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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L248	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Dobsonia viridis	Dobsonia viridis	Dobsonia viridis	Dobsonia viridis	Dobsonia viridis	Dobsonia viridis	Dobsonia viridis	Dobsonia viridis	Dobsonia viridis	Dobsonia viridis	Dobsonia viridis	Dobsonia viridis	Dobsonia viridis	Dobsonia viridis	Dobsonia viridis		[MSW2] Does not include chapmani (see Bergmans, 1978:12). Includes crenulata (see Hill, 1991:171); but see Bergmans and Rozendaal (1988:33).; [MSW3] viridis species group. Does not include chapmani; see Bergmans (1978). Does not include crenulata; see Bergmans and Rozendaal (1988). See also Hill (1983) and Flannery (1995b).; [HMW] Cephalotes viridis Heude, 1896 , Kai Islands, Maluku Islands , Indonesia . Records from southern Sulawesi , Buton, and Kabaena islands are assigned here to D. crenulata (previously treated by some authors as subspecies of a more widespread D. viridis ). Monotypic.; [batnames2022]  viridis species group. Does not include chapmani ; see Bergmans (1978). Does not include crenulata ; see Bergmans and Rozendaal (1988). See also Hill (1983) and Flannery (1995).; [batnames2023]  viridis species group. Does not include chapmani ; see Bergmans (1978). Does not include crenulata ; see Bergmans and Rozendaal (1988). See also Hill (1983) and Flannery (1995).; [batnames2025_1.7] viridis species group. Does not include chapmani; see Bergmans (1978). Does not include crenulata; see Bergmans and Rozendaal (1988). See also Hill (1983) and Flannery (1995).			crenulata			crenulata, umbrosa.	viridis, crenulata		umbrosa			viridis 	viridis - umbrosa	viridis, umbrosus		viridis 	viridis - umbrosa	viridis, umbrosus	viridis, umbrosa	viridis 	viridis - umbrosa	viridis (Heude, 1897)|umbrosa O. Thomas, 1910		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Greenish naked-backed bat	Negros I, Philippines; S Moluccas	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Dobsonia viridis	Indonesia, Molucca Isis., Kei Isis.	Heude	1896	Mem. Hist. Nat. Emp. Chin., 3:176.	Distribution: Known from Celebes, the Moluccas, and nearby small islands, including Misol in the west Papuan islands.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	(? crenulata)	Greenish naked-backed bat Sulawesi; Moluccas; ref. 4.12	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Heude	1896	Mem. Hist. Nat. Emp. Chin., 3:176.	Does not include chapmani (see Bergmans, 1978:12). Includes crenulata (see Hill, 1991:171); but see Bergmans and Rozendaal (1988:33).	Sulawesi, Muluccas, and nearby islands.	Indonesia, Molucca Isis, Kei Isis.		HEUDE	1896	A well-marked antero-internal basal ledge or cusp on the last upper and lower premolars as well as the first upper and lower molars. Notches gen erally present on longitudinal ridges of molars and premolars. Size medium to large for group (fore arm length, 108-129 mm; total length of skull, 45-54 mm). Rostrum relatively lighter (lacrimal breadth, 11.5-12.8 mm). Teeth variable in size (maxillary toothrow length, 18-21 mm). Interor bital width medium (9-10 mm).	Distribution: Known from Celebes, the Moluccas, and nearby small islands, including Misol in the west Papuan islands.	Two currently recognized subspecies:	D. v. viridis (Celebes, central Moluccas, Misol); D. v. crenulata (Sanghir and Togian islands, near Celebes, to the northern Moluccas, Halmahera group).	29	species	D. viridis	HEUDE	1896	Dobsonia	genus	Dobsonia viridis				A well-marked antero-internal basal ledge or cusp on the last upper and lower premolars as well as the first upper and lower molars. Notches gen erally present on longitudinal ridges of molars and premolars. Size medium to large for group (fore arm length, 108-129 mm; total length of skull, 45-54 mm). Rostrum relatively lighter (lacrimal breadth, 11.5-12.8 mm). Teeth variable in size (maxillary toothrow length, 18-21 mm). Interor bital width medium (9-10 mm).	Two currently recognized subspecies:		8. D. viridis (HEUDE 1896) [viridis group].	8	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Dobsonia viridis	Dobsonia		viridis	Heude	y	1896		Mem. Hist. Nat. Emp. Chin.	3		176		Greenish Naked-backed Fruit Bat	Indonesia, Maluku (Moluccas), Kai Isls.	C and S Moluccas including Seram, Ambon, and Buru; Banda, and Kai Isls (Indonesia). A closely related but undescribed species occurs in the Tanimbar Isls (K. Helgen, pers. comm.).	IUCN/SSC Action Plan (1992) – Not Threatened. IUCN 2003 – Lower Risk (lc).	umbrosa Thomas, 1910.	viridis species group. Does not include chapmani; see Bergmans (1978). Does not include crenulata; see Bergmans and Rozendaal (1988). See also Hill (1983) and Flannery (1995b).	03AD87FAFFF1F61F896B386CF9B5F984	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	111	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFF1F61F896B386CF9B5F984.xml	Dobsonia viridis	Pteropodidae	Dobsonia	viridis	Heude	1897	Roussette verdatre @fr | Griner Nacktrickenflughund @de | Dobsonia verdoso @es	Cephalotes viridis Heude, 1896 , Kai Islands, Maluku Islands , Indonesia . Records from southern Sulawesi , Buton, and Kabaena islands are assigned here to D. crenulata (previously treated by some authors as subspecies of a more widespread D. viridis ). Monotypic.	Sula Is (Mangole), C Moluccas (Buru, Seram, and Ambon ), Banda Is (Banda Neira), and Kai Is; there are specimens that likely represent this species or closely related undescribed species from the TanimbarIs.	Head-body 154-— 176 mm , tail 25-39 mm , ear 21-25 mm , forearm 109-118 mm ; weight 173-229 g . Fur of the Greenish Naked-backed Fruit Bat is greenish. Wings attach along midline of back with underlying fur. Index claw (second digit of wing) is absent. Nostrils are slightly tubular, and each flares outward from centerline. Eyes are large, with brown irises. Short tail is free of narrow uropatagium that runs along legs to short stiff calcars. Molars are more cuspidate than in other species of Dobsonia . Anterointernal corner of M| is sharply marked off by distinct cusp or ledge. There are notched longitudinal ridges on upper and lower last premolars and first molars.	[Lowland and hill tropical moist forests, disturbed forests, coconut and cocoa plantations, and gardens from sea level up to elevations of ¢. 1000 m .	The Greenish Naked-backed Fruit Bat is frugivorous. It presumably feeds heavily on native figs and commonly forages in orchards and gardens.	Pregnant females and non-volant young Greenish Naked-backed Fruit Bats have been observed in November on Mangole.	Greenish Naked-backed Fruit Bats are nocturnal and roost in caves and trees.	No information.	Classified as Least Concern on The IUCN Red List. The Greenish Naked-backed Fruit Bat is abundant throughout its distribution, and populations appear to be stable. It also appears to be well adapted to disturbed habitats. Hunting for local consumption is a threat in some areas, but main threat is continuing decline in extent and quality of lowland and hill forests.	Bergmans (1978b) | Bergmans & Rozendaal (1988) | Flannery (1995a) | Mubarok et al. (2018) | Tsang (2016b)		90. Greenish Naked-backed Fruit Bat Dobsonia viridis French: Roussette verdatre / German: Griner Nacktrickenflughund / Spanish: Dobsonia verdoso Taxonomy. Cephalotes viridis Heude, 1896 , Kai Islands, Maluku Islands , Indonesia . Records from southern Sulawesi , Buton, and Kabaena islands are assigned here to D. crenulata (previously treated by some authors as subspecies of a more widespread D. viridis ). Monotypic. Distribution. Sula Is (Mangole), C Moluccas (Buru, Seram, and Ambon ), Banda Is (Banda Neira), and Kai Is; there are specimens that likely represent this species or closely related undescribed species from the TanimbarIs. Descriptive notes. Head-body 154-— 176 mm , tail 25-39 mm , ear 21-25 mm , forearm 109-118 mm ; weight 173-229 g . Fur of the Greenish Naked-backed Fruit Bat is greenish. Wings attach along midline of back with underlying fur. Index claw (second digit of wing) is absent. Nostrils are slightly tubular, and each flares outward from centerline. Eyes are large, with brown irises. Short tail is free of narrow uropatagium that runs along legs to short stiff calcars. Molars are more cuspidate than in other species of Dobsonia . Anterointernal corner of M| is sharply marked off by distinct cusp or ledge. There are notched longitudinal ridges on upper and lower last premolars and first molars. Habitat. [Lowland and hill tropical moist forests, disturbed forests, coconut and cocoa plantations, and gardens from sea level up to elevations of ¢. 1000 m . Food and Feeding. The Greenish Naked-backed Fruit Bat is frugivorous. It presumably feeds heavily on native figs and commonly forages in orchards and gardens. Breeding. Pregnant females and non-volant young Greenish Naked-backed Fruit Bats have been observed in November on Mangole. Activity patterns. Greenish Naked-backed Fruit Bats are nocturnal and roost in caves and trees. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Greenish Naked-backed Fruit Bat is abundant throughout its distribution, and populations appear to be stable. It also appears to be well adapted to disturbed habitats. Hunting for local consumption is a threat in some areas, but main threat is continuing decline in extent and quality of lowland and hill forests. Bibliography. Bergmans (1978b), Bergmans & Rozendaal (1988), Flannery (1995a), Mubarok et al. (2018), Tsang (2016b).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Dobsonia viridis	Dobsonia		viridis	Heude	1896	1	Mem. Hist. Nat. Emp. Chin.	0.2472	Greenish Naked-backed Fruit Bat	 umbrosa Thomas, 1910.	Indonesia, Maluku (Moluccas), Kai Isls.	C and S Moluccas including Seram, Ambon, and Buru; Banda, and Kai Isls (Indonesia). A closely related but undescribed species occurs in the Tanimbar Isls (K. Helgen, pers. comm.).	Not listed.	Least Concern	 viridis species group. Does not include chapmani ; see Bergmans (1978). Does not include crenulata ; see Bergmans and Rozendaal (1988). See also Hill (1983) and Flannery (1995).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Dobsonia viridis	23	Greenish Naked-backed Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	HARPYIONYCTERINAE	DOBSONIINI	Dobsonia	NA	viridis	Heude	1896	1						Kai Islands, Maluku Islands, Indonesia.			viridis (Heude, 1896)|umbrosus O. Thomas, 1910	NA	NA	Indonesia	Oceania	Australasia/Oceania	LC	0	0	0	Dobsonia_viridis	0	sciname match	Dobsonia_viridis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	6780	Dobsonia viridis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Dobsonia	viridis	(Heude, 1896)		20000000	Dobsonia viridis	Least Concern		2016	2016-01-18 00:00:00 UTC	3.1	English	Listed as Least Concern as the species is considered common and abundant throughout its range, and populations are stable. The species also appears to be well adapted to disturbed habitat.	It is common in disturbed forest, coconut plantations, and gardens. It roosts in caves and trees. It can commonly be found foraging in orchards.	There are no major threats to the species throughout its range. Hunting, especially in caves (Boadi pers. comm.), represents a localised threat for some populations.	This species is common and widespread throughout its range.	Stable	This species is found on the Kai Islands, central and south Moluccas including Seram, Ambon, and Buru, Banda Islands, all in Indonesia. A closely related but undescribed species occurs in the Tanimbar Islands (K. Helgen pers. comm.).		Terrestrial	This species occurs in some protected areas throughout its range, such as Manusela National Park on Seram.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Dobsonia		viridis	Heude	1896	1	Mem. Hist. Nat. Emp. Chin.	0.247222	Greenish Naked-backed Fruit Bat	 umbrosa Thomas, 1910.	Indonesia, Maluku (Moluccas), Kai Isls.	C and S Moluccas including Seram, Ambon, and Buru; Banda, and Kai Isls (Indonesia). A closely related but undescribed species occurs in the Tanimbar Isls (K. Helgen, pers. comm.).	Not listed.	Least Concern	 viridis species group. Does not include chapmani ; see Bergmans (1978). Does not include crenulata ; see Bergmans and Rozendaal (1988). See also Hill (1983) and Flannery (1995).	Dobsonia viridis	1004403	23	Greenish Naked-backed Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	HARPYIONYCTERINAE	DOBSONIINI	Dobsonia	NA	viridis	Heude	1896	1						Kai Islands, Maluku Islands, Indonesia.			viridis (Heude, 1896)|umbrosus O. Thomas, 1910	NA	NA				Indonesia	Oceania	Australasia/Oceania	LC	0	0	0	Dobsonia_viridis	0	sciname match	Dobsonia_viridis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Dobsonia_viridis	1004403	23	Greenish Naked-backed Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Harpyionycterinae	Dobsoniini	Dobsonia	NA	viridis	Heude	1	Cephalotes viridis	Heude, P.M. 1897. Études odontologiques. Seconde partie. Mammifères carnassiers. Mémoires concernant l'histoire naturelle de l'empire chinois 3(4):157-188.					Kai Islands, Maluku Islands, Indonesia.			NA	NA				Indonesia	Oceania (Continent)	Australasia	LC	0	0	0	Dobsonia_viridis	0	sciname match	Dobsonia_viridis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Dobsonia		viridis	Heude	1896	1	Mem. Hist. Nat. Emp. Chin.	0.247222	Greenish Naked-backed Fruit Bat	umbrosa Thomas, 1910.	Indonesia, Maluku (Moluccas), Kai Isls.	C and S Moluccas including Seram, Ambon, and Buru; Banda, and Kai Isls (Indonesia). A closely related but undescribed species occurs in the Tanimbar Isls (K. Helgen, pers. comm.).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/6780/22033412/' target='_blank'>Least Concern</a>	viridis species group. Does not include chapmani; see Bergmans (1978). Does not include crenulata; see Bergmans and Rozendaal (1988). See also Hill (1983) and Flannery (1995).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Dobsonia viridis; Dobsonia viridis; Dobsonia viridis; Dobsonia viridis; Dobsonia viridis; Dobsonia viridis; umbrosa; umbrosa; viridis; umbrosus; Roussette verdatre; Griner Nacktrickenflughund; Dobsonia verdoso; Greenish Naked-backed Fruit Bat; Greenish Naked-backed Fruit Bat; Greenish Naked-backed Fruit Bat; D. viridis