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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L235	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Dobsonia beauforti	Dobsonia beauforti	Dobsonia beauforti	Dobsonia beauforti	Dobsonia beauforti	Dobsonia beauforti	Dobsonia beauforti	Dobsonia beauforti	Dobsonia beauforti	Dobsonia beauforti	Dobsonia beauforti	Dobsonia beauforti	Dobsonia beauforti	Dobsonia beauforti	Dobsonia beauforti		[MSW2] Closely related to viridis (see Bergmans, 1975).; [MSW3] viridis species group. Closely related to viridis; see Bergmans (1975). Also see Flannery (1995b).; [HMW] Dobsonia beauforti Bergmans, 1975 , “from a cave near Nja-njef, Waigeo [ Island , Sorong Division, Papua Province], New Guinea ,” Indonesia . This species is monotypic.; [batnames2022]  viridis species group. Closely related to viridis ; see Bergmans (1975). Also see Flannery (1995).; [IUCN] This may possibly be conspecific with Dobsonia viridis .; [batnames2023]  viridis species group. Closely related to viridis ; see Bergmans (1975). Also see Flannery (1995).; [batnames2025_1.7] viridis species group. Closely related to viridis; see Bergmans (1975). Bonaccorso (2019) suggests that the record from Sulawati represents a vagrant and that this species may be present on Misool Island as well. Also see Flannery (1995).														beauforti	This may possibly be conspecific with Dobsonia viridis .			beauforti	beauforti			beauforti Bergmans, 1975		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Waigeo I, off NW New Guinea	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Dobsonia beauforti	Indonesia, Irian Jaya, So Rong Div., Waigeo Isl., Njanjef.	Bergmans	1975	Beaufortia, 23(295):3.	Distribution: Confined to Waigeo in the west Papuan islands and to Biak and Owii islands in Geelvink Bay.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Biak I, Owii I, Waigeo I, off NW New Guinea	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Bergmans	1975	Beaufortia, 23(295):3.	Closely related to viridis (see Bergmans, 1975).	Waigeo Isl (off west end of New Guinea).	Indonesia, Irian Jaya, So Rong Div., Waigeo Isl, Njanjef.		BERGMANS	1975	Size small for group (forearm length, 99 -112 mm; total length of skull, 41 -4 6 mm). Teeth relatively small (maxillary toothrow length, 16-17 mm). In terorbital width relatively great (7.9-8.8 mm).	Distribution: Confined to Waigeo in the west Papuan islands and to Biak and Owii islands in Geelvink Bay.	No supspecies.		29	species	D. beauforti	BERGMANS	1975	Dobsonia	genus	Dobsonia beauforti				Size small for group (forearm length, 99 -112 mm; total length of skull, 41 -4 6 mm). Teeth relatively small (maxillary toothrow length, 16-17 mm). In terorbital width relatively great (7.9-8.8 mm).	No supspecies.		9. D. beauforti BERGMANS 1975 [viridis group],	9	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Dobsonia beauforti	Dobsonia		beauforti	Bergmans		1975		Beaufortia	23	295	3		Beaufort’s Naked-backed Fruit Bat	Indonesia, Prov. of Papua, Sorong Div., Waigeo Isl, Njanjef.	Waigeo, Batanta, Salawati, Gebe, Gag, and Biak Isls (off Vogelkop Peninsula, New Guinea).	IUCN/SSC Action Plan (1992) – No Data: Limited Distribution. IUCN 2003 – Endangered.		viridis species group. Closely related to viridis; see Bergmans (1975). Also see Flannery (1995b).	03AD87FAFFF1F61C8C6D3274FAA0FCAE	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	111	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFF1F61C8C6D3274FAA0FCAE.xml	Dobsonia beauforti	Pteropodidae	Dobsonia	beauforti	Bergmans	1975	Roussette de Beaufort @fr | \Waigeo-Nacktriickenflughund @de | Dobsonia de Beaufort @es | Beaufort's Bare-backed Fruit Bat @en	Dobsonia beauforti Bergmans, 1975 , “from a cave near Nja-njef, Waigeo [ Island , Sorong Division, Papua Province], New Guinea ,” Indonesia . This species is monotypic.	Raja Ampat Is (Gebe, Gag, Waigeo, and Batanta), Schouten Is (Biak-Supiori and Owi) off NWNew Guinea ;it possibly may be found on MisoolI, a single individual reported from Salawati I probably represents a vagrant.	Head-body 120-155 mm , tail 20-28 mm , ear 23-27 mm , forearm 100-114 mm ; weight 116-163 g . Males are appreciably larger than females in mean weight (males 158 g and females 133 g ) and slightly larger in mean forearm length (males 108 mm and females 106 mm ). Breeding males differ from females in having brighter greenish yellow fur, proportionately more massive heads with prominent white cheek glands, and almost naked skin on rest of face. Females tend to be more brownish, have more gracile head that is more heavily furred, and lack inflated cheek glands. Young have green-yellow wash to fur, more similar to adult males. Fur on top and side of head is bone brown to olive-brown (greenish yellow in some descriptions). Nape, shoulders, and furred parts of back are olive-brown and somewhat darker in occipital region and bordering naked skin including narrow strip along spine. Chin is sparsely furred blackish brown. Hairs on throat are drab, and central region of throat has exposed pale skin. Ventral parts of upper arms, shoulders, and flanks are drab, with median patch of Sudan brown on breast and belly. Undersides of propatagium and plagiopatagium have Sudan brown hairs near forearm and between upper arm and body. Claws are light yellowish brown but darkest basally. Index claw (second digit of wing) is absent. Ears, wings, and naked dorsal skin are blackish brown. Nostrils are slightly tubular, and each flares outward from centerline.	Primary and secondary tropical moist forests, pandanus palms, coconut and cocoa plantations, and village gardens.	Beaufort’s Naked-backed Fruit Bat is frugivorous. Native cluster figs ( Ficus spp. , Moraceae ) and invasive Piper aduncum ( Piperaceae ) are included and probably quite important in its diet. It forages on carambolas ( Oxalidaceae ) and other fruits in village gardens.	Seven of 14 Beaufort’s Naked-backed Fruit Bats examined from Waigeo in December were lactating or pregnant. Females observed on Biak in September were in the first trimester of pregnancy. On Batanta, many pregnant females were observed in October.	Beaufort’s Naked-backed Fruit Bat is nocturnal and roosts in sea and inland caves.	Beaufort’s Naked-backed Fruit Bat is highly gregarious, sometimes roosting in large colonies of thousands of individuals. It will also roost in smaller groups of tens of individuals in trees.	Classified as Least Concern on The IUCN Red List. Beaufort’s Naked-backed Fruit Bat is distributed on a number of islands, presumably has a large population, and is unlikely to be declining at a rate required to qualify for listing in a threatened category. Cave disturbanceis a potential threat. Research is needed on extent of occurrence throughout its distribution and monitoring of population sizes over time to establish whether numbers are stable or experiencing decline. Studies also are needed on habitat requirements, food habits, and effects of forest loss and degradation. Additional taxonomic studies are needed to determine how Beaufort’s Nakedbacked Fruit Bat might be related (or even if some populations thought to represent it) or conspecific with closely related Greenish Naked-backed Fruit Bat (D. viridas).	Bergmans (1975) | Flannery (1995a) | Maryanto & Kitchener (1999) | Mildenstein (2016d)		91. Beaufort’s Naked-backed Fruit Bat Dobsonia beauforti French: Roussette de Beaufort / German: \Waigeo-Nacktriickenflughund / Spanish: Dobsonia de Beaufort Other common names: Beaufort's Bare-backed Fruit Bat Taxonomy. Dobsonia beauforti Bergmans, 1975 , “from a cave near Nja-njef, Waigeo [ Island , Sorong Division, Papua Province], New Guinea ,” Indonesia . This species is monotypic. Distribution. Raja Ampat Is (Gebe, Gag, Waigeo, and Batanta), Schouten Is (Biak-Supiori and Owi) off NWNew Guinea ;it possibly may be found on MisoolI, a single individual reported from Salawati I probably represents a vagrant. Descriptive notes. Head-body 120-155 mm , tail 20-28 mm , ear 23-27 mm , forearm 100-114 mm ; weight 116-163 g . Males are appreciably larger than females in mean weight (males 158 g and females 133 g ) and slightly larger in mean forearm length (males 108 mm and females 106 mm ). Breeding males differ from females in having brighter greenish yellow fur, proportionately more massive heads with prominent white cheek glands, and almost naked skin on rest of face. Females tend to be more brownish, have more gracile head that is more heavily furred, and lack inflated cheek glands. Young have green-yellow wash to fur, more similar to adult males. Fur on top and side of head is bone brown to olive-brown (greenish yellow in some descriptions). Nape, shoulders, and furred parts of back are olive-brown and somewhat darker in occipital region and bordering naked skin including narrow strip along spine. Chin is sparsely furred blackish brown. Hairs on throat are drab, and central region of throat has exposed pale skin. Ventral parts of upper arms, shoulders, and flanks are drab, with median patch of Sudan brown on breast and belly. Undersides of propatagium and plagiopatagium have Sudan brown hairs near forearm and between upper arm and body. Claws are light yellowish brown but darkest basally. Index claw (second digit of wing) is absent. Ears, wings, and naked dorsal skin are blackish brown. Nostrils are slightly tubular, and each flares outward from centerline. Habitat. Primary and secondary tropical moist forests, pandanus palms, coconut and cocoa plantations, and village gardens. Food and Feeding. Beaufort’s Naked-backed Fruit Bat is frugivorous. Native cluster figs ( Ficus spp. , Moraceae ) and invasive Piper aduncum ( Piperaceae ) are included and probably quite important in its diet. It forages on carambolas ( Oxalidaceae ) and other fruits in village gardens. Breeding. Seven of 14 Beaufort’s Naked-backed Fruit Bats examined from Waigeo in December were lactating or pregnant. Females observed on Biak in September were in the first trimester of pregnancy. On Batanta, many pregnant females were observed in October. Activity patterns. Beaufort’s Naked-backed Fruit Bat is nocturnal and roosts in sea and inland caves. Movements, Home range and Social organization. Beaufort’s Naked-backed Fruit Bat is highly gregarious, sometimes roosting in large colonies of thousands of individuals. It will also roost in smaller groups of tens of individuals in trees. Status and Conservation. Classified as Least Concern on The IUCN Red List. Beaufort’s Naked-backed Fruit Bat is distributed on a number of islands, presumably has a large population, and is unlikely to be declining at a rate required to qualify for listing in a threatened category. Cave disturbanceis a potential threat. Research is needed on extent of occurrence throughout its distribution and monitoring of population sizes over time to establish whether numbers are stable or experiencing decline. Studies also are needed on habitat requirements, food habits, and effects of forest loss and degradation. Additional taxonomic studies are needed to determine how Beaufort’s Nakedbacked Fruit Bat might be related (or even if some populations thought to represent it) or conspecific with closely related Greenish Naked-backed Fruit Bat (D. viridas). Bibliography. Bergmans (1975), Flannery (1995a), Maryanto & Kitchener (1999), Mildenstein (2016d).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Dobsonia beauforti	Dobsonia		beauforti	Bergmans	1975	0	Beaufortia	23(295): 3	Beaufort&apos;s Naked-backed Fruit Bat	None.	Indonesia, Prov. of Papua, Sorong Div., Waigeo Isl, Njanjef.	Waigeo, Batanta, Salawati, Gebe, Gag, and Biak Isls (off Vogelkop Peninsula, New Guinea).	Not listed.	Least Concern	 viridis species group. Closely related to viridis ; see Bergmans (1975). Also see Flannery (1995).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Dobsonia beauforti	23	Beaufort's Naked-backed Fruit Bat	Beaufort's Bare-backed Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	HARPYIONYCTERINAE	DOBSONIINI	Dobsonia	NA	beauforti	Bergmans	1975	0						"from a cave near Nja-njef, Waigeo [Island, Sorong Division, Papua Province], New Guinea," Indonesia.			beauforti Bergmans, 1975	NA	NA	Indonesia	Oceania	Australasia/Oceania	LC	0	0	0	Dobsonia_beauforti	0	sciname match	Dobsonia_beauforti	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	6772	Dobsonia beauforti	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Dobsonia	beauforti	Bergmans, 1975	This may possibly be conspecific with Dobsonia viridis .	20000000	Dobsonia beauforti	Least Concern		2016	2015-04-20 00:00:00 UTC	3.1	English	Listed as Least Concern because it is distributed on a number of islands, presumably has large population, occurs in a number of protected areas, and it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	This species roosts in large colonies in both caves and trees. It is commonly found foraging in secondary forest, Pandanus, plantations, and gardens. Flannery (1995) thought that the similar -sized Dobsonia emersa excludes it from caves where the two species co-occur. Of the 14 specimens in the type series collected in December 1909 from Waigeo, seven females were lactating or pregnant. Females observed on Biak-Supiori in September were in an early stage of pregnancy, while on Batanta many pregnant females were observed in October (Flannery1995).	There appear to be no major threats to this species. It is unlikely to be hunted within this range (F. Bonaccorso pers. comm.).	This species is widespread and common within its range.	Unknown	This species is endemic to the Raja Ampat Islands (Batanta, Gebe, Salawati, Gag, Waigeo (plus possibly Misool)), and the Biak and Supiori islands (including Owi) in Geelvinck Bay, Indonesia. It is not present on the island of New Guinea (Flannery 1995).		Terrestrial	It is present in a number of protected areas. <u> Research needs</u> Studies are needed on the species’ population sizes, distribution, and extent of occurrence throughout its range. Monitoring of population sizes and locations over time are also important to establish whether these are stable or experiencing trends of decline. The threats to these bats are poorly understood. Studies are needed on the species’ habitat requirements and on the effects of forest loss and degradation on the species’ population sizes/distribution. Further taxonomic studies are needed to determine if this species is conspecific with Dobsonia viridis. 	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Dobsonia		beauforti	Bergmans	1975	0	Beaufortia	23(295): 3	Beaufort&apos;s Naked-backed Fruit Bat	None.	Indonesia, Prov. of Papua, Sorong Div., Waigeo Isl, Njanjef.	Waigeo, Batanta, Salawati, Gebe, Gag, and Biak Isls (off Vogelkop Peninsula, New Guinea).	Not listed.	Least Concern	 viridis species group. Closely related to viridis ; see Bergmans (1975). Also see Flannery (1995).	Dobsonia beauforti	1004392	23	Beaufort's Naked-backed Fruit Bat	Beaufort's Bare-backed Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	HARPYIONYCTERINAE	DOBSONIINI	Dobsonia	NA	beauforti	Bergmans	1975	0						"from a cave near Nja-njef, Waigeo [Island, Sorong Division, Papua Province], New Guinea," Indonesia.			beauforti Bergmans, 1975	NA	NA				Indonesia	Oceania	Australasia/Oceania	LC	0	0	0	Dobsonia_beauforti	0	sciname match	Dobsonia_beauforti	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Dobsonia_beauforti	1004392	23	Beaufort's Naked-backed Fruit Bat	Beaufort's Bare-backed Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Harpyionycterinae	Dobsoniini	Dobsonia	NA	beauforti	Bergmans	0	Dobsonia beauforti	Bergmans, W. 1975-01-16. A new species of _Dobsonia_ Palmer, 1898 (Mammalia, Megachiroptera) from Waigeo, with notes on other members of the genus. Beaufortia 23(295):1-13.		ZMA.MAM.16476.a, ZMA.MAM.16476.b	holotype	https://data.biodiversitydata.nl/naturalis/specimen/ZMA.MAM.16476.a | https://data.biodiversitydata.nl/naturalis/specimen/ZMA.MAM.16476.b	"from a cave near Nja-njef, Waigeo [Island, Sorong Division, Papua Province], New Guinea," Indonesia.	-0.33333	130.7167	NA	NA				Indonesia	Oceania (Continent)	Australasia	LC	0	0	0	Dobsonia_beauforti	0	sciname match	Dobsonia_beauforti	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Dobsonia		beauforti	Bergmans	1975	0	Beaufortia	23(295): 3	Beaufort's Naked-backed Fruit Bat	None.	Indonesia, Prov. of Papua, Sorong Div., Waigeo Isl, Njanjef	Waigeo, Batanta, Salawati, Gebe, Gag, possibly Misool, and Biak Isls (off Vogelkop Peninsula, New Guinea)	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/6772/22034699/' target='_blank'>Least Concern</a>	viridis species group. Closely related to viridis; see Bergmans (1975). Bonaccorso (2019) suggests that the record from Sulawati represents a vagrant and that this species may be present on Misool Island as well. Also see Flannery (1995).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Dobsonia beauforti; Dobsonia beauforti; Dobsonia beauforti; Dobsonia beauforti; Dobsonia beauforti; Dobsonia beauforti; beauforti; Roussette de Beaufort; \Waigeo-Nacktriickenflughund; Dobsonia de Beaufort; Beaufort's Bare-backed Fruit Bat; Beaufort's Naked-backed Fruit Bat; Beaufort's Bare-backed Fruit Bat; Beaufort’s Naked-backed Fruit Bat; Beaufort&apos;s Naked-backed Fruit Bat; D. beauforti