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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L249 application/vnd.openxmlformats-officedocument.spreadsheetml.sheet Hipposideros camerunensis Hipposideros camerunensis Hipposideros camerunensis Hipposideros camerunensis Hipposideros camerunensis Hipposideros camerunensis Doryrhina camerunensis Doryrhina camerunensis Doryrhina camerunensis Hipposideros camerunensis Doryrhina camerunensis Doryrhina camerunensis Doryrhina camerunensis Doryrhina camerunensis Doryrhina camerunensis [MSW3] cyclops species group.; [HMW] Hipposideros camerunensis Eisentraut, 1956 , near Buea, Cameroon . This species is only known from two isolated areas separated by a large gap, suggesting there may be some taxonomic distinction between the two populations, but this has not yet been investigated. Monotypic.; [batnames2022] cyclops species group. Previously included in Hipposideros .; [MDD2022] moved from Hipposideros to Doryrhina; [IUCN] There is debate as to whether all the records comprise a single species, or whether there might be two or three different species involved. The records from the Democratic Republic of Congo and Kenya could refer to other, possibly undescribed, species.; [batnames2023] cyclops species group. Previously included in Hipposideros .; [MDD2023] moved from Hipposideros to Doryrhina; [MDD2025_2.0] moved from Hipposideros to Doryrhina; [batnames2025_1.7] cyclops species group. Previously included in Hipposideros.; [MDD2025_2.2] moved from Hipposideros to Doryrhina camerunensis There is debate as to whether all the records comprise a single species, or whether there might be two or three different species involved. The records from the Democratic Republic of Congo and Kenya could refer to other, possibly undescribed, species. camerunensis camerunensis camerunensis (Eisentraut, 1956) Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp. Greater cyclops bat Cameroun, E Zaire Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp. Hipposideros camerunensis Cameroun, near Buea. Eisentraut 1956 Zool. Jahrb. Abt. Syst. Oekol. Geogr. Tiere, 84:526. Distribution: Known only from southern Cameroon, eastern Zaire and western Kenya. Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5 Greater cyclops bat Cameroun, E Zaire, Kenya Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp. Eisentraut 1956 Zool. Jahrb. Abt. Syst. Oekol. Geogr. Tiere, 84:526. Cameroon, E Zaire, W Kenya. Cameroon, near Buea. EISENTRAUT 1956 Posterior lateral supplementary leaflet not extending anteriorly beneath anterior nose leaf. Premaxillae wide, enclosing the anterior palatal foramina, making a broad junction with the maxillae. Antorbital foramen relatively small. Size relatively large (forearm length, 74-76 mm). Distribution: Known only from southern Cameroon, eastern Zaire and western Kenya. No subspecies. 64 species H. camerunensis EISENTRAUT 1956 Hipposideros genus Hipposideros camerunensis Posterior lateral supplementary leaflet not extending anteriorly beneath anterior nose leaf. Premaxillae wide, enclosing the anterior palatal foramina, making a broad junction with the maxillae. Antorbital foramen relatively small. Size relatively large (forearm length, 74-76 mm). No subspecies. 34. H. camerunensis EISENTRAUT 1956 [cyclops group]. 34 NA Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu). CHIROPTERA Hipposideridae Hipposideros camerunensis Hipposideros camerunensis Eisentraut 1956 Zool. Jahrb. Abt. Syst. Oekol. Geogr. Tiere 84 526 Cameroon Leaf-nosed Bat Cameroon, near Buea. Cameroon, E Dem. Rep. Congo, W Kenya. IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt). cyclops species group. 03BD87A2C675A207FF29FA7AFD544EC5 Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions 978-84-16728-19-0 hbmw_9_Hipposideridae_210.pdf.imf hash://md5/ff84ffdac676a204fff8ff9affef4346 230 zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C675A207FF29FA7AFD544EC5.xml Doryrhina camerunensis Hipposideridae Doryrhina camerunensis 1956 Phyllorhine du Cameroun @fr | Kamerun-Rundblattnase @de | Doryrina de Camenjn @es | Cameroon Leaf-nosed Bat @en | Greater Roundleaf Bat @en Hipposideros camerunensis Eisentraut, 1956 , near Buea, Cameroon . This species is only known from two isolated areas separated by a large gap, suggesting there may be some taxonomic distinction between the two populations, but this has not yet been investigated. Monotypic. Patchily recorded from SW Cameroon , E DR Congo , S Uganda , and W Kenya . No information. Twelve pregnant females were found in October on Mount Cameroon . The Greater Leaf-nosed Bat roosts in caves and hollow trees. Echolocation call is not known. No information. Classified as Data Deficient on IUCN Red List (as Hipposideros camerunensis ). Due to its limited geographical distribution and its dependence on undisturbed rainforest, the Greater Leaf-nosed Bat may be threatened by deforestation. Eisentraut (1956, 1973) | Happold, M. (2013w) | Hill (1963a) | Rahm (1965) | Schütter et al. (1986) | Thorn & Kerbis Peterhans (2009) https://zenodo.org/record/3749049/files/figure.png 5. Greater Leaf-nosed Bat Doryrhina camerunensis French: Phyllorhine du Cameroun I German: Kamerun-Rundblattnase / Spanish: Doryrina de Camenjn Other common names: Cameroon Leaf-nosed Bat , Greater Roundleaf Bat Taxonomy. Hipposideros camerunensis Eisentraut, 1956 , near Buea, Cameroon . This species is only known from two isolated areas separated by a large gap, suggesting there may be some taxonomic distinction between the two populations, but this has not yet been investigated. Monotypic. Distribution. Patchily recorded from SW Cameroon , E DR Congo , S Uganda , and W Kenya . Descriptive notes. Head—body 92—97 mm , tail 23—45 mm , ear 30-38 mm, hindfoot 18-22 mm, forearm 74—80 mm ; weight 39-53 g. The Greater Leaf-nosed Bat has a distinctive noseleaf with two median clubshaped processes. Pelage is dense, long and soft blackish brown with pale frosting dorsally, and somewhat paler with inconspicuous frosting ventrally. Ears are long, narrow and pointed at tip. A frontal sac is present in both sexes. An anal sac is present between penis and anus in males. Females do not have this sac but fur around vagina has been replaced by long stiff hairs. Dental formula for all species of Doryhina is I 1/2, C 1/1, P 2/2, M 3/3 (x2) = 30. Habitat Inhabits Afromontane and lowland rainforest. It occurs at 1200-1400 m on Mount Cameroon , and at 1600 m above sea level in Kenya , but at much lower altitudes in Uganda and DR Congo . Food and Feeding. No information. Breeding. Twelve pregnant females were found in October on Mount Cameroon . Activity patterns. The Greater Leaf-nosed Bat roosts in caves and hollow trees. Echolocation call is not known. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on IUCN Red List (as Hipposideros camerunensis ). Due to its limited geographical distribution and its dependence on undisturbed rainforest, the Greater Leaf-nosed Bat may be threatened by deforestation. Bibliography. Eisentraut (1956, 1973), Happold, M. (2013w), Hill (1963a), Rahm (1965), Schütter eta/. (1986), Thorn & Kerbis Peterhans (2009). Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022. Hipposideridae Doryrhina camerunensis Doryrhina camerunensis Eisentraut 1956 1 Zool. Jahrb. Abt. Syst. Oekol. Geogr. Tiere 92:46:00 Cameroon Leaf-nosed Bat None. Cameroon, near Buea. Cameroon, E Dem. Rep. Congo, W Kenya. Not listed. Data Deficient as Hipposideros camerunensis cyclops species group. Previously included in Hipposideros . Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023 Doryrhina camerunensis 23 Greater Leaf-nosed Bat Cameroon Leaf-nosed Bat|Greater Roundleaf Bat Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA PTEROPODIFORMES NA NA RHINOLOPHOIDEA HIPPOSIDERIDAE NA NA Doryrhina NA camerunensis Eisentraut 1956 1 Hipposideros_camerunensis Eisentraut, M. (1956). Beitrag zur Chiropteren-Fauna von Kamerun. Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere, 84, 526. SMNS 5194 near Buea, Cameroon. camerunensis (Eisentraut, 1956) moved from Hipposideros to Doryrhina Foley, N. M., Goodman, S. M., Whelan, C. V., Puechmaille, S. J., & Teeling, E. (2017). Towards navigating the Minotaur's labyrinth: cryptic diversity and taxonomic revision within the speciose genus Hipposideros (Hipposideridae). Acta Chiropterologica, 19(1), 1-18. Cameroon|Democratic Republic of the Congo|Uganda|Kenya Africa Afrotropic DD 0 0 0 Doryrhina_camerunensis 0 sciname match Hipposideros_camerunensis 0 IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022]. 10117 Hipposideros camerunensis ANIMALIA CHORDATA MAMMALIA CHIROPTERA HIPPOSIDERIDAE Hipposideros camerunensis Eisentraut, 1956 There is debate as to whether all the records comprise a single species, or whether there might be two or three different species involved. The records from the Democratic Republic of Congo and Kenya could refer to other, possibly undescribed, species. 20000000 Hipposideros camerunensis Data Deficient 2019 2018-08-31 00:00:00 UTC 3.1 English Confirmed as Data Deficient in view of continuing uncertainties as to its taxonomic status, extent of occurrence and ecological requirements. Little known about this species. The Kakamega specimen was captured in 'Intermediate Evergreen Forest' (Schlitter et al. 1986). The original specimens were taken from a hollow tree (Eisentraut 1963). If it is a forest-dependent species, then it is probably suffering from ongoing forest loss. If future taxonomic studies show it to be restricted to Mount Cameroon, then it could be seriously threatened by habitat destruction. Very little known about the species. It is known form only three records in a general region that is not widely surveyed. Unknown On the basis of current knowledge, this species is known only from three localities: Buea in Cameroon (type locality); Shabunda in eastern Democratic Republic of Congo; and at the Kakamega Forest in western Kenya close to Lake Victoria. If it is a single species, then it is likely to be found in future in intervening locations. Its elevational range is from sea-level to 500 m, and probably higher. Terrestrial None conservation measures are currently in place. It is not known from any protected areas. Taxonomic studies are urgently needed to determine whether or not the three recorded localities all refer to the same species. Afrotropical FALSE FALSE Global Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 Hipposideridae Doryrhina camerunensis Eisentraut 1956 1 Zool. Jahrb. Abt. Syst. Oekol. Geogr. Tiere 92:46:00 Cameroon Leaf-nosed Bat None. Cameroon, near Buea. Cameroon, E Dem. Rep. Congo, W Kenya. Not listed. Data Deficient as Hipposideros camerunensis cyclops species group. Previously included in Hipposideros . Doryrhina camerunensis 1004566 23 Greater Leaf-nosed Bat Cameroon Leaf-nosed Bat|Greater Roundleaf Bat Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA PTEROPODIFORMES NA NA RHINOLOPHOIDEA Hipposideridae NA NA Doryrhina NA camerunensis Eisentraut 1956 1 Hipposideros_camerunensis Eisentraut, M. (1956). Beitrag zur Chiropteren-Fauna von Kamerun. Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere, 84, 526. SMNS 5194 near Buea, Cameroon. camerunensis (Eisentraut, 1956) moved from Hipposideros to Doryrhina Foley, N. M., Goodman, S. M., Whelan, C. V., Puechmaille, S. J., & Teeling, E. (2017). Towards navigating the Minotaur's labyrinth: cryptic diversity and taxonomic revision within the speciose genus Hipposideros (Hipposideridae). Acta Chiropterologica, 19(1), 1-18. Cameroon|Democratic Republic of the Congo|Uganda|Kenya Africa Afrotropic DD 0 0 0 Doryrhina_camerunensis 0 oldname match Hipposideros_camerunensis 0 Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393 Doryrhina_camerunensis 1004566 23 Greater Roundleaf Bat Cameroon Leaf-nosed Bat|Greater Roundleaf Bat|Greater Leaf-nosed Bat Theria Placentalia Boreoeutheria Laurasiatheria Chiroptera Yinpterochiroptera NA NA Rhinolophoidea Hipposideridae NA NA Doryrhina NA camerunensis Eisentraut 1 Hipposideros camerunensis Eisentraut, M. 1956. Beitrag zur Chiropteren-Fauna von Kamerun (Westafrika). Zoologische Jahrbücher 85:505-540. SMNS 5194 holotype near Buea, Cameroon. moved from Hipposideros to Doryrhina Foley, N. M., Goodman, S. M., Whelan, C. V., Puechmaille, S. J., & Teeling, E. (2017). Towards navigating the Minotaur's labyrinth: cryptic diversity and taxonomic revision within the speciose genus Hipposideros (Hipposideridae). Acta Chiropterologica, 19(1), 1-18. Cameroon|Democratic Republic of the Congo|Uganda|Kenya Africa Afrotropic DD 0 0 0 Doryrhina_camerunensis 0 oldname match Hipposideros_camerunensis 0 Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586 Hipposideridae Doryrhina camerunensis Eisentraut 1956 1 Zool. Jahrb. Abt. Syst. Oekol. Geogr. Tiere 92:46:00 Cameroon Leaf-nosed Bat None. Cameroon, near Buea. Cameroon, E Dem. Rep. Congo, W Kenya. Not Listed Data Deficient cyclops species group. Previously included in Hipposideros. Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505 NA Hipposideros camerunensis; Doryrhina camerunensis; Doryrhina camerunensis; Doryrhina camerunensis; Hipposideros camerunensis; Doryrhina camerunensis; camerunensis; Phyllorhine du Cameroun; Kamerun-Rundblattnase; Doryrina de Camenjn; Cameroon Leaf-nosed Bat; Greater Roundleaf Bat; Greater Leaf-nosed Bat; Cameroon Leaf-nosed Bat; Greater Roundleaf Bat; Cameroon Leaf-nosed Bat; Cameroon Leaf-nosed Bat; D. camerunensis