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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L247	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Dobsonia praedatrix	Dobsonia praedatrix	Dobsonia praedatrix	Dobsonia praedatrix	Dobsonia praedatrix	Dobsonia praedatrix	Dobsonia praedatrix	Dobsonia praedatrix	Dobsonia praedatrix	Dobsonia praedatrix	Dobsonia praedatrix	Dobsonia praedatrix	Dobsonia praedatrix	Dobsonia praedatrix	Dobsonia praedatrix		[MSW3] viridis species group; see discussion in Bergmans (1978) and Hill (1983). See also Flannery (1995b) and Bonaccorso (1998).; [HMW] Dobsonia praedatrix K. Andersen, 1909 , “Duke of York group,” Bismarck Archipelago , Papua New Guinea . This species is monotypic.; [batnames2022]  viridis species group; see discussion in Bergmans (1978) and Hill (1983). See also Flannery (1995b) and Bonaccorso (1998).; [batnames2023]  viridis species group; see discussion in Bergmans (1978) and Hill (1983). See also Flannery (1995b) and Bonaccorso (1998).; [batnames2025_1.7] viridis species group; see discussion in Bergmans (1978) and Hill (1983). See also Flannery (1995b) and Bonaccorso (1998).														praedatrix				praedatrix	praedatrix			praedatrix Andersen, 1909		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	New Britain naked-backed	Bismarck Archipelago bat	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Dobsonia praedatrix	Papua New Guinea, Bismarck Arch., "Duke of York group".	K. Andersen	1909	Ann. Mag. Nat. Hist., ser. 8, 4:532.	Distribution: Confined to the Bismarcks.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		New Britain naked-backed Bismarck Archipelago bat	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	K. Andersen	1909	Ann. Mag. Nat. Hist., ser. 8, 4:532.		Bismarck Arch.	Papua New Guinea, Bismarck Arch., "Duke of York group".		ANDERSEN	1900	Size medium for group (forearm length, 111-122mm; total length of skull 49-52mm). Rostrum relatively heavier (lacrimal breadth, 12.8-13.8 mm). Teeth relatively large (maxillary toothrow length, 18-21 mm). Interorbital width great (10.2-10.5 mm).	Distribution: Confined to the Bismarcks.	No subspecies.		29	species	D. praedatrix	ANDERSEN	1900	Dobsonia	genus	Dobsonia praedatrix				Size medium for group (forearm length, 111-122mm; total length of skull 49-52mm). Rostrum relatively heavier (lacrimal breadth, 12.8-13.8 mm). Teeth relatively large (maxillary toothrow length, 18-21 mm). Interorbital width great (10.2-10.5 mm).	No subspecies.		10. D. praedatrix ANDERSEN 1900 [viridis group].	10	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Dobsonia praedatrix	Dobsonia		praedatrix	K. Andersen		1909		Ann. Mag. Nat. Hist., ser. 8	4		532		New Britain Naked-backed Fruit Bat	Papua New Guinea, Bismarck Arch., “Duke of York group”.	Bismarck Arch. (Papua New Guinea).	IUCN/SSC Action Plan (1992) – No Data. IUCN 2003 – Lower Risk (nt).		viridis species group; see discussion in Bergmans (1978) and Hill (1983). See also Flannery (1995b) and Bonaccorso (1998).	03AD87FAFFF2F61D8CB43E03F859FBD4	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	112	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFF2F61D8CB43E03F859FBD4.xml	Dobsonia praedatrix	Pteropodidae	Dobsonia	praedatrix	K. Andersen	1909	Roussette de Nouvelle-Bretagne @fr | Neubritannien-Nacktriickenflughund @de | Dobsonia de @es | Bismarck Bare-backed Fruit Bat @en | New Britain Bare-backed Fruit Bat @en	Dobsonia praedatrix K. Andersen, 1909 , “Duke of York group,” Bismarck Archipelago , Papua New Guinea . This species is monotypic.	Bismarck Archipelago including Umboi I and nearbyIs groups.	Head-body 116- 180 mm , tail 21-37 mm , car 21-28 mm , hindfoot 22-33 mm , forearm 101- - 125 mm ; weight 141-260 g . Pelage on crown ofhead and face of the NewBritain Naked-backed Fruit Bat is blackish brown; shoulders and upper back are paler brown. Ventral fur is pale brown, with yellow-green wash. Greenish color, which fades to a dull gray/tan in museum specimens, is probably attributable to photosynthetic bluegreen algae based on iodine tests that reveal starch-containing globs coating hairs. Wing membranes meet along spine with underlying fur. Ears are long and slender compared with other species of Dobsonia . Nostrils are slightly tubular, and each flares outward from centerline. Claws are bicolored, brown at bases and paler tan at tips. Index claw (second digit of wing) is absent. Skull has well-developed sagittal and nuchal crests. The New Britain Naked-backed Fruit Bat has exceptionally large pineal organ, which is associated with providing seasonal cues for production of hormones controlling reproductive timing. Upperincisors are unilobed, and molars have weakly developed cusps. Chromosomal complement has 2n = 28 and FN = 68.	Lowland primary and secondary tropical moist forests, fruit and coconut plantations, and rural gardens from sea level up to elevations of ¢. 300 m .	The New Britain Naked-backed Fruit Batis frugivorous. It can often be found foraging in the same fruiting trees as the much larger Andersen’s Nakedbacked Fruit Bat ( D. anderseni ). Diet includes native figs ( Ficus , Moraceae ), garden fruits such as papaya, plantation grown cashews, and unidentified forest canopy flowers.	Pregnant New Britain Naked-backed Fruit Bats have been observed in January and July at Kerevat, New Britain , suggesting two annual birth peaks. July observations of pregnancy on New Ireland also exist. Dependent young have been observed in January-February at various locations. Nearly full-grown individuals have been observed in January and July.	The New Britain Naked-backed Fruit Bat is nocturnal; flight activity begins well after sunset in complete darkness. It commonly roosts in foliage, often coconut plantations among dead palm fronds, and rarely limestone caves.	Roosting group size tends to be c.10-15 individuals, although solitary individuals are encountered. The New Britain Naked-backed Fruit Bat is mainly a foliage-roosting species that responds to commonly encountered ambient daily temperatures (25-27°C) of foliage roosts, having basal metabolic rates and thermal conductance near mass-specific mammalian averages, and by maintaining homeothermy by increasing metabolism to regulate constant body temperature in cool shade temperatures that are slightly below thermoneutral zone. Maintaining alertness with high body temperature while roosting in tree canopies with potential predators including arboreal snakes and aerial raptors is critical to permit instantaneous flight. Although no studies on predation exist, brown tree snakes (Boiga irregularis) likely prey on New Britain Naked-backed Fruit Bats because they are known to feed on Marianas Flying Foxes ( Pteropus mariannus ) of similarsize.	Classified as Least Concern on The IUCN Red List. The New Britain Naked-backed Fruit Bat has a large and presumably stable population and large distribution occurring on many islands in the Bismarck Archipelago and surrounding region. It tolerates some degree of habitat degradation and is resilient to traditional small-scale agriculture in which small forest patches are maintained by clearing and abandoned gardens are allowed to return to secondary forest.	Bhatnagar et al. (1990) | Bonaccorso (1998) | Flannery (1995b) | Helgen, Bonaccorso & Hamilton (2008) | McNab & Bonaccorso (2001) | Smith & Hood (1981)		94. New Britain Naked-backed Fruit Bat Dobsonia praedatrix French: Roussette de Nouvelle-Bretagne / German: Neubritannien-Nacktriickenflughund / Spanish: Dobsonia de Nueva Bretana Other common names: Bismarck Bare-backed Fruit Bat , New Britain Bare-backed Fruit Bat Taxonomy. Dobsonia praedatrix K. Andersen, 1909 , “Duke of York group,” Bismarck Archipelago , Papua New Guinea . This species is monotypic. Distribution. Bismarck Archipelago including Umboi I and nearbyIs groups. Descriptive notes. Head-body 116- 180 mm , tail 21-37 mm , car 21-28 mm , hindfoot 22-33 mm , forearm 101- - 125 mm ; weight 141-260 g . Pelage on crown ofhead and face of the NewBritain Naked-backed Fruit Bat is blackish brown; shoulders and upper back are paler brown. Ventral fur is pale brown, with yellow-green wash. Greenish color, which fades to a dull gray/tan in museum specimens, is probably attributable to photosynthetic bluegreen algae based on iodine tests that reveal starch-containing globs coating hairs. Wing membranes meet along spine with underlying fur. Ears are long and slender compared with other species of Dobsonia . Nostrils are slightly tubular, and each flares outward from centerline. Claws are bicolored, brown at bases and paler tan at tips. Index claw (second digit of wing) is absent. Skull has well-developed sagittal and nuchal crests. The New Britain Naked-backed Fruit Bat has exceptionally large pineal organ, which is associated with providing seasonal cues for production of hormones controlling reproductive timing. Upperincisors are unilobed, and molars have weakly developed cusps. Chromosomal complement has 2n = 28 and FN = 68. Habitat. Lowland primary and secondary tropical moist forests, fruit and coconut plantations, and rural gardens from sea level up to elevations of ¢. 300 m . Food and Feeding. The New Britain Naked-backed Fruit Batis frugivorous. It can often be found foraging in the same fruiting trees as the much larger Andersen’s Nakedbacked Fruit Bat ( D. anderseni ). Diet includes native figs ( Ficus , Moraceae ), garden fruits such as papaya, plantation grown cashews, and unidentified forest canopy flowers. Breeding. Pregnant New Britain Naked-backed Fruit Bats have been observed in January and July at Kerevat, New Britain , suggesting two annual birth peaks. July observations of pregnancy on New Ireland also exist. Dependent young have been observed in January-February at various locations. Nearly full-grown individuals have been observed in January and July. Activity patterns. The New Britain Naked-backed Fruit Bat is nocturnal; flight activity begins well after sunset in complete darkness. It commonly roosts in foliage, often coconut plantations among dead palm fronds, and rarely limestone caves. Movements, Home range and Social organization. Roosting group size tends to be c.10-15 individuals, although solitary individuals are encountered. The New Britain Naked-backed Fruit Bat is mainly a foliage-roosting species that responds to commonly encountered ambient daily temperatures (25-27°C) of foliage roosts, having basal metabolic rates and thermal conductance near mass-specific mammalian averages, and by maintaining homeothermy by increasing metabolism to regulate constant body temperature in cool shade temperatures that are slightly below thermoneutral zone. Maintaining alertness with high body temperature while roosting in tree canopies with potential predators including arboreal snakes and aerial raptors is critical to permit instantaneous flight. Although no studies on predation exist, brown tree snakes (Boiga irregularis) likely prey on New Britain Naked-backed Fruit Bats because they are known to feed on Marianas Flying Foxes ( Pteropus mariannus ) of similarsize. Status and Conservation. Classified as Least Concern on The IUCN Red List. The New Britain Naked-backed Fruit Bat has a large and presumably stable population and large distribution occurring on many islands in the Bismarck Archipelago and surrounding region. It tolerates some degree of habitat degradation and is resilient to traditional small-scale agriculture in which small forest patches are maintained by clearing and abandoned gardens are allowed to return to secondary forest. Bibliography. Bhatnagar et al. (1990), Bonaccorso (1998), Flannery (1995b), Helgen, Bonaccorso & Hamilton (2008), McNab & Bonaccorso (2001), Smith & Hood (1981).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Dobsonia praedatrix	Dobsonia		praedatrix	K. Andersen	1909	0	Ann. Mag. Nat. Hist.	ser. 8, 4: 532	New Britain Naked-backed Fruit Bat	None.	Papua New Guinea, Bismarck Arch., “Duke of York group”.	Bismarck Arch. (Papua New Guinea).	Not listed.	Least Concern	 viridis species group; see discussion in Bergmans (1978) and Hill (1983). See also Flannery (1995b) and Bonaccorso (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Dobsonia praedatrix	23	New Britain Naked-backed Fruit Bat	Bismarck Bare-backed Fruit Bat|New Britain Bare-backed Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	HARPYIONYCTERINAE	DOBSONIINI	Dobsonia	NA	praedatrix	K. Andersen	1909	0						"Duke of York group," Bismarck Archipelago, Papua New Guinea.			praedatrix K. Andersen, 1909	NA	NA	Papua New Guinea	Oceania	Australasia/Oceania	LC	0	0	0	Dobsonia_praedatrix	0	sciname match	Dobsonia_praedatrix	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	6777	Dobsonia praedatrix	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Dobsonia	praedatrix	K. Andersen, 1909		20000000	Dobsonia praedatrix	Least Concern		2020	2019-05-10 00:00:00 UTC	3.1	English	Listed as Least Concern due to its large population and tolerance of some degree of habitat degradation, and because it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category. If the trend towards forest conversion for oil palms and re-entry logging on less than 35 year rotations continues, this species may become threatened.	This species is found in lowland primary and secondary tropical moist forest, plantations, and subsistence gardens (Bonaccorso 1998). It normally roosts in small groups, or as solitary individuals, in foliage. It has been recorded roosting in coconut plantations, and there are some records of animals roosting in caves including a maternity colony (Bonaccorso 1998, Flannery 1995). It forages in the canopy for fruit and flowers. Females give birth to a single young and at least in New Britain, birth peaks may occur in January and July (Bonaccorso 1998).	Conversion of lowland forest habitats to oil palm plantation is a major threat. May be threatened by deforestation and forest degradation, for the timber trade, but it is reportedly very resilient as long as there is not complete forest conversion (i.e. if small forest patches are maintained during clearing).  Deforestation and forest degradation are on-going in the Southern and Northern Bismarck Island Arc bioregions with by far the largest percentage change (45.5% and 42.9% of forests) recorded in PNG forests between 1972 and 2002, largely in the lowlands (Shearman and Bryan 2011). New Britain and New Ireland continued to record high rates of deforestation and degradation from logging between 2002 and 2014 (Bryan et al. 2015). A better understanding of the species habitat requirements is required to determine the threat that on-going deforestation and forest degradation poses.	The population trend is unknown. The Bismarck Bare-backed Fruit-bat can be locally common (Bonaccorso 1998, Anthony et al. 2001).	Unknown	The Bismarck Bare-backed Fruit Bat is restricted to the Bismarck Archipelago of Papua New Guinea. It has been recorded from the islands of Duke of York, Dyaul, Mioko, New Britain, New Ireland, Ulu, Umboi (and nearby small islands), and the New Hanover Islands (Flannery 1995, Bonaccorso 1998). It is found from sea level to 300 m asl.	Unlikely that the species is specifically hunted for food given the species small body size, that it often roosts cryptically amongst foliage, and roosting groups are generally small. It may be killed if opportunistically encountered.	Terrestrial	It is not known if the species is present in any protected areas, and even if it is, the level of protection this affords the species is unclear. For example, the majority of the protected areas within the species range were either logged or converted to subsistence gardens or plantations between 1972 and 2002 (derived from supplementary information in Shearman and Bryan 2011).	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Dobsonia		praedatrix	K. Andersen	1909	0	Ann. Mag. Nat. Hist.	ser. 8, 4: 532	New Britain Naked-backed Fruit Bat	None.	Papua New Guinea, Bismarck Arch., “Duke of York group”.	Bismarck Arch. (Papua New Guinea).	Not listed.	Least Concern	 viridis species group; see discussion in Bergmans (1978) and Hill (1983). See also Flannery (1995b) and Bonaccorso (1998).	Dobsonia praedatrix	1004402	23	New Britain Naked-backed Fruit Bat	Bismarck Bare-backed Fruit Bat|New Britain Bare-backed Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	HARPYIONYCTERINAE	DOBSONIINI	Dobsonia	NA	praedatrix	K. Andersen	1909	0						"Duke of York group," Bismarck Archipelago, Papua New Guinea.			praedatrix K. Andersen, 1909	NA	NA				Papua New Guinea	Oceania	Australasia/Oceania	LC	0	0	0	Dobsonia_praedatrix	0	sciname match	Dobsonia_praedatrix	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Dobsonia_praedatrix	1004402	23	New Britain Naked-backed Fruit Bat	Bismarck Bare-backed Fruit Bat|New Britain Bare-backed Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Harpyionycterinae	Dobsoniini	Dobsonia	NA	praedatrix	Andersen	0	Dobsonia praedatrix	Andersen, K.C. 1909-12-01. On the fruit-bats of the genus _Dobsonia_. Annals and Magazine of Natural History (8)4(24):528-536.	https://www.biodiversitylibrary.org/page/22097913	BMNH:Mamm:1877.7.18.5	holotype	https://data.nhm.ac.uk/object/82ffbf22-7926-4acd-9109-eaecce2b541f	"Duke of York group," Bismarck Archipelago, Papua New Guinea.			NA	NA				Papua New Guinea	Oceania (Continent)	Australasia	LC	0	0	0	Dobsonia_praedatrix	0	sciname match	Dobsonia_praedatrix	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Dobsonia		praedatrix	K. Andersen	1909	0	Ann. Mag. Nat. Hist.	ser. 8, 4: 532	New Britain Naked-backed Fruit Bat	None.	Papua New Guinea, Bismarck Arch., “Duke of York group”.	Bismarck Arch. (Papua New Guinea).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/6777/22033332/' target='_blank'>Least Concern</a>	viridis species group; see discussion in Bergmans (1978) and Hill (1983). See also Flannery (1995b) and Bonaccorso (1998).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Dobsonia praedatrix; Dobsonia praedatrix; Dobsonia praedatrix; Dobsonia praedatrix; Dobsonia praedatrix; Dobsonia praedatrix; praedatrix; Roussette de Nouvelle-Bretagne; Neubritannien-Nacktriickenflughund; Dobsonia de; Bismarck Bare-backed Fruit Bat; New Britain Bare-backed Fruit Bat; New Britain Naked-backed Fruit Bat; Bismarck Bare-backed Fruit Bat; New Britain Bare-backed Fruit Bat; New Britain Naked-backed Fruit Bat; New Britain Naked-backed Fruit Bat; D. praedatrix