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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L246	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Dobsonia peroni	Dobsonia peroni	Dobsonia peronii	Dobsonia peroni	Dobsonia peroni	Dobsonia peronii	Dobsonia peronii	Dobsonia peronii	Dobsonia peronii	Dobsonia peronii	Dobsonia peronii	Dobsonia peronii	Dobsonia peronii	Dobsonia peronii	Dobsonia peronii		[MSW2] Reviewed by Bergmans (1978).; [MSW3] peronii species group. Reviewed by Bergmans (1978) and Kitchener et al. (1997a). Sometimes spelled peroni (e.g., Andersen, 1912; Koopman, 1993).; [HMW] Cephalotes peronii E. Geoffroy Saint-Hilaire, 1810 , “L’ile de Timor,” Lesser Sunda Islands , Indonesia . "Two subspecies are recognized.; [batnames2022]  peronii species group. Reviewed by Bergmans (1978) and Kitchener et al . (1997). Sometimes spelled peroni (e.g., Andersen, 1912; Koopman,1993).; [batnames2023]  peronii species group. Reviewed by Bergmans (1978) and Kitchener et al . (1997). Sometimes spelled peroni (e.g., Andersen, 1912; Koopman,1993).; [batnames2025_1.7] peronii species group. Reviewed by Bergmans (1978) and Kitchener et al. (1997). Sometimes spelled peroni (e.g., Andersen, 1912; Koopman,1993).						grandis, sumbanus.	peroni, sumbana, grandis	peronii, grandis	desmarestii, paliatus, sumbanus	peronii, grandis		peronii, grandis	peronii - desmarestii, paliatus, sumbanus	peronii , palliatus, desmarestii, sumbanus, grandis		peronii, grandis	peronii - desmarestii, paliatus, sumbanus	peronii, palliatus, desmarestii, sumbanus, grandis	peronii, desmarestii, sumbana, desmaresti, grandis, peroni	grandis, peronii	peronii - desmarestii, paliatus, sumbanus	peronii (Ã‰. Geoffroy Saint-Hilaire, 1810)|desmarestii (Burnett, 1829) [nomen nudum]|sumbana Andersen, 1909|peroni E. Schwarz, 1914 [incorrect subsequent spelling]|desmaresti (Laurie & J. Edwards Hill, 1954) [incorrect subsequent spelling | not used as valid]|grandis Bergmans, 1978		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Western naked-backed bat	Lesser Sunda Is; ref. 4.31	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Dobsonia peroni	Indonesia, Lesser Sunda Isis., Timor.	E. Geoffroy	1810	Ann. Mus. Hist. Nat. Paris, 15:104.	Distribution: Confined to the Lesser Sundas from Nusa Penida (just east of Bali) to Babar island (between Timor and Tim orlaut).		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Western naked-backed bat Lesser Sunda Is, Timor	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	E. Geoffroy	1810	Ann. Mus. Hist. Nat. Paris, 15:104.	Reviewed by Bergmans (1978).	Sumba, Timor, Flores, Sumbawa, Nusa Penida (near Bali), Komodo, Alor, Wetar, Babar (Indonesia).	Indonesia, Lesser Sunda Isis, Timor.		E. GEOFFROY	1810	A well-marked antero-internal basal ledge or cusp on the last upper and lower premolars as well as the first lower molar but not on the first upper molar. Longitudinal ridges of molars and premolars without notches. Size relatively small (forearm length, 105-130 mm; total length of skull, 46-54 mm).	Distribution: Confined to the Lesser Sundas from Nusa Penida (just east of Bali) to Babar island (between Timor and Tim orlaut).	Three subspecies are currently recog nized (BERGMANS 1978):	D. p. peroni (Timor, probably also nearby Alor, Wetar, and Babar), D. p. sumbana (Sumba), D. p. grandis (western Les ser Sundas from Nusa Penida to Komodo and probably Flores).	29	species	D. peroni	E. GEOFFROY	1810	Dobsonia	genus	Dobsonia peroni				A well-marked antero-internal basal ledge or cusp on the last upper and lower premolars as well as the first lower molar but not on the first upper molar. Longitudinal ridges of molars and premolars without notches. Size relatively small (forearm length, 105-130 mm; total length of skull, 46-54 mm).	Three subspecies are currently recog nized (BERGMANS 1978):		7. D. peroni (E. GEOFFROY 1810) [peroni group].	7	_D. p. grandis_ Bergmans, 1978; _D. p. peronii_ (Geoffroy Saint-Hilaire, 1810) (synonyms: _desmarestii_ (Burnett, 1829), _sumbana_ Andersen, 1909)			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Dobsonia peronii	Dobsonia		peronii	E. Geoffroy	y	1810		Ann. Mus. Natn. Hist. Nat. Paris	15		104		Western Naked-backed Fruit Bat	Indonesia, Lesser Sunda Isls, Timor.	Bali, Nusa Penida, Lombok, Moyo, Sangeang, Komodo, Sumbawa, Rinca, Flores, Lembata, Pantar, Alor, Wetar, Babar, Timor, Sematu, Roti, Savu, and Sumba Isls (Indonesia).	IUCN/SSC Action Plan (1992) – Indeterminate. IUCN 2003 – Vulnerable.	desmarestii Burnett, 1829 [nomen nudum]; paliatus Geoffroy, 1810; sumbanus K. Andersen, 1909; grandis Bergmans, 1978.	peronii species group. Reviewed by Bergmans (1978) and Kitchener et al. (1997a). Sometimes spelled peroni (e.g., Andersen, 1912; Koopman, 1993).	03AD87FAFFF7F61989653E59FA7AF925	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	109	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFF7F61989653E59FA7AF925.xml	Dobsonia peronii	Pteropodidae	Dobsonia	peronii	E. Geoffroy Saint-Hilaire	1810	Roussette de Péron @fr | Westlicher Nacktriickenflughund @de | Dobsonia de Péron @es	Cephalotes peronii E. Geoffroy Saint-Hilaire, 1810 , “L’ile de Timor,” Lesser Sunda Islands , Indonesia . "Two subspecies are recognized.	D. p. peronii E. Geoffroy Saint-Hilaire, 1810 — E Lesser Sundas (Sumba, Sawu, Roti, Timor, Wetar, Sermata, and Babar Is). D. p. grandis Bergmans, 1978 — W Lesser Sundas ( Bali , Nusa Penida, Lombok, Sumbawa, Moyo, Sangeang, Komodo, Rinca, Flores , Lembata, Pantar, and AlorIs).	Head-body 141-158 mm , tail 29-32 mm , ear 24-26 mm , hindfoot 24-30 mm , forearm 114-121 mm ; weight 162-235 g . Adult male Western Nakedbacked Fruit Bats have back of neck olive, shading to dark olive-bistre on head; rump is brownish olive; flanks are drab, mixed with tawny-olive; breast and belly are suffused with light golden tawny-olive. Females are drab dark brown on head, back, and rump; shoulders are lighter brown; and back and rump are very sparsely haired, appearing almost naked on back. Female ventral fur from upper chest extending down each side is light grayish olive, mixed with drab brown hairs having buffy tips; central chest and abdomen are drab dark brown, tipped with honey-yellow. Young males are lighter than adult females, with shoulders being brown and upper chest and sides light brown. Adult males have very strong musky odor. Index claw (second digit of wing) is absent. There is well-marked anterointernal basal ledge on upper and lower third premolars, a posterior basal ledge in second upper and lower premolars as on third lower premolar. Median surface ridge is present on M' and M, and often also on P° and M,. Anterointernal corner of M' is sharply marked offas a distinct ledge; however, there is no well-marked anterointernal basal ledge in M. Longitudinal ridges of cheekteeth are simple.	Monsoon forests, dry woodland savannas, and moist tropical forests from sea level up to elevations of ¢. 880 m .	Western Naked-backed Fruit Bats eat fruits of Borassus ( Arecaceae ), Muntingia ( Muntingiaceae ), and Ficus ( Moraceae ). They commonly feed from the same trees visited by Geoffroy’s Rousettes ( Rousettus amplexicaudatus ) and Wallacean Gray Flying Foxes ( Pteropus griseus ) .	Two Western Naked-backed Fruit Bats captured in October on Lombok were lactating and had left and right uterine horns slightly swollen. Five adult males on Timor were in breeding condition in March-April, and a female was pregnant in March, with near-term fetus weighing 48 g .	Western Naked-backed Fruit Bats typically emerge ¢.90 minutes after dark and forage throughout the night. Groups roost in limestone caves, rock fissures, crevices, hollow trees, and tree canopies and under rock ledges. Occasionally, individuals roost beneath large fan-shaped fronds of Corypha and Borassus palms ( Arecaceae ).	Up to 300 adults and young Western Naked-backed Fruit Bats can occupy dimly lit large cave chambers, sometimes jointly occupied with Black-bearded Tomb Bats (7aphozous melanopogon ). Smaller groups of young might roost in chambers of a cave not occupied by adults.	Classified as Least Concern on The IUCN Red List. The Western Naked-backed Fruit Bat has a large distribution and is abundant, presumably with stable population. Although hunted, often by smoking them out of caves, consumption is restricted to local indigenous people. The Western Naked-backed Fruit Bat seems to be tolerant of disturbance from human activities. Conservation awareness programs to avoid disturbance at roost caves would be beneficial .	Bergmans (1978b) | Goodwin (1979) | Hutson, Suyanto & Helgen (2008) | Kitchener, Boeadi et al. (1990)		84. Western Naked-backed Fruit Bat Dobsonia peronii French: Roussette de Péron / German: Westlicher Nacktriickenflughund / Spanish: Dobsonia de Péron Taxonomy . Cephalotes peronii E. Geoffroy Saint-Hilaire, 1810 , “L’ile de Timor,” Lesser Sunda Islands , Indonesia . "Two subspecies are recognized. Subspecies and Distribution. D. p. peronii E. Geoffroy Saint-Hilaire, 1810 — E Lesser Sundas (Sumba, Sawu, Roti, Timor, Wetar, Sermata, and Babar Is). D. p. grandis Bergmans, 1978 — W Lesser Sundas ( Bali , Nusa Penida, Lombok, Sumbawa, Moyo, Sangeang, Komodo, Rinca, Flores , Lembata, Pantar, and AlorIs). Descriptive notes . Head-body 141-158 mm , tail 29-32 mm , ear 24-26 mm , hindfoot 24-30 mm , forearm 114-121 mm ; weight 162-235 g . Adult male Western Nakedbacked Fruit Bats have back of neck olive, shading to dark olive-bistre on head; rump is brownish olive; flanks are drab, mixed with tawny-olive; breast and belly are suffused with light golden tawny-olive. Females are drab dark brown on head, back, and rump; shoulders are lighter brown; and back and rump are very sparsely haired, appearing almost naked on back. Female ventral fur from upper chest extending down each side is light grayish olive, mixed with drab brown hairs having buffy tips; central chest and abdomen are drab dark brown, tipped with honey-yellow. Young males are lighter than adult females, with shoulders being brown and upper chest and sides light brown. Adult males have very strong musky odor. Index claw (second digit of wing) is absent. There is well-marked anterointernal basal ledge on upper and lower third premolars, a posterior basal ledge in second upper and lower premolars as on third lower premolar. Median surface ridge is present on M' and M, and often also on P° and M,. Anterointernal corner of M' is sharply marked offas a distinct ledge; however, there is no well-marked anterointernal basal ledge in M. Longitudinal ridges of cheekteeth are simple. Habitat . Monsoon forests, dry woodland savannas, and moist tropical forests from sea level up to elevations of ¢. 880 m . Food and Feeding . Western Naked-backed Fruit Bats eat fruits of Borassus ( Arecaceae ), Muntingia ( Muntingiaceae ), and Ficus ( Moraceae ). They commonly feed from the same trees visited by Geoffroy’s Rousettes ( Rousettus amplexicaudatus ) and Wallacean Gray Flying Foxes ( Pteropus griseus ) . Breeding . Two Western Naked-backed Fruit Bats captured in October on Lombok were lactating and had left and right uterine horns slightly swollen. Five adult males on Timor were in breeding condition in March-April, and a female was pregnant in March, with near-term fetus weighing 48 g . Activity patterns . Western Naked-backed Fruit Bats typically emerge ¢.90 minutes after dark and forage throughout the night. Groups roost in limestone caves, rock fissures, crevices, hollow trees, and tree canopies and under rock ledges. Occasionally, individuals roost beneath large fan-shaped fronds of Corypha and Borassus palms ( Arecaceae ). Movements, Home range and Social organization . Up to 300 adults and young Western Naked-backed Fruit Bats can occupy dimly lit large cave chambers, sometimes jointly occupied with Black-bearded Tomb Bats (7aphozous melanopogon ). Smaller groups of young might roost in chambers of a cave not occupied by adults. Status and Conservation . Classified as Least Concern on The IUCN Red List. The Western Naked-backed Fruit Bat has a large distribution and is abundant, presumably with stable population. Although hunted, often by smoking them out of caves, consumption is restricted to local indigenous people. The Western Naked-backed Fruit Bat seems to be tolerant of disturbance from human activities. Conservation awareness programs to avoid disturbance at roost caves would be beneficial . Bibliography . Bergmans (1978b), Goodwin (1979), Hutson, Suyanto & Helgen (2008), Kitchener, Boeadi et al. (1990).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Dobsonia peronii	Dobsonia		peronii	E. Geoffroy	1810	1	Ann. Mus. Natn. Hist. Nat. Paris	0.6972	Western Naked-backed Fruit Bat	 desmarestii Burnett, 1829 [nomen nudum]; paliatus Geoffroy, 1810; sumbanus K. Andersen, 1909; <b>grandis</b> Bergmans, 1978.	Indonesia, Lesser Sunda Isls, Timor.	Bali, Nusa Penida, Lombok, Moyo, Sangeang, Komodo, Sumbawa, Rinca, Flores, Lembata, Pantar, Alor, Wetar, Babar, Timor, Sematu, Roti, Savu, and Sumba Isls (Indonesia).	Not listed.	Least Concern	 peronii species group. Reviewed by Bergmans (1978) and Kitchener et al . (1997). Sometimes spelled peroni (e.g., Andersen, 1912; Koopman,1993).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Dobsonia peronii	23	Western Naked-backed Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	HARPYIONYCTERINAE	DOBSONIINI	Dobsonia	NA	peronii	Ã‰. Geoffroy Saint-Hilaire	1810	1	Cephalotes_peronii	Geoffroy Saint-Hilaire, Ã‰. (1810). Description de roussettes et des cÃ©phalotes, deux nouveaux genres de la famille des chauve-souris. Annales du MusÃ©um d'histoire naturelle, 15, 104.	https://www.biodiversitylibrary.org/item/23421#page/124/mode/1up	MNHN 1996-2116 [lectotype]		"L'Ã®le de Timor," Lesser Sunda Islands, Indonesia.			peronii (Ã‰. Geoffroy Saint-Hilaire, 1810)|palliatus (Ã‰. Geoffroy Saint-Hilaire, 1810)|desmarestii (Burnett, 1829) [nomen nudum]|sumbanus K. Andersen, 1909|grandis Bergmans, 1978	NA	NA	Indonesia|East Timor	Asia	Indomalaya|Australasia/Oceania	LC	0	0	0	Dobsonia_peronii	0	sciname match	Dobsonia_peronii	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	6771	Dobsonia peronii	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Dobsonia	peronii	(Ã‰. Geoffroy, 1810)		20000000	Dobsonia peronii	Least Concern		2019	2019-07-06 00:00:00 UTC	3.1	English	Listed as Least Concern as the species is widespread in the Lesser Sunda Islands, and considered common throughout its range. Although hunted, the species seems to be quite tolerant of disturbance due to human activities and the population of this species is currently considered stable.	It roosts in caves and rock crevices. A large series was collected from a hollow tree. Colonies up to 300 individuals have been found on Timor (Kitchener et al . 1990). This species seems to be quite tolerant of disturbance.	There are no major threats to this species throughout its range. Hunting (by smoking them out of caves), and cave loss can be localised threats to the species.	This species is common throughout its range.	Stable	This species is found in the Lesser Sunda Islands, Indonesia. It has been recorded from Bali, Nusa Penida, Lombok, Moyo, Sangeang, Komodo, Sumbawa, Rinca, Flores, Lembata, Pantar, Alor, Wetar, Babar, Timor, Sematu, Roti, Savu, and Sumba islands.		Terrestrial	It probably occurs in protected areas within its range.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Dobsonia		peronii	Ã‰. Geoffroy Saint-Hilaire	1810	1	Ann. Mus. Natn. Hist. Nat. Paris	0.697222	Western Naked-backed Fruit Bat	 desmarestii Burnett, 1829 [nomen nudum]; paliatus Geoffroy, 1810; sumbanus K. Andersen, 1909; <b>grandis</b> Bergmans, 1978.	Indonesia, Lesser Sunda Isls, Timor.	Bali, Nusa Penida, Lombok, Moyo, Sangeang, Komodo, Sumbawa, Rinca, Flores, Lembata, Pantar, Alor, Wetar, Babar, Timor, Sematu, Roti, Savu, and Sumba Isls (Indonesia).	Not listed.	Least Concern	 peronii species group. Reviewed by Bergmans (1978) and Kitchener et al . (1997). Sometimes spelled peroni (e.g., Andersen, 1912; Koopman,1993).	Dobsonia peronii	1004401	23	Western Naked-backed Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	HARPYIONYCTERINAE	DOBSONIINI	Dobsonia	NA	peronii	Ã‰. Geoffroy Saint-Hilaire	1810	1	Cephalotes_peronii	Geoffroy Saint-Hilaire, Ã‰. (1810). Description de roussettes et des cÃ©phalotes, deux nouveaux genres de la famille des chauve-souris. Annales du MusÃ©um d'histoire naturelle, 15, 104.	https://www.biodiversitylibrary.org/item/23421#page/124/mode/1up	MNHN 1996-2116 [lectotype]		"L'Ã®le de Timor," Lesser Sunda Islands, Indonesia.			peronii (Ã‰. Geoffroy Saint-Hilaire, 1810)|palliatus (Ã‰. Geoffroy Saint-Hilaire, 1810)|desmarestii (Burnett, 1829) [nomen nudum]|sumbanus K. Andersen, 1909|grandis Bergmans, 1978	NA	NA				Indonesia|East Timor	Asia	Indomalaya|Australasia/Oceania	LC	0	0	0	Dobsonia_peronii	0	sciname match	Dobsonia_peronii	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Dobsonia_peronii	1004401	23	Western Naked-backed Fruit Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Harpyionycterinae	Dobsoniini	Dobsonia	NA	peronii	Ã‰. Geoffroy Saint-Hilaire	1	Cephalotes Peronii	Geoffroy Saint-Hilaire, Ã‰. 1810. Description des roussettes et des cÃ©phalotes, deux nouveaux genres de la famille des chauve-souris. Annales du MusÃ©um d'histoire naturelle 15:86-108.	https://www.biodiversitylibrary.org/page/3546736	MNHN-ZM-MO-1996-2116	holotype	http://coldb.mnhn.fr/catalognumber/mnhn/zm/mo-1996-2116	"L'Ã®le de Timor," Lesser Sunda Islands, Indonesia.			NA	NA				Indonesia|East Timor	Asia	Indomalaya|Australasia	LC	0	0	0	Dobsonia_peronii	0	sciname match	Dobsonia_peronii	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Dobsonia		peronii	Ã‰. Geoffroy Saint-Hilaire	1810	1	Ann. Mus. Natn. Hist. Nat. Paris	0.697222	Western Naked-backed Fruit Bat	desmarestii Burnett, 1829 [nomen nudum]; paliatus Geoffroy, 1810; sumbanus K. Andersen, 1909; grandis Bergmans, 1978.	Indonesia, Lesser Sunda Isls, Timor.	Bali, Nusa Penida, Lombok, Moyo, Sangeang, Komodo, Sumbawa, Rinca, Flores, Lembata, Pantar, Alor, Wetar, Babar, Timor, Sematu, Roti, Savu, and Sumba Isls (Indonesia).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/6771/22034782/' target='_blank'>Least Concern</a>	peronii species group. Reviewed by Bergmans (1978) and Kitchener et al. (1997). Sometimes spelled peroni (e.g., Andersen, 1912; Koopman,1993).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Dobsonia peronii; Dobsonia peronii; Dobsonia peronii; Dobsonia peronii; Dobsonia peronii; Dobsonia peronii; peronii; grandis; desmarestii; paliatus; sumbanus; peronii; grandis; grandis; desmarestii; paliatus; sumbanus; peronii; palliatus; desmarestii; sumbanus; grandis; Roussette de Péron; Westlicher Nacktriickenflughund; Dobsonia de Péron; Western Naked-backed Fruit Bat; Western Naked-backed Fruit Bat; Western Naked-backed Fruit Bat; D. peronii
