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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L244	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Dobsonia pannietensis	Dobsonia pannietensis	Dobsonia pannietensis	Dobsonia pannietensis	Dobsonia pannietensis	Dobsonia pannietensis	Dobsonia pannietensis	Dobsonia pannietensis	Dobsonia pannietensis	Dobsonia pannietensis	Dobsonia pannietensis	Dobsonia pannietensis	Dobsonia pannietensis	Dobsonia pannietensis	Dobsonia pannietensis		[MSW2] Considered a subspecies of moluccensis by Laurie and Hill (1954:42), but as a separate species by Bergmans (1979). Includes remota (see Koopman, 1982:6). A record of remota from Bougainville Isl is based on a misidentified inermis (see Bergmans, 1979).; [MSW3] moluccensis species group. Considered a subspecies of moluccensis by Laurie and Hill (1954), but apparently distinct; see Bergmans (1979) and Bonaccorso (1998). Includes remota; see Koopman (1982). A record of remota from Bougainville Isl is based on a misidentified inermis (see Bergmans, 1979).; [HMW] Cephalotes pannietensis De Vis, 1905 , “Panniet [= Panaeadl] Island, 1.ouisiades [= Louisiade Archipelago, Papua New Guinea ].” Two subspecies are recognized.; [batnames2022]  moluccensis species group. Considered a subspecies of moluccensis by Laurie and Hill (1954), but apparently distinct; see Bergmans (1979) and Bonaccorso (1998). Includes remota ; see Koopman (1982). A record of remota from Bougainville Isl is based on a misidentified inermis (see Bergmans, 1979).; [batnames2023]  moluccensis species group. Considered a subspecies of moluccensis by Laurie and Hill (1954), but apparently distinct; see Bergmans (1979) and Bonaccorso (1998). Includes remota ; see Koopman (1982). A record of remota from Bougainville Isl is based on a misidentified inermis (see Bergmans, 1979).; [batnames2025_1.7] moluccensis species group. Considered a subspecies of moluccensis by Laurie and Hill (1954), but apparently distinct; see Bergmans (1979) and Bonaccorso (1998). Includes remota; see Koopman (1982). A record of remota from Bougainville Isl is based on a misidentified inermis (see Bergmans, 1979).			remota		(anderseni)	remota.	pannietensis, remota			pannietensis, remota		pannietensis, remota		pannietensis, remota		pannietensis, remota		pannietensis, remota	pannietensis, remota	pannietensis, remota		pannietensis (De Vis, 1905)|remota Cabrera, 1920		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Trobriand, D'Entrecasteaux, Louisiade Is	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Dobsonia pannietensis	Papua New Guinea, Louisiade Arch., Panniet Isl.	De Vis	1905	Ann. Queensl. Mus., 6:36.	Distribution: Confined to the east Papuan islands.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	(? remota)	Bismarck Arch., Trobriand, D'Entrecasteaux, Louisiade Is, N Queensland; ref. 4.19	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	De Vis	1905	Ann. Queensl. Mus., 6:36.	Considered a subspecies of moluccensis by Laurie and Hill (1954:42), but as a separate species by Bergmans (1979). Includes remota (see Koopman, 1982:6). A record of remota from Bougainville Isl is based on a misidentified inermis (see Bergmans, 1979).	Louisiade Arch., D'Entrecasteaux Isis, and Trobriand Isis (Papua New Guinea).	Papua New Guinea, Louisiade Arch., Panniet Isl.		DE VIS	1905	Size small for group (forearm length, 97-125 mm; total length of skull, 41-56 mm; maxillary toothrow length, 17-24 mm). Fore head relatively flat. Maxillary dentition relatively weak.	Distribution: Confined to the east Papuan islands.	Two currently recognized subspecies (KOOPMAN 1982):	D. p. pannietensis (Louisiade and D'Entrecasteaux archi pelagos), D. p. remota (Kiriwina and Woodlark islands).	28	species	D. pannietensis	DE VIS	1905	Dobsonia	genus	Dobsonia pannietensis				Size small for group (forearm length, 97-125 mm; total length of skull, 41-56 mm; maxillary toothrow length, 17-24 mm). Fore head relatively flat. Maxillary dentition relatively weak.	Two currently recognized subspecies (KOOPMAN 1982):		6. D. pannietensis (DE VIS 1905) [moluccensis group],	6	_D. p. pannietensis_ (De Vis, 1905); _D. p. remota_ Cabrera, 1920			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Dobsonia pannietensis	Dobsonia		pannietensis	De Vis	y	1905		Ann. Queensl. Mus.	6		36		Panniet Naked-backed Fruit Bat	Papua New Guinea, Louisiade Arch., Panniet Isl.	Louisiade Arch., D'Entrecasteaux Isls, and Trobriand Isls.	IUCN/SSC Action Plan (1992) – No Data. IUCN 2003 – Lower Risk (lc).	remota Cabrera, 1920.	moluccensis species group. Considered a subspecies of moluccensis by Laurie and Hill (1954), but apparently distinct; see Bergmans (1979) and Bonaccorso (1998). Includes remota; see Koopman (1982). A record of remota from Bougainville Isl is based on a misidentified inermis (see Bergmans, 1979).	03AD87FAFFF0F61E89B7308EFB10F464	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	110	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFF0F61E89B7308EFB10F464.xml	Dobsonia pannietensis	Pteropodidae	Dobsonia	pannietensis		1905	Roussette de Panaeati @fr | Panniet-Nacktriickenflughund @de | Dobsonia de Panniet @es | De Vis's Bare-backed Fruit Bat @en | Panaeati Bare-backed Fruit Bat @en	Cephalotes pannietensis De Vis, 1905 , “Panniet [= Panaeadl] Island, 1.ouisiades [= Louisiade Archipelago, Papua New Guinea ].” Two subspecies are recognized.	D.p.pannietensisDeVis,1905—D’Entre-casteauxIs(Goodenough,Fergusson,andNormanby)andLouisiadeArchi-pelago(Misima,Panaeati,Rossel,andVanatinaiIs). D. p. remota Cabrera, 1920 — Trobriand Is (Kiriwina) and Woodlark Is. A record attributed to Bougainville I is a misidentification of the Solomons Nakedbacked Fruit Bat (D. inermas).	Head-body 150-203 mm , tail 21-36 mm , ear 21-28 mm , hindfoot 25-40 mm , forearm 108-125 mm ; weight 180-304 g . Adult male Panniet Nakedbacked Fruit Bats are much heavier than females. Upper back is gray-brown; venteris medium brown, with yellow wash along midline. Head is black-brown and tends to be darker on young individuals. Coarse, erect, brown, or red-brown hairs cover nape and shoulders. Wings meet along spine with underlying fur (naked-backed condition). Claws are pale tan. Ears are long and narrow. Short tail is free of narrow uropatagium that runs along legs to short stiff calcars. Index claw (second digit of wing) is absent. Upper incisors are small, lower pair is minute, and P|is reduced in size.	Tropical moist lowland to lower montane forests from sea level up ¢. 1300 m .	The Panniet Naked-backed Fruit Bat is frugivorous. Native figs are probably important dietary components.	Pregnant Panniet Naked-backed Fruit Bats were observed in August on Goodenough (first trimester with 9-g fetus) and October on Woodlark. A lactating female was recorded in August on Woodlark.	The Panniet Naked-backed Fruit Bat is nocturnal and roosts in tree hollows and limestone caves.	The Panniet Naked-backed Fruit Bat is gregarious, roosting in small groups in cave twilight zones. It is host to the ectoparasitic streblid bat fly Megastrebla gigantea.	Classified as Near Threatened on The IUCN Red List. The Panniet Naked-backed Fruit Bat has a distribution ofless than 6000 km *, and quality of its habitat is declining. It is close to qualifying as Vulnerable. Important roosts should be protected. Panniet Naked-backed Fruit Bats occur in protected wildlife management areas on Goodenough and at Lake LLawma on Fergusson. It is likely more vulnerable on the Trobriands and Woodlark Island (subspecies remota). It is threatened by deforestation in parts of its distribution. Hunting might be a threat becauseit is large enough to be a good source of meat. Additional studies are needed on its population trends, ecology, and conservation threats.	Bonaccorso (1998) | Flannery (1995a) | Helgen, Allison et al. (2008b) | Maa (1971)	https://zenodo.org/record/6803758/files/figure.png	86. Panniet Naked-backed Fruit Bat Dobsonia pannietensis French: Roussette de Panaeati / German: Panniet-Nacktriickenflughund / Spanish: Dobsonia de Panniet Other common names: De Vis's Bare-backed Fruit Bat , Panaeati Bare-backed Fruit Bat Taxonomy. Cephalotes pannietensis De Vis, 1905 , “Panniet [= Panaeadl] Island, 1.ouisiades [= Louisiade Archipelago, Papua New Guinea ].” Two subspecies are recognized. Subspecies and Distribution. D.p.pannietensisDeVis,1905—D’Entre-casteauxIs(Goodenough,Fergusson,andNormanby)andLouisiadeArchi-pelago(Misima,Panaeati,Rossel,andVanatinaiIs). D. p. remota Cabrera, 1920 — Trobriand Is (Kiriwina) and Woodlark Is. A record attributed to Bougainville I is a misidentification of the Solomons Nakedbacked Fruit Bat (D. inermas). Descriptive notes. Head-body 150-203 mm , tail 21-36 mm , ear 21-28 mm , hindfoot 25-40 mm , forearm 108-125 mm ; weight 180-304 g . Adult male Panniet Nakedbacked Fruit Bats are much heavier than females. Upper back is gray-brown; venteris medium brown, with yellow wash along midline. Head is black-brown and tends to be darker on young individuals. Coarse, erect, brown, or red-brown hairs cover nape and shoulders. Wings meet along spine with underlying fur (naked-backed condition). Claws are pale tan. Ears are long and narrow. Short tail is free of narrow uropatagium that runs along legs to short stiff calcars. Index claw (second digit of wing) is absent. Upper incisors are small, lower pair is minute, and P|is reduced in size. Habitat. Tropical moist lowland to lower montane forests from sea level up ¢. 1300 m . Food and Feeding. The Panniet Naked-backed Fruit Bat is frugivorous. Native figs are probably important dietary components. Breeding. Pregnant Panniet Naked-backed Fruit Bats were observed in August on Goodenough (first trimester with 9-g fetus) and October on Woodlark. A lactating female was recorded in August on Woodlark. Activity patterns. The Panniet Naked-backed Fruit Bat is nocturnal and roosts in tree hollows and limestone caves. Movements, Home range and Social organization. The Panniet Naked-backed Fruit Bat is gregarious, roosting in small groups in cave twilight zones. It is host to the ectoparasitic streblid bat fly Megastrebla gigantea. Status and Conservation. Classified as Near Threatened on The IUCN Red List. The Panniet Naked-backed Fruit Bat has a distribution ofless than 6000 km *, and quality of its habitat is declining. It is close to qualifying as Vulnerable. Important roosts should be protected. Panniet Naked-backed Fruit Bats occur in protected wildlife management areas on Goodenough and at Lake LLawma on Fergusson. It is likely more vulnerable on the Trobriands and Woodlark Island (subspecies remota). It is threatened by deforestation in parts of its distribution. Hunting might be a threat becauseit is large enough to be a good source of meat. Additional studies are needed on its population trends, ecology, and conservation threats. Bibliography. Bonaccorso (1998), Flannery (1995a), Helgen, Allison et al. (2008b), Maa (1971).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Dobsonia pannietensis	Dobsonia		pannietensis	De Vis	1905	1	Ann. Queensl. Mus.	6:36	Panniet Naked-backed Fruit Bat	<b> remota </b>Cabrera, 1920.	Papua New Guinea, Louisiade Arch., Panniet Isl.	Louisiade Arch., D'Entrecasteaux Isls, and Trobriand Isls.	Not listed.	Near Threatened	 moluccensis species group. Considered a subspecies of moluccensis by Laurie and Hill (1954), but apparently distinct; see Bergmans (1979) and Bonaccorso (1998). Includes remota ; see Koopman (1982). A record of remota from Bougainville Isl is based on a misidentified inermis (see Bergmans, 1979).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Dobsonia pannietensis	23	Panniet Naked-backed Fruit Bat	De Vis's Bare-backed Fruit Bat|Panaeati Bare-backed Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	HARPYIONYCTERINAE	DOBSONIINI	Dobsonia	NA	pannietensis	De Vis	1905	1						"Panniet [= Panaeati] Island, Louisiades [= Louisiade Archipelago, Papua New Guinea]."			pannietensis (De Vis, 1905)|remota Cabrera, 1920	NA	NA	Papua New Guinea	Oceania	Australasia/Oceania	NT	0	0	0	Dobsonia_pannietensis	0	sciname match	Dobsonia_pannietensis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	6776	Dobsonia pannietensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Dobsonia	pannietensis	(De Vis, 1905)		20000000	Dobsonia pannietensis	Near Threatened	A2c	2020	2019-05-30 00:00:00 UTC	3.1	English	Dobsonia pannietensis is assessed as Near Threatened under criterion A2c as the species is suspected to have experienced a past decline of 25-29% over the last three (3) generations (17.4 years; GL = 5.8 years; Pacifici et al. , 2013) due to the loss of forests from logging, conversion to oil palm plantations, mining, and hunting. The species’ range is restricted to 10 islands and is assumed to be in ca. 6 subpopulations due to the spatial distribution and proximity of the islands.	This little-known species roosts in groups in caves and tree hollows (Flannery 1995, Bonaccorso 1998). Nothing is known of the diet of the species but it is presumably largely frugivorous like other members of the genus. It is unknown whether the species utilises degraded secondary rainforest. Little is known of reproduction in the species but pregnant and lactating females have been recorded in August and October (Flannery 1995, Bonaccorso 1998).	Dobsonia pannietensis is threatened by continuing degradation and loss of primary and secondary forests over much of its range. For example, between 1972 and 2002 the D’Entrecasteaux Island Group had the highest rate of deforestation (28% of forests) in Papua New Guinea, and the East Papuan islands similarly recorded high rates of deforestation and degradation (Shearman and Bryan 2011). Almost half of Woodlark Island (which makes up the majority of the area of the East Papuan islands) has been allocated to the logging industry (Shearman and Bryan 2011). Logging and clearing of forest for oil palm plantations is occurring on many of the larger islands and further areas are likely to be impacted in the future. Hunting, including the disturbance of important roosts, is likely a significant threat as it is a large species, similar species of Dobsonia are hunted at their cave roosts. Open-cut mining is currently proposed for Woodlark Island, as is re-opening of the Misima mine, presenting additional potential threats to the species from both deforestation and destabilisation of roosting caves (T. Leary pers. comm.).	Dobsonia pannietensis is suspected to have experienced a past decline of 25–29% over the last three (3) generations (17.4 years; GL = 5.8 years; Pacifici et al. , 2013) due to the loss of forests from logging, conversion to oil palm plantations, mining, and hunting. The species’ range is restricted to 10 islands and is assumed to be in ca. 6 subpopulations due to the spatial distribution and proximity of the islands. It was reportedly very common in some parts of its range (e.g., Woodlark and Normanby Islands) as recent as the 1980s and 1990s (Flannery 1995).	Decreasing	This species is endemic to Milne Bay Province, Papua New Guinea. It has been recorded from the islands of Fergusson, Goodenough, Normanby, Misima, Panaeati, Rossel, Sudest, Sideia, Kiriwina, and Woodlark (Flannery 1995, Bonaccorso 1998, T. Leary pers. comm.). It has been recorded at elevations of between sea level and 1,800 m asl.	The species is assumed to be hunted as similar species in the genus are hunted at their cave roosts.	Terrestrial	Dobsonia pannietensis is not known to be present in any protected areas, except perhaps a wildlife management area on Goodenough Island and Lake Lawma on Fergusson Island. Immediate protection of ;important roosts, and the remaining primary forest tracts within the species range, is required for the conservation of this poorly-known species. Further studies are needed into the species ecology, habitat use and distribution, population trends and threats the species. It is likely more vulnerable on the Trobriands and Woodlark Island (as subspecies, D. p. remota ).	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Dobsonia		pannietensis	De Vis	1905	1	Ann. Queensl. Mus.	6:36	Panniet Naked-backed Fruit Bat	<b> remota </b>Cabrera, 1920.	Papua New Guinea, Louisiade Arch., Panniet Isl.	Louisiade Arch., D'Entrecasteaux Isls, and Trobriand Isls.	Not listed.	Near Threatened	 moluccensis species group. Considered a subspecies of moluccensis by Laurie and Hill (1954), but apparently distinct; see Bergmans (1979) and Bonaccorso (1998). Includes remota ; see Koopman (1982). A record of remota from Bougainville Isl is based on a misidentified inermis (see Bergmans, 1979).	Dobsonia pannietensis	1004400	23	Panniet Naked-backed Fruit Bat	De Vis's Bare-backed Fruit Bat|Panaeati Bare-backed Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	HARPYIONYCTERINAE	DOBSONIINI	Dobsonia	NA	pannietensis	De Vis	1905	1						"Panniet [= Panaeati] Island, Louisiades [= Louisiade Archipelago, Papua New Guinea]."			pannietensis (De Vis, 1905)|remota Cabrera, 1920	NA	NA				Papua New Guinea	Oceania	Australasia/Oceania	NT	0	0	0	Dobsonia_pannietensis	0	sciname match	Dobsonia_pannietensis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Dobsonia_pannietensis	1004400	23	Panniet Naked-backed Fruit Bat	De Vis's Bare-backed Fruit Bat|Panaeati Bare-backed Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Harpyionycterinae	Dobsoniini	Dobsonia	NA	pannietensis	De Vis	1	Cephalotes pannietensis	De Vis, C.W. 1905. Bats. Annals of the Queensland Museum 6:36-40.	https://www.biodiversitylibrary.org/page/48070439	QM J1964	lectotype	https://collections.qm.qld.gov.au/objects/VE122921/dobsonia-pannietensis	"Panniet [= Panaeati] Island, Louisiades [= Louisiade Archipelago, Papua New Guinea]."			NA	NA				Papua New Guinea	Oceania (Continent)	Australasia	NT	0	0	0	Dobsonia_pannietensis	0	sciname match	Dobsonia_pannietensis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Dobsonia		pannietensis	De Vis	1905	1	Ann. Queensl. Mus.	6:36	Panniet Naked-backed Fruit Bat	remota Cabrera, 1920.	Papua New Guinea, Louisiade Arch., Panniet Isl.	Louisiade Arch., D'Entrecasteaux Isls, and Trobriand Isls.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/6776/22034157/' target='_blank'>Near Threatened</a>	moluccensis species group. Considered a subspecies of moluccensis by Laurie and Hill (1954), but apparently distinct; see Bergmans (1979) and Bonaccorso (1998). Includes remota; see Koopman (1982). A record of remota from Bougainville Isl is based on a misidentified inermis (see Bergmans, 1979).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Dobsonia pannietensis; Dobsonia pannietensis; Dobsonia pannietensis; Dobsonia pannietensis; Dobsonia pannietensis; Dobsonia pannietensis; pannietensis; remota; remota; pannietensis; remota; Roussette de Panaeati; Panniet-Nacktriickenflughund; Dobsonia de Panniet; De Vis's Bare-backed Fruit Bat; Panaeati Bare-backed Fruit Bat; Panniet Naked-backed Fruit Bat; De Vis's Bare-backed Fruit Bat; Panaeati Bare-backed Fruit Bat; Panniet Naked-backed Fruit Bat; Panniet Naked-backed Fruit Bat; D. pannietensis