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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L25	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Pipistrellus angulatus	Pipistrellus tenuis [synonym of]	Pipistrellus angulatus	Pipistrellus tenuis [synonym of]	Pipistrellus tenuis angulatus	Pipistrellus angulatus	Pipistrellus angulatus	Pipistrellus angulatus	Pipistrellus angulatus	Pipistrellus angulatus	Pipistrellus angulatus	Pipistrellus angulatus	Pipistrellus angulatus	Pipistrellus angulatus	Alionoctula angulata		[MSW3] Subgenus Pipistrellus. Included in tenuis by Koopman (1993, 1994), but see Kitchener et al. (1986). Also see Flannery (1995a, b) and Bonaccorso (1998).; [HMW] Vesperugo angulatus Peters, 1880 , “Insula Papuana, ‘Duke of York’ [Islands],” East New Britain Province , Papua New Guinea . Pipustrellus angulatus has been included under P. tenuis , but it and subspecies ponceleti are considered distinct species based on morphology. Two subspecies recognized.; [batnames2022] Subgenus Pipistrellus . Included in tenuis by Koopman (1993, 1994), but see Kitchener et al. (1986). Also see Flannery (1995 a , b ) and Bonaccorso (1998).; [IUCN] Koopman in Wilson and Reeder (1993) treat this as a synonym of Pipistrellus tenuis whereas others consider it a good species (Simmons 2005).; [batnames2023] Subgenus Pipistrellus . Included in tenuis by Koopman (1993, 1994), but see Kitchener et al. (1986). Also see Flannery (1995 a , b ) and Bonaccorso (1998).; [batnames2025_1.7] Subgenus Pipistrellus. Included in tenuis by Koopman (1993, 1994), but see Kitchener et al. (1986). Also see Flannery (1995 a, b) and Bonaccorso (1998).; [MDD2025_2.2] moved from Pipistrellus to Alionoctula		(collinus) (papuanus)						angulatus, ponceleti		angulatus, ponceleti		angulatus, ponceleti		angulatus, ponceleti	Koopman in Wilson and Reeder (1993) treat this as a synonym of Pipistrellus tenuis whereas others consider it a good species (Simmons 2005).	angulatus, ponceleti		angulatus, ponceleti 	angulatus, ponceleti 	angulatus, ponceleti		angulata (W. C. H. Peters, 1880)|ponceleti (Troughton, 1936)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Greater New Guinea pipistrelle	New Guinea etc., Solomons, S Moluccas, N, ? W Australia, ? N Celebes; possibly conspecific with P. tenuis; ref. 4.84		N/A							Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Greater New Guinea pipistrelle	New Guinea, Bismarck Arch., Solomons; refs. 4.12, 72, 128																															_A. a. angulata_ (Peters, 1880); _A. a. ponceleti_ (Troughton, 1936)			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Pipistrellini	Pipistrellus angulatus	Pipistrellus	Pipistrellus	angulatus	Peters		1880		Sitz. Ges. Naturf. Freunde			122		Angulate Pipistrelle	Duke of York Isl, between New Britain and New Ireland (New Hebrides, = Vanuatu).	New Guinea; Bismarck Arch.; Bougainville Isl and Solomon Isls; adjacent small islands.	IUCN 2003 – Not listed (lapsus); IUCN/SSC Action Plan (2001) – Lower Risk (lc).	ponceleti Troughton, 1936.	Subgenus Pipistrellus. Included in tenuis by Koopman (1993, 1994), but see Kitchener et al. (1986). Also see Flannery (1995a, b) and Bonaccorso (1998).	4C3D87E8FFE36A5DFA4F90F71DBEBAFA	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	782	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFE36A5DFA4F90F71DBEBAFA.xml	Pipistrellus angulatus	Vespertilionidae	Pipistrellus	angulatus	Peters	1880	Pipistrelle anguleuse @fr | Neu-Guinea-Zwergfledermaus @de | Pipistrelade Nueva Guinea @es | Angulate Pipistrelle @en	Vesperugo angulatus Peters, 1880 , “Insula Papuana, ‘Duke of York’ [Islands],” East New Britain Province , Papua New Guinea . Pipustrellus angulatus has been included under P. tenuis , but it and subspecies ponceleti are considered distinct species based on morphology. Two subspecies recognized.	P.a. angulatus Peters, 1880 - E New Guinea and Biak-Supiori, Mioko, Manus, Karkar, Emirau, New Ireland, Duke of York, New Britain, Goodenough, Fergusson, Normanby, Misima, and Sudest Is. P. a. ponceleti Troughton, 1936 — Solomon Is, including Bougainville, Fauro, New Georgia , Guadalcanal, Santa Isabel and Nendo Is.	Head-body 33-4— 48 mm , tail 27-41 mm, ear 9-3-15-5 mm, hindfoot 6-10 mm, forearm 31-37-4 mm; weight 3-2—4-3 g. Dorsal pelage of the New Guinea Pipistrelle is dark blackish brown; ventral pelage is pale brown, with some whitish hairs intermixed. Skin on face, ears, arms, and wing membranes are medium brown. Uropatagium is lightly furred, with cinnamon-brown hair along venation; forearm is lightly furred under humerus. Ears are triangular, with rounded tips, and posterior edge is conspicuously concave; tragus narrows slightly at rounded tip and has slightly concave anterior edge, gently convex posterior edge, and small basal lobe. Uropatagium stretches nearly to end of tail (only extreme tip is free), and postcalcarial lobe is elongated. Glans penisis narrow, with slight ventral folds forming shallow mesial groove. Baculum is long, with narrow base and shaft, and is bifurcated at tip (10% of total length). Skull is of moderate size, with very concave rostral profile; sagittal crest is absent; lambdoidal crest is moderately developed laterally and becomes less apparent toward apex where it is absent; zygomatic arches are moderately wide basally but weaker in middle; I* is bicuspid, and I” is unicuspid and subequal in height to second cusp of I*; P* is less than one-half the height of P* and is slightly displaced from tooth row; and lower molars are nyctalodont.	Tall primary and secondary tropical moist forests and some urban areas from sea level up to elevations of ¢. 2400 m .	New Guinea Pipistrelles are insectivorous.	Two pregnant New Guinea Pipistrelles were captured in June.	New Guinea Pipistrelles emerge from roosts at dusk. They roost in caves, buildings, and bamboo clumps. They are seemingly slow fliers, with many erratic turns. Search-call shape is FM/QCEF, with a characteristic frequency of 48 kHz on Manus Island.	A maternity colony of 200 individuals was found in June on New Ireland in a very close-knit cluster in a 500 x 66 cm cave ceiling. Males have often been found alone.	Classified as Least Concern on The IUCN Red List. The New Guinea Pipistrelle does not seem to face any major threats.	Aplin & Opiang (2009) | Aplin et al. (2015) | Armstrong, Novera & Aplin (2015) | Bonaccorso (1998) | Bonaccorso & Leary (2008d) | Flannery (1995a, 1995b) | Kitchener et al. (1986)	https://zenodo.org/record/6397864/files/figure.png	45. New Guinea Pipistrelle Pipistrellus angulatus French: Pipistrelle anguleuse / German: Neu-Guinea-Zwergfledermaus / Spanish: Pipistrela de Nueva Guinea Other common names: Angulate Pipistrelle Taxonomy. Vesperugo angulatus Peters, 1880 , “Insula Papuana, ‘Duke of York’ [Islands],” East New Britain Province , Papua New Guinea . Pipustrellus angulatus has been included under P. tenuis , but it and subspecies ponceleti are considered distinct species based on morphology. Two subspecies recognized. Subspecies and Distribution. P.a. angulatus Peters, 1880 - E New Guinea and Biak-Supiori, Mioko, Manus, Karkar, Emirau, New Ireland, Duke of York, New Britain, Goodenough, Fergusson, Normanby, Misima, and Sudest Is. P. a. ponceleti Troughton, 1936 — Solomon Is, including Bougainville, Fauro, New Georgia , Guadalcanal, Santa Isabel and Nendo Is. Descriptive notes. Head-body 33-4— 48 mm , tail 27-41 mm, ear 9-3-15-5 mm, hindfoot 6-10 mm, forearm 31-37-4 mm; weight 3-2—4-3 g. Dorsal pelage of the New Guinea Pipistrelle is dark blackish brown; ventral pelage is pale brown, with some whitish hairs intermixed. Skin on face, ears, arms, and wing membranes are medium brown. Uropatagium is lightly furred, with cinnamon-brown hair along venation; forearm is lightly furred under humerus. Ears are triangular, with rounded tips, and posterior edge is conspicuously concave; tragus narrows slightly at rounded tip and has slightly concave anterior edge, gently convex posterior edge, and small basal lobe. Uropatagium stretches nearly to end of tail (only extreme tip is free), and postcalcarial lobe is elongated. Glans penisis narrow, with slight ventral folds forming shallow mesial groove. Baculum is long, with narrow base and shaft, and is bifurcated at tip (10% of total length). Skull is of moderate size, with very concave rostral profile; sagittal crest is absent; lambdoidal crest is moderately developed laterally and becomes less apparent toward apex where it is absent; zygomatic arches are moderately wide basally but weaker in middle; I* is bicuspid, and I” is unicuspid and subequal in height to second cusp of I*; P* is less than one-half the height of P* and is slightly displaced from tooth row; and lower molars are nyctalodont. Habitat. Tall primary and secondary tropical moist forests and some urban areas from sea level up to elevations of ¢. 2400 m . Food and Feeding. New Guinea Pipistrelles are insectivorous. Breeding. Two pregnant New Guinea Pipistrelles were captured in June. Activity patterns. New Guinea Pipistrelles emerge from roosts at dusk. They roost in caves, buildings, and bamboo clumps. They are seemingly slow fliers, with many erratic turns. Search-call shape is FM/QCEF, with a characteristic frequency of 48 kHz on Manus Island. Movements, Home range and Social organization. A maternity colony of 200 individuals was found in June on New Ireland in a very close-knit cluster in a 500 x 66 cm cave ceiling. Males have often been found alone. Status and Conservation. Classified as Least Concern on The IUCN Red List. The New Guinea Pipistrelle does not seem to face any major threats. Bibliography. Aplin & Opiang (2009), Aplin et al. (2015), Armstrong, Novera & Aplin (2015), Bonaccorso (1998), Bonaccorso & Leary (2008d), Flannery (1995a, 1995b), Kitchener et al. (1986).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Pipistrellus angulatus	Pipistrellus	Pipistrellus	angulatus	Peters	1880	0	Sitz. Ges. Naturf. Freunde	p. 122	Angulate Pipistrelle	<b> ponceleti </b>Troughton, 1936.	Duke of York Isl, between New Britain and New Ireland (New Hebrides, = Vanuatu).	New Guinea; Bismarck Arch.; Bougainville Isl and Solomon Isls; adjacent small islands.	Not listed.	Least Concern	Subgenus Pipistrellus . Included in tenuis by Koopman (1993, 1994), but see Kitchener et al. (1986). Also see Flannery (1995 a , b ) and Bonaccorso (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Pipistrellus angulatus	23	New Guinea Pipistrelle	Angulate Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	PIPISTRELLINI	Pipistrellus	NA	angulatus	W. Peters	1880	1	Vesperugo_angulatus	Peters, W. C. H. (1880). Mittheilung über eine neue Art von Flederthieren, Vesperugo angulatus, von der Papua-Insel Duke of York. Sitzungsberichte der naturforschenden Freunde zu Berlin, 1880, 122.	https://www.biodiversitylibrary.org/item/35623#page/136/mode/1up	ZMB 5492		"Insula Papuana, ‘Duke of York' [Islands]," East New Britain Province, Papua New Guinea.			angulatus (W. Peters, 1880)|ponceleti Troughton, 1936	NA	NA	Indonesia|Papua New Guinea|Solomon Islands	Asia|Oceania	Australasia/Oceania	LC	0	0	0	Pipistrellus_angulatus	0	sciname match	Pipistrellus_angulatus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	17326	Pipistrellus angulatus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Pipistrellus	angulatus	Peters, 1880	Koopman in Wilson and Reeder (1993) treat this as a synonym of Pipistrellus tenuis whereas others consider it a good species (Simmons 2005).	20000000	Pipistrellus angulatus	Least Concern		2020	2019-07-31 00:00:00 UTC	3.1	English	<p>This species is listed as Least Concern in view of its wide distribution, tolerance of a high degree of habitat modification, presumed large population, and because it is not believed to be declining.</p>	<p>This species roosts in caves, buildings, and bamboo clumps. Unlike Pipistrellus papuanus , it does not roost in holes in coconut palms. The species forages in tall primary and secondary tropical moist forest, areas recently logged, gardens, villages and urban areas. On New Guinea, this species has been recorded emerging from their roost at dusk to forage on aerial insects. A maternity colony of 200 bats in a cave was active in June on the island of New Ireland in the Bismarck Archipelago (Flannery 1995b, Bonaccorso 1998). In the Solomon Islands it has a short (4-5 millisecond) frequency modulated echolocation call dropping from around 70 to 47 kHz with a brief quasi-constant frequency section around 47.5 kHz (Pennay and Lavery 2017), similar calls have also been observed from the species in Manus Island in Papua New Guinea (Armstrong et al. 2015).</p>	<p>There is widespread logging and habitat clearance for gardens throughout the range of this species, however the species appears tolerant of these disturbances. Feral cats and invasive species, particularly fire ants may pose a threat to the species in island populations.</p>	<p>Poorly known but presumed to be large. It was recorded as “super abundant” and found in all habitats on Manus Island by Armstrong et al. (2015). Acoustic sampling in the Solomon Islands found this species to be one of the most commonly recorded species in a range of habitats including villages, gardens and patches of recently logged forest (Pennay and Lavery 2017).</p>	Stable	<p>This widespread species ranges from the island of Biak (Indonesia), through parts of the island of New Guinea (Indonesia and Papua New Guinea), the Bismark Archipelago, many islands of Milne Bay Province, and the island of Bougainville (all Papua New Guinea), to the Solomon Islands as far south as the island of Nendo (Flannery 1995a,b; Bonaccorso 1998). Its distribution on the island of New Guinea is poorly known. It has been recorded between sea level and 2,400 m asl.</p>		Terrestrial	<p>It is presumed to occur in some protected areas. Important roosting and foraging sites should be identified and protected.</p>	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Pipistrellus	Pipistrellus	angulatus	Peters	1880	0	Sitz. Ges. Naturf. Freunde	p. 122	Angulate Pipistrelle	<b> ponceleti </b>Troughton, 1936.	Duke of York Isl, between New Britain and New Ireland (New Hebrides, = Vanuatu).	New Guinea; Bismarck Arch.; Bougainville Isl and Solomon Isls; adjacent small islands.	Not listed.	Least Concern	Subgenus Pipistrellus . Included in tenuis by Koopman (1993, 1994), but see Kitchener et al. (1986). Also see Flannery (1995 a , b ) and Bonaccorso (1998).	Pipistrellus angulatus	1005612	23	New Guinea Pipistrelle	Angulate Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	PIPISTRELLINI	Pipistrellus	NA	angulatus	W. Peters	1880	1	Vesperugo_angulatus	Peters, W. C. H. (1880). Mittheilung über eine neue Art von Flederthieren, Vesperugo angulatus, von der Papua-Insel Duke of York. Sitzungsberichte der naturforschenden Freunde zu Berlin, 1880, 122.	https://www.biodiversitylibrary.org/item/35623#page/136/mode/1up	ZMB 5492		"Insula Papuana, ‘Duke of York' [Islands]," East New Britain Province, Papua New Guinea.			angulatus (W. Peters, 1880)|ponceleti Troughton, 1936	NA	NA				Indonesia|Papua New Guinea|Solomon Islands	Asia|Oceania	Australasia/Oceania	LC	0	0	0	Pipistrellus_angulatus	0	sciname match	Pipistrellus_angulatus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Alionoctula_angulata	1005612	23	New Guinea Pipistrelle	Angulate Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Pipistrellini	Alionoctula	NA	angulata	W. C. H. Peters	1	Vesperugo angulatus	Peters, W.C.H. 1880. Mittheilung über eine neue Art von Flederthieren, Vesperugo angulatus, von der Papua-Insel Duke of York. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1880:122.	https://www.biodiversitylibrary.org/page/8807354	ZMB 5492	holotype		"Insula Papuana, ‘Duke of York' [Islands]," East New Britain Province, Papua New Guinea.			moved from Pipistrellus to Alionoctula	Zhukova, S. S., Yuzefovich, A. P., Lebedev, V. S., & Kruskop, S. V. (2025). Reassessment of the Taxonomic Borders Within Pipistrellus (Chiroptera, Vespertilionidae, Pipistrellini). Diversity, 17(5), 317.				Indonesia|Papua New Guinea|Solomon Islands	Asia|Oceania (Continent)	Australasia	LC	0	0	0	Pipistrellus_angulatus	0	sciname match	Pipistrellus_angulatus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Pipistrellus	Pipistrellus	angulatus	Peters	1880	1	Sitz. Ges. Naturf. Freunde	p. 122	Angulate Pipistrelle	ponceleti Troughton, 1936.	Duke of York Isl, between New Britain and New Ireland (New Hebrides, = Vanuatu).	New Guinea; Bismarck Arch.; Bougainville Isl and Solomon Isls; adjacent small islands.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/17326/22131495/' target='_blank'>Least Concern</a>	Subgenus Pipistrellus. Included in tenuis by Koopman (1993, 1994), but see Kitchener et al. (1986). Also see Flannery (1995 a, b) and Bonaccorso (1998).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Pipistrellus angulatus; Pipistrellus angulatus; Pipistrellus angulatus; Pipistrellus angulatus; Pipistrellus angulatus; Pipistrellus angulatus; angulatus; ponceleti; angulatus; ponceleti; ponceleti; angulatus; ponceleti; Pipistrelle anguleuse; Neu-Guinea-Zwergfledermaus; Pipistrelade Nueva Guinea; Angulate Pipistrelle; New Guinea Pipistrelle; Angulate Pipistrelle; Angulate Pipistrelle; Angulate Pipistrelle; P. angulatus