDD2025_2.0] recently described genus and species; [MDD2025_2.2] recently described genus and species														capixaba				capixaba	capixaba			capixaba Nogueira, I. P. Lima, A. L. Peracchi, & Simmons, 2012						N/A																																								NA																											03A687BCFF99FF9913AAFD5FFC09F83D	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	524	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFF99FF9913AAFD5FFC09F83D.xml	Dryadonycteris capixaba	Phyllostomidae	Dryadonycteris	capixaba	Nogueira	2012	Dryadonyctére d’Espirito Santo @fr | Capixaba-Dryades-Fledermaus @de | Driadonicterio de Capixaba @es	Dryadonycteris capixaba Nogueira et al, 2012 , “ Brazil , state of Espirito Santo , municipality of Linhares, Reserva Natural Vale, 19°08'29” S and 40°04'08” W , approximately 60 m elevation.” This species is monotypic.	NE & SE Brazil ( Pernambuco , Alagoas , Sergipe , Minas Gerais , and Espirito Santo ); it probably occurs in Bahia .	Head-body 49-9-56-5 mm, tail 4-4-6-4 mm, ear 9-12-1 mm, hindfoot 7-5-8: 3 mm , forearm 29-1-32.3 mm; weight 4-1-6 g. Dorsal fur of the Capixaba Nectarfeeding Bat is dark brown. Dorsal hairs are tricolored, with pale brown bases, beige middles, and dark brown tips. Venter is brown, slightly lighter than dorsum. Rostrum is elongated but shorter than braincase. Mandibular prognathism is evident. Chin has deep medial cleft, bordered by pair of dermal pads. Upper and lower lips are ornamented with well-developed vibrissae. Ears are round and small. Noseleaf is dark brown and relatively small, with no central rib. Wing membranes and uropatagium are dark brown. Plagiopatagium is inserted at bases of toes. Uropatagium is long and reaches distal part of tibia. Calcaris about the same length as foot. Tail is short, completely inserted in uropatagium. Rostrum is elongated and smaller than braincase, forming smooth slope in profile. Maxilla is inflated posteriorly. Basisphenoid pits are shallow, separated by wide septum. Zygomatic arches are incomplete. Palate is relatively long, broad, and U-shaped posteriorly. Mandibular condyleis level with coronoid process. Dental formula is 12/0,C1/1,P 2/3, M 3/3 ( x2 ) = 30. Upper incisors are diminutive and separated from each other and from canines by a gap. C! is laterally compressed, and M' and M? are longer than M>.	Humid, lowland Atlantic Forest. In Minas Gerais and Espirito Santo , the Capixaba Nectarfeeding Bat was captured in mature forest, with relatively open understory, canopy heights of 25-30 m, and emergent trees up to 40 m . One specimen from Sergipe was captured in well-structured forest surrounded by sugarcane plantations. It is also found in humid forest enclaves (“brejos de altitude”) in the Caatinga biome at elevations of ¢. 750 m .	Skull morphology and dentition of the Capixaba Nectar-feeding Bat suggest that it is a nectarivore.	No information.	Capixaba Nectar-feeding Bats were captured using ground-level mist nets set in clearings and borders and inside the forest. In Espirito Santo , one individual was captured at 18:05 h and another at 22:30 h.	No information.	Not assessed on The IUCN Red List. Recent studies of the Capixaba Nectar-feeding Bat showed thatits distribution is much larger than previously assumed, and it has been found in relatively large protected areas in Minas Gerais and Espirito Santo .	Gregorin et al. (2015) | Nogueira et al. (2012) | Rocha et al. (2014)	https://zenodo.org/record/6458764/files/figure.png	78. Capixaba Nectarfeeding Bat Dryadonycteris capixaba French: Dryadonyctére d’Espirito Santo / German: Capixaba-Dryades-Fledermaus / Spanish: Driadonicterio de Capixaba Taxonomy. Dryadonycteris capixaba Nogueira et al, 2012 , “ Brazil , state of Espirito Santo , municipality of Linhares, Reserva Natural Vale, 19°08'29” S and 40°04'08” W , approximately 60 m elevation.” This species is monotypic. Distribution. NE & SE Brazil ( Pernambuco , Alagoas , Sergipe , Minas Gerais , and Espirito Santo ); it probably occurs in Bahia . Descriptive notes. Head-body 49-9-56-5 mm, tail 4-4-6-4 mm, ear 9-12-1 mm, hindfoot 7-5-8: 3 mm , forearm 29-1-32.3 mm; weight 4-1-6 g. Dorsal fur of the Capixaba Nectarfeeding Bat is dark brown. Dorsal hairs are tricolored, with pale brown bases, beige middles, and dark brown tips. Venter is brown, slightly lighter than dorsum. Rostrum is elongated but shorter than braincase. Mandibular prognathism is evident. Chin has deep medial cleft, bordered by pair of dermal pads. Upper and lower lips are ornamented with well-developed vibrissae. Ears are round and small. Noseleaf is dark brown and relatively small, with no central rib. Wing membranes and uropatagium are dark brown. Plagiopatagium is inserted at bases of toes. Uropatagium is long and reaches distal part of tibia. Calcaris about the same length as foot. Tail is short, completely inserted in uropatagium. Rostrum is elongated and smaller than braincase, forming smooth slope in profile. Maxilla is inflated posteriorly. Basisphenoid pits are shallow, separated by wide septum. Zygomatic arches are incomplete. Palate is relatively long, broad, and U-shaped posteriorly. Mandibular condyleis level with coronoid process. Dental formula is 12/0,C1/1,P 2/3, M 3/3 ( x2 ) = 30. Upper incisors are diminutive and separated from each other and from canines by a gap. C! is laterally compressed, and M' and M? are longer than M>. Habitat. Humid, lowland Atlantic Forest. In Minas Gerais and Espirito Santo , the Capixaba Nectarfeeding Bat was captured in mature forest, with relatively open understory, canopy heights of 25-30 m, and emergent trees up to 40 m . One specimen from Sergipe was captured in well-structured forest surrounded by sugarcane plantations. It is also found in humid forest enclaves (“brejos de altitude”) in the Caatinga biome at elevations of ¢. 750 m . Food and Feeding. Skull morphology and dentition of the Capixaba Nectar-feeding Bat suggest that it is a nectarivore. Breeding. No information. Activity patterns. Capixaba Nectar-feeding Bats were captured using ground-level mist nets set in clearings and borders and inside the forest. In Espirito Santo , one individual was captured at 18:05 h and another at 22:30 h. Movements, Home range and Social organization. No information. Status and Conservation. Not assessed on The IUCN Red List. Recent studies of the Capixaba Nectar-feeding Bat showed thatits distribution is much larger than previously assumed, and it has been found in relatively large protected areas in Minas Gerais and Espirito Santo . Bibliography. Gregorin et al. (2015), Nogueira et al. (2012), Rocha et al. (2014).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Dryadonycteris capixaba	Dryadonycteris		capixaba	Nogueira, Lima, Peracchi, & Simmons	2012	0	Am. Mus. Novitates	######	Espirito Santo Atlantic Forest Bat	None.	Brazil, Espirito Santo, municipality of Linhares, Reserva Natural Vale	E Brazil.	Not listed.	Data Deficient		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Dryadonycteris capixaba	23	Capixaba Nectar-feeding Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	GLOSSOPHAGINAE	CHOERONYCTERINI	Dryadonycteris	NA	capixaba	Nogueira, I. P. Lima, Peracchi, & Simmons	2012	0	Dryadonycteris_capixaba	Nogueira, M. R., Lima, I. P., Peracchi, A. L. & Simmons, N. B. (2012). New genus and species of nectar-feeding bat from the Atlantic Forest of Southeastern Brazil (Chiroptera: Phyllostomidae: Glossophaginae). American Museum Novitates, 3747, 5.	http://digitallibrary.amnh.org/bitstream/handle/2246/6180/N3747.pdf?sequence=1&isAllowed=y	UFRRJ 9667		"Brazil, state of Espírito Santo, municipality of Linhares, Reserva Natural Vale, 19°08'29" S and 40°04'08" W, approximately 60 m elevation."	-19.14	-40.07	capixaba Nogueira, I. P. Lima, Peracchi, & Simmons, 2012	recently described genus and species	Nogueira, M. R., Lima, I. P., Peracchi, A. L., & Simmons, N. B. (2012). New genus and species of nectar-feeding bat from the Atlantic Forest of southeastern Brazil (Chiroptera: Phyllostomidae: Glossophaginae). American Museum Novitates, 3747, 1-30.	Brazil	South America	Neotropic	DD	0	0	0	Dryadonycteris_capixaba	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Dryadonycteris capixaba	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Dryadonycteris	capixaba	Nogueira, Lima, Peracchi &; Simmons, 2012		200000000	Dryadonycteris capixaba	Data Deficient		2020	2016-07-12 00:00:00 UTC	3.1	English	This species is listed as Data Deficient as it has only recently been described and its ecological requirements are poorly known. The current known distribution has been extended to the north and west, suggesting our current knowledge on this species is limited. There has been extensive habitat change and fragmentation within its known range. More surveys and review of existing collection are required. Nothing has been published on its ecological characteristics and requirements, and that is a key missing component for an accurate assessment.	The original locality where D. capixaba was caught is covered by tabuleiro forests, which in ;northern Espírito Santo includes the 'mata alta' (high forest) formation. The mata alta occurs on clay or sandy-clay soils and exhibits a thick canopy layer with most trees reaching around 30 m, and emergent trees towering up to 40 m; there are two other lower arboreal strata, with average heights of 14 and 9 m, respectively. The understory is relatively open, probably as a result of the intense shade from the closed canopy (Nogueira et al. 2012). The northernmost sites are 1,200 km north of the type locality, but on a similar habitat with canopy above 20 m (Rocha et al. 2014). The extent to which this species is dependent on specific floras or forest fragments of particular sizes remains to be determined, as does the full extent of its geographic range.	The potential of D. capixaba to persist in the fragmented landscapes of Atlantic Forest on eastern Brazil needs to be further investigated, particularly considering the fact that the forests covering the region are subject to intense pressure and transformation for resources.	Known from less than ten specimens from fives states in eastern Brazil. ;Assuming that the known localities do not represent two isolated populations in the north and south of the Atlantic Forest, it would seem reasonable to conclude that the species is distributed throughout most of this biome on the tropical eastern coast of Brazil, although it is presumably ;rare at most sites (Rocha et al.  2014).	Unknown	This species is only known from the Atlantic Forest of southeastern Brazil, in the states of Espirito Santo, Minas Gerais, Sergipe, Alagoas and Pernambuco (Nogueira et al.  2012, Rocha et al.  2014, Gregorin et al.  2015).		Terrestrial	This is the most threatened Brazilian biome, with the remains of its forest highly fragmented and representing no more than 10% of the original distribution. These humid lowland forest are highly fragmented on eastern Brazil, and only few protected areas extend over this region.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Dryadonycteris		capixaba	Nogueira, Lima, Peracchi, & Simmons	2012	0	Am. Mus. Novitates	########	Espirito Santo Atlantic Forest Bat	None.	Brazil, Espirito Santo, municipality of Linhares, Reserva Natural Vale	E Brazil.	Not listed.	Data Deficient		Dryadonycteris capixaba	1004903	23	Capixaba Nectar-feeding Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	GLOSSOPHAGINAE	CHOERONYCTERINI	Dryadonycteris	NA	capixaba	Nogueira, I. P. Lima, Peracchi, & Simmons	2012	0	Dryadonycteris_capixaba	Nogueira, M. R., Lima, I. P., Peracchi, A. L. & Simmons, N. B. (2012). New genus and species of nectar-feeding bat from the Atlantic Forest of Southeastern Brazil (Chiroptera: Phyllostomidae: Glossophaginae). American Museum Novitates, 3747, 5.	http://digitallibrary.amnh.org/bitstream/handle/2246/6180/N3747.pdf?sequence=1&isAllowed=y	UFRRJ 9667		"Brazil, state of Espírito Santo, municipality of Linhares, Reserva Natural Vale, 19°08'29" S and 40°04'08" W, approximately 60 m elevation."	-19.1414	-40.0689	capixaba Nogueira, I. P. Lima, Peracchi, & Simmons, 2012	recently described genus and species	Nogueira, M. R., Lima, I. P., Peracchi, A. L., & Simmons, N. B. (2012). New genus and species of nectar-feeding bat from the Atlantic Forest of southeastern Brazil (Chiroptera: Phyllostomidae: Glossophaginae). American Museum Novitates, 3747, 1-30.				Brazil	South America	Neotropic	DD	0	0	0	Dryadonycteris_capixaba	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Dryadonycteris_capixaba	1004903	23	Capixaba Nectar-feeding Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Glossophaginae	Choeronycterini	Dryadonycteris	NA	capixaba	Nogueira, I. P. Lima, A. L. Peracchi, & Simmons	0	Dryadonycteris capixaba	Nogueira, M.R., Lima, I.P., Peracchi, A.L. and Simmons, N.B. 2012-06. New genus and species of nectar-feeding bat from the Atlantic Forest of southeastern Brazil (Chiroptera: Phyllostomidae: Glossophaginae). American Museum Novitates 3747:1-30.	https://www.biodiversitylibrary.org/page/59317395	ALP 9667	holotype		"Brazil, state of Espírito Santo, municipality of Linhares, Reserva Natural Vale, 19°08'29" S and 40°04'08" W, approximately 60 m elevation."	-19.1414	-40.0689	recently described genus and species	Nogueira, M. R., Lima, I. P., Peracchi, A. L., & Simmons, N. B. (2012). New genus and species of nectar-feeding bat from the Atlantic Forest of southeastern Brazil (Chiroptera: Phyllostomidae: Glossophaginae). American Museum Novitates, 3747, 1-30.				Brazil	South America	Neotropic	DD	0	0	0	Dryadonycteris_capixaba	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Dryadonycteris		capixaba	Nogueira, Lima, Peracchi & Simmons	2012	0	Am. Mus. Novitates	########	Espirito Santo Atlantic Forest Bat	None.	Brazil, Espirito Santo, municipality of Linhares, Reserva Natural Vale	E Brazil.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/88120233/166613008/' target='_blank'>Data Deficient</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	Choeronycterina	Dryadonycteris capixaba; Dryadonycteris capixaba; Dryadonycteris capixaba; Dryadonycteris capixaba; Dryadonycteris capixaba; capixaba; Dryadonyctére d’Espirito Santo; Capixaba-Dryades-Fledermaus; Driadonicterio de Capixaba; Capixaba Nectar-feeding Bat; Espirito Santo Atlantic Forest Bat; D. capixaba