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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L232	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Diclidurus scutatus	Diclidurus scutatus	Diclidurus scutatus	Diclidurus scutatus	Diclidurus scutatus	Diclidurus scutatus	Diclidurus scutatus	Diclidurus scutatus	Diclidurus scutatus	Diclidurus scutatus	Diclidurus scutatus	Diclidurus scutatus	Diclidurus scutatus	Diclidurus scutatus	Diclidurus scutatus		[MSW2] Subgenus Diclidurus.; [MSW3] Subgenus Diclidurus.; [HMW] Diclidurus scutatus Peters, 1869 , “Südamerica.” Restricted by A. M. Husson in 1962 to “Para [= Belém] , Para , Brazil . Diclidurus scutatus is in the subgenus Depanycteris . Monotypic.; [batnames2022] Subgenus Diclidurus .; [batnames2023] Subgenus Diclidurus .; [batnames2025_1.7] Subgenus Diclidurus.														scutatus				scutatus	scutatus, scuttatus			scutatus W. C. H. Peters, 1869|scuttatus SodrÃ© & Uieda, 2006 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Venezuela – Surinam, Peru, Brazil	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Diclidurus scutatus	Brazil, Para, Belem.	Peters	1869	Monatsb. Preuss. Akad. Wiss. Berlin, p. 400.	Distribution: Con fined to southern Venezuela, the Guianas, and Amazonian parts of Peru and Brazil.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Venezuela – Surinam, Peru, Brazil	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Peters	1869	Monatsb. K. Preuss. Akad. Wiss. Berlin, 1869:400.	Subgenus Diclidurus.	Amazonian Brazil, Venezuela, Peru, Guyana, Surinam.	Brazil, Para, Belem.		PETERS	1869	Size relatively small (forearm length, 51-58 mm). Upper incisor with anterior and posterior cingular secondary cusps. Upper premolars in contact.	Distribution: Con fined to southern Venezuela, the Guianas, and Amazonian parts of Peru and Brazil.	No sub species.		48	species	D. scutatus	PETERS	1869	Diclidurus	subgenus	Diclidurus scutatus				Size relatively small (forearm length, 51-58 mm). Upper incisor with anterior and posterior cingular secondary cusps. Upper premolars in contact.	No sub species.		2. D. scutatus PETERS 1869.	2	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Emballonuridae	Emballonurinae		Diclidurus scutatus	Diclidurus	Diclidurus	scutatus	Peters		1869		Monatsb. K. Preuss. Akad. Wiss. Berlin	1869		400		Lesser Ghost Bat	Brazil, Pará, Belem.	Amazonian Brazil, Venezuela, Peru, Guyana, Surinam, French Guiana.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).		Subgenus Diclidurus.	03D587F2FFDC4C17FF7D36B4FDB6F385	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Emballorunidae.pdf.imd	hash://md5/ffecff8affcf4c04ffa53577fff8ffe9	368	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/D5/87/03D587F2FFDC4C17FF7D36B4FDB6F385.xml	Diclidurus scutatus	Emballonuridae	Diclidurus	scutatus	Peters	1869	Lesser Ghost Bat @en | Diclidure des cyclanthes @fr | Kleine Gespenstfledermaus @de | Dicliduro pequeno @es	Diclidurus scutatus Peters, 1869 , “Südamerica.” Restricted by A. M. Husson in 1962 to “Para [= Belém] , Para , Brazil . Diclidurus scutatus is in the subgenus Depanycteris . Monotypic.	Amazonian Basin in NE Ecuador (Orellana Province ), NE Peru (Loreto Department), E Colombia (Vaupés Department), S & W Venezuela, the Guianas, and N Brazil (Amazonas, Amapa, and Para states), with one additional record from Atlantic Dry Forest of SE Brazil (Sâo Paulo State).	Head—body 57-68 mm, tail 17—19 mm, ear 10—15 mm, hindfoot 7-11 mm, forearm 51-58-9 mm; weight c.13 g. Dorsal fur of the Lesser Ghost Bat is long, soft, and either pure white or pale brown at bases of hairs; venter is uniformly white. Claws are black. Flight membranes are unpigmented. Naked skin of wings and legs are pink. Eyes are relatively large, with dark brown irises. Adult males have two glandular pouches on their uropatagia that are particularly prominent during breeding season. Ear is yellowish, rounded at apex, and c.15 mm long; tragus is.6 mm. Greatest length of skull is c.15 mm. Diagnostic skull characteristics include large palatine foramina, with posterior palatal emargination, and no space between premolars.	Multi-strata evergreen forests (rainforest), Atlantic Dry Forest (one record), and towns and villages from sea level to elevations of c.1000 m.	The Lesser Ghost Bat preys on insects. One specimen was collected with a large moth in its mouth, and another was captured in a canopy mist net.	No information.	Lesser Ghost Bats are crepuscular and fly high in open spaces above rivers, streams, lagoons, towns, and forest canopies. They are attracted to insects around lights.	No information.	Classified as Least Concern on The IUCNRed List. The Lesser Ghost Bat has a large distribution in areas with intact evergreen forests. Nevertheless, little is known about its taxonomy, distribution, and ecology. Conservation actions should include scientific studies of distribution, roosting behavior, reproduction, diet, and foraging behavior.	Albuja (1999) | Carter & Dolan (1978) | Eisenberg (1989) | Escobedo &Velazco (2012) | Fonseca ef al. (1996) | Handley (1976) | Husson (1962, 1978) | Lim & Engstrom (2005) | Linares (1998) | Novaes, Clâudio & Kuniy (2017) | Tirira (2017)	https://zenodo.org/record/3747976/files/figure.png	40 . Lesser Ghost Bat Diclidurus scutatus French: Diclidure des cyclanthes / German: Kleine Gespenstfledermaus / Spanish: Dicliduro pequeno Taxonomy . Diclidurus scutatus Peters, 1869 , “Südamerica.” Restricted by A. M. Husson in 1962 to “Para [= Belém] , Para , Brazil . Diclidurus scutatus is in the subgenus Depanycteris . Monotypic. Distribution. Amazonian Basin in NE Ecuador (Orellana Province ), NE Peru (Loreto Department), E Colombia (Vaupés Department), S & W Venezuela, the Guianas, and N Brazil (Amazonas, Amapa, and Para states), with one additional record from Atlantic Dry Forest of SE Brazil (Sâo Paulo State). Descriptive notes. Head—body 57-68 mm, tail 17—19 mm, ear 10—15 mm, hindfoot 7-11 mm, forearm 51-58-9 mm; weight c.13 g. Dorsal fur of the Lesser Ghost Bat is long, soft, and either pure white or pale brown at bases of hairs; venter is uniformly white. Claws are black. Flight membranes are unpigmented. Naked skin of wings and legs are pink. Eyes are relatively large, with dark brown irises. Adult males have two glandular pouches on their uropatagia that are particularly prominent during breeding season. Ear is yellowish, rounded at apex, and c.15 mm long; tragus is.6 mm. Greatest length of skull is c.15 mm. Diagnostic skull characteristics include large palatine foramina, with posterior palatal emargination, and no space between premolars. Habitat . Multi-strata evergreen forests (rainforest), Atlantic Dry Forest (one record), and towns and villages from sea level to elevations of c.1000 m. Food and Feeding . The Lesser Ghost Bat preys on insects. One specimen was collected with a large moth in its mouth, and another was captured in a canopy mist net. Breeding . No information. Activity patterns. Lesser Ghost Bats are crepuscular and fly high in open spaces above rivers, streams, lagoons, towns, and forest canopies. They are attracted to insects around lights. Movements, Home range and Social organization. No information. Status and Conservation . Classified as Least Concern on The IUCNRed List. The Lesser Ghost Bat has a large distribution in areas with intact evergreen forests. Nevertheless, little is known about its taxonomy, distribution, and ecology. Conservation actions should include scientific studies of distribution, roosting behavior, reproduction, diet, and foraging behavior. Bibliography. Albuja (1999), Carter & Dolan (1978), Eisenberg (1989), Escobedo &Velazco (2012), Fonseca ef al. (1996), Handley (1976), Husson (1962, 1978), Lim & Engstrom (2005), Linares (1998), Novaes, Clâudio & Kuniy (2017), Tirira (2017).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Emballonuridae	Diclidurus scutatus	Diclidurus	Diclidurus	scutatus	Peters	1869	0	Monatsb. K. Preuss. Akad. Wiss. Berlin	1875:40:00	Lesser Ghost Bat	None.	Brazil, Par&aacute;, Belem.	Amazonian Brazil, Venezuela, Peru, Guyana, Surinam, French Guiana.	Not listed.	Least Concern	Subgenus Diclidurus .	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Diclidurus scutatus	23	Lesser Ghost Bat	Little Ghost Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	EMBALLONURIDAE	EMBALLONURINAE	DICLIDURINI	Diclidurus	Depanycteris	scutatus	W. Peters	1869	0	Diclidurus_scutatus	Peters, W. (1869). Bemerkungen Ã¼ber neue oder weniger bekannte Flederthiere, besonders des Pariser Museums. Monatsberichte der KÃ¶niglich Preussischen Akademie der Wissenschaften zu Berlin, 1869, 400.	https://www.biodiversitylibrary.org/item/111869#page/434/mode/1up	MNHN 722		"SÃ¼damerica." Restricted by A. M. Husson in 1962 to "ParÃ¡ [= BelÃ©m], ParÃ¡, Brazil.			scutatus W. Peters, 1869	NA	NA	Ecuador|Peru|Colombia|Venezuela|Guyana|Suriname|French Guiana|Brazil	South America	Neotropic	LC	0	0	0	Diclidurus_scutatus	0	sciname match	Diclidurus_scutatus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	6564	Diclidurus scutatus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	EMBALLONURIDAE	Diclidurus	scutatus	Peters, 1869		20000000	Diclidurus scutatus	Least Concern		2016	2016-07-01 00:00:00 UTC	3.1	English	This species is listed as Least Concern because it is relatively widely distributed although known only from few specimens and is unlikely to be declining fast enough to qualify for inclusion in any of the threat categories in the near future.	Ecology is little known. These bats fly high in open space, such as above rivers, streams, and lagoons, and above the forest canopy. They are attracted to concentrations of insects around high floodlights and are among the many species swooping around such lights, even in towns (Emmons and Feer 1997). They are associated with wet habitats and multistratal evergreen forest (Eisenberg 1989). Their roosts are not well known, perhaps they are found in or between the leaves of palm trees (Emmons and Feer 1997).	Deforestation is a localized threat.	The high roosting and flying behaviour of these bats makes detecting them extremely difficult, and it is not possible to say whether they are common or rare (Emmons and Feer 1997).	Unknown	This species is distributed in the Amazonia portions of Peru (Escobedo and Velazco 2012), Colombia (Escobedo and Velazco 2012), Ecuador (Albuja 1999), Brazil (Fonseca et al . 1996), adjacent Venezuela (Linares 1998), Guianas and Surinam (Lim and Engstron 2005). Many specimens have been taken below 200 m elevation (Hood and Gardner 2008) and upper elevation limit is below 1,000 m in Venezuela (Handley 1976).		Terrestrial	Retention of forest is needed.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Emballonuridae	Diclidurus	Diclidurus	scutatus	Peters	1869	0	Monatsb. K. Preuss. Akad. Wiss. Berlin	1875:40:00	Lesser Ghost Bat	None.	Brazil, Par&aacute;, Belem.	Amazonian Brazil, Venezuela, Peru, Guyana, Surinam, French Guiana.	Not listed.	Least Concern	Subgenus Diclidurus .	Diclidurus scutatus	1004786	23	Lesser Ghost Bat	Little Ghost Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	Emballonuridae	EMBALLONURINAE	DICLIDURINI	Diclidurus	Depanycteris	scutatus	W. Peters	1869	0	Diclidurus_scutatus	Peters, W. (1869). Bemerkungen Ã¼ber neue oder weniger bekannte Flederthiere, besonders des Pariser Museums. Monatsberichte der KÃ¶niglich Preussischen Akademie der Wissenschaften zu Berlin, 1869, 400.	https://www.biodiversitylibrary.org/item/111869#page/434/mode/1up	MNHN 722		"SÃ¼damerica." Restricted by A. M. Husson in 1962 to "ParÃ¡ [= BelÃ©m], ParÃ¡, Brazil.			scutatus W. Peters, 1869	NA	NA				Ecuador|Peru|Colombia|Venezuela|Guyana|Suriname|French Guiana|Brazil	South America	Neotropic	LC	0	0	0	Diclidurus_scutatus	0	sciname match	Diclidurus_scutatus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Diclidurus_scutatus	1004786	23	Lesser Ghost Bat	Little Ghost Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Emballonuroidea	Emballonuridae	Emballonurinae	Diclidurini	Diclidurus	NA	scutatus	W. C. H. Peters	0	Diclidurus scutatus	Peters, W.C.H. 1869. Bemerkungen Ã¼ber neue oder weniger bekannte Flederthiere, besonders des Pariser Museums. Monatsberichte der KÃ¶niglichen Preussischen Akademie der Wissenschaften zu Berlin 1869:393-408.	https://www.biodiversitylibrary.org/page/36276613	MNHN-ZM-MO-1859-340 (= MNHN ? 722)	holotype	http://coldb.mnhn.fr/catalognumber/mnhn/zm/mo-1859-340	"SÃ¼damerica." Restricted by A. M. Husson in 1962 to "ParÃ¡ [= BelÃ©m], ParÃ¡, Brazil.			NA	NA				Ecuador|Peru|Colombia|Venezuela|Guyana|Suriname|French Guiana|Brazil	South America	Neotropic	LC	0	0	0	Diclidurus_scutatus	0	sciname match	Diclidurus_scutatus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Emballonuridae	Diclidurus	Diclidurus	scutatus	Peters	1869	0	Monatsb. K. Preuss. Akad. Wiss. Berlin	1875:40:00	Lesser Ghost Bat	None.	Brazil, Par&aacute;, Belem.	Amazonian Brazil, Venezuela, Peru, Guyana, Surinam, French Guiana.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/6564/21986499/' target='_blank'>Least Concern</a>	Subgenus Diclidurus.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Diclidurus scutatus; Diclidurus scutatus; Diclidurus scutatus; Diclidurus scutatus; Diclidurus scutatus; Diclidurus scutatus; scutatus; Lesser Ghost Bat; Diclidure des cyclanthes; Kleine Gespenstfledermaus; Dicliduro pequeno; Lesser Ghost Bat; Little Ghost Bat; Lesser Ghost Bat; Lesser Ghost Bat; D. scutatus
