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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L230	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Diclidurus ingens	Diclidurus ingens	Diclidurus ingens	Diclidurus ingens	Diclidurus ingens	Diclidurus ingens	Diclidurus ingens	Diclidurus ingens	Diclidurus ingens	Diclidurus ingens	Diclidurus ingens	Diclidurus ingens	Diclidurus ingens	Diclidurus ingens	Diclidurus ingens		[MSW2] Subgenus Diclidurus.; [MSW3] Subgenus Diclidurus.; [HMW] Diclidurus ingens Hemândez- Camacho, 1955 , “ Puerto Legufzamo, Intendencia del Caqueta [= Putumayo] ,” Colombia . Diclidurus ingens is in the subgenus Diclidurus . Monotypic.; [batnames2022] Subgenus Diclidurus .; [batnames2023] Subgenus Diclidurus .; [batnames2025_1.7] Subgenus Diclidurus.														ingens				ingens	ingens			ingens Hernández-Camacho, 1955		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Colombia, Venezuela	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Diclidurus ingens	Colombia, Caqueta, Rio Putumayo, Puerto Leguizamo.	Hernandez-Camacho	1955	Caldasia, 7:87.	Distribution: Known only from eastern Colombia, southern and eastern Venezuela, Guyana, and western Amazonian Brazil.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Colombia – Guyana, NW Brazil	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Hernandez-Camacho	1955	Caldasia, 7:87.	Subgenus Diclidurus.	Venezuela, SE Colombia, Guyana, NW Brazil.	Colombia, Caqueta, Rio Putumayo, Puerto Leguizamo.		HERNANDEZ CAMACHO	1955	Size relatively large (forearm length, 70-7 3 mm). Up per incisor with prominent posterior secondary cusp. Upper premolars separate.	Distribution: Known only from eastern Colombia, southern and eastern Venezuela, Guyana, and western Amazonian Brazil.	No subspecies.		48	species	D. ingens	HERNANDEZ CAMACHO	1955	Diclidurus	subgenus	Diclidurus ingens				Size relatively large (forearm length, 70-7 3 mm). Up per incisor with prominent posterior secondary cusp. Upper premolars separate.	No subspecies.		4. D. ingens HERNANDEZ CAMACHO 1955.	4	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Emballonuridae	Emballonurinae		Diclidurus ingens	Diclidurus	Diclidurus	ingens	Hernandez-Camacho		1955		Caldasia	7		87		Greater Ghost Bat	Colombia, Caqueta, Río Putumayo, Puerto Leguizamo.	Venezuela, SE Colombia, Guyana, NW Brazil.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.		Subgenus Diclidurus.	03D587F2FFDD4C17F8F53FA8F921FCBA	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Emballorunidae.pdf.imd	hash://md5/ffecff8affcf4c04ffa53577fff8ffe9	367	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/D5/87/03D587F2FFDD4C17F8F53FA8F921FCBA.xml	Diclidurus ingens	Emballonuridae	Diclidurus	ingens	Hemândez- Camacho	1955	Greater Ghost Bat @en | Grande Diclidure @fr | Grosse Gespenstfledermaus @de | Dicliduro grande @es	Diclidurus ingens Hemândez- Camacho, 1955 , “ Puerto Legufzamo, Intendencia del Caqueta [= Putumayo] ,” Colombia . Diclidurus ingens is in the subgenus Diclidurus . Monotypic.	Scattered localities from Colombia (Chocó and Putumayo departments) and adjacent Venezuela to Guyana, extending S to N & C Brazil (Para and Mato Grosso). Future research likely will extend distributional limits and establish occurrence in Peru and Ecuador.	Head-body c.83 mm, tail 22 mm, ear 14 mm, hindfoot 12 mm, forearm 67-9-71-9 mm; weight 20-26 g (most measurements are from a single specimen). Dorsal and ventral fur of the Greater Ghost Bat is composed of bicolored hairs that are cream to pale gray at bases and gray to blackish distally . Flight membranes are unpigmented and appear pinkish from heavy vascularization. Bare skin of face and ears also appears pinkish. Eyes are large and conspicuous, with dark brown irises. An inter-femoral sac is present along tail. P1 and P2 are not in contact, which is diagnostic . Presence of four lobes on the I2 of this species is diagnostic and unique among emballonurids.	Vertically stratified primary and secondary evergreen forests and banana plantations from sea level to elevations of c.200 m.	Greater Ghost Bats prey on insects high in open spaces above rivers, streams, and lagoons and above forest canopies. They are attracted to concentrations of insects around streetlights in towns. On following pages: 40. Lesser Ghost Bat ( Diclidurus scutatus ); 41. Isabelline Ghost Bat ( Diclidurus isabella ); 42. Lesser Dog-like Bat ( Peropteryx macrotis ); 43. Greater Dog-like Bat ( Peropteryx kappleri ); 44. Trinidad Dog-like Bat ( Peropteryx trinitatis ); 45. White-winged Dog-like Bat ( Peropteryx leucoptera ); 46. Pale-winged Dog-like Bat ( Peropteryx pallidoptera ); 47. Thomas's Shaggy Bat ( Centronycteris centrali); 48. Common Shaggy Bat ( Centronycteris maximiliani ); 49. Proboscis Bat ( Rhynchonycteris naso ); 50. Greater Sac-winged Bat ( Saccopteryx bilineata }; 51. Lesser Sac-winged Bat ( Saccopteryx leptura ); 52. rosted Sac-winged Bat ( Saccopteryx canescens ); 53. Antioquian Sac-winged Bat ( Saccopteryx antioquensis ); 54. Amazonian Sac-winged Bat ( Saccopteryx gymnura ).	No information.	No information.	The only roosting record of the Greater Ghost Bat is of a solitary female hanging from the ceiling of a house in a suburban area in Colombia.	Classified as Data Deficient on The IUCN Red List. The Greater Ghost Bat is known from very few specimens and few locations. Information on distribution, status, and conservation threats is lacking.	Dalponte & Aguiar (2009) | Eisenberg (1989) | Emmons & eer (1997) | Hood & Gardner (2008) | Lim, B.K. et al. (1999) | Mantilla-Meluk, Jiménez-Ortega, Palacios & Baker (2009) | Ojasti & Linares (1971)	https://zenodo.org/record/3740275/files/figure.png	39 . Greater Ghost Bat Diclidurus ingens French: Grande Diclidure / German: Grosse Gespenstfledermaus / Spanish: Dicliduro grande Taxonomy . Diclidurus ingens Hemândez- Camacho, 1955 , “ Puerto Legufzamo, Intendencia del Caqueta [= Putumayo] ,” Colombia . Diclidurus ingens is in the subgenus Diclidurus . Monotypic. Distribution. Scattered localities from Colombia (Chocó and Putumayo departments) and adjacent Venezuela to Guyana, extending S to N & C Brazil (Para and Mato Grosso). Future research likely will extend distributional limits and establish occurrence in Peru and Ecuador. Descriptive notes. Head-body c.83 mm, tail 22 mm, ear 14 mm, hindfoot 12 mm, forearm 67-9-71-9 mm; weight 20-26 g (most measurements are from a single specimen). Dorsal and ventral fur of the Greater Ghost Bat is composed of bicolored hairs that are cream to pale gray at bases and gray to blackish distally . Flight membranes are unpigmented and appear pinkish from heavy vascularization. Bare skin of face and ears also appears pinkish. Eyes are large and conspicuous, with dark brown irises. An inter-femoral sac is present along tail. P1 and P2 are not in contact, which is diagnostic . Presence of four lobes on the I2 of this species is diagnostic and unique among emballonurids. Habitat . Vertically stratified primary and secondary evergreen forests and banana plantations from sea level to elevations of c.200 m. Food and Feeding . Greater Ghost Bats prey on insects high in open spaces above rivers, streams, and lagoons and above forest canopies. They are attracted to concentrations of insects around streetlights in towns. On following pages: 40. Lesser Ghost Bat ( Diclidurus scutatus ); 41. Isabelline Ghost Bat ( Diclidurus isabella ); 42. Lesser Dog-like Bat ( Peropteryx macrotis ); 43. Greater Dog-like Bat ( Peropteryx kappleri ); 44. Trinidad Dog-like Bat ( Peropteryx trinitatis ); 45. White-winged Dog-like Bat ( Peropteryx leucoptera ); 46. Pale-winged Dog-like Bat ( Peropteryx pallidoptera ); 47. Thomas's Shaggy Bat ( Centronycteris centrali); 48. Common Shaggy Bat ( Centronycteris maximiliani ); 49. Proboscis Bat ( Rhynchonycteris naso ); 50. Greater Sac-winged Bat ( Saccopteryx bilineata }; 51. Lesser Sac-winged Bat ( Saccopteryx leptura ); 52. rosted Sac-winged Bat ( Saccopteryx canescens ); 53. Antioquian Sac-winged Bat ( Saccopteryx antioquensis ); 54. Amazonian Sac-winged Bat ( Saccopteryx gymnura ). Breeding . No information. Activity patterns. No information. Movements, Home range and Social organization. The only roosting record of the Greater Ghost Bat is of a solitary female hanging from the ceiling of a house in a suburban area in Colombia. Status and Conservation . Classified as Data Deficient on The IUCN Red List. The Greater Ghost Bat is known from very few specimens and few locations. Information on distribution, status, and conservation threats is lacking. Bibliography. Dalponte & Aguiar (2009), Eisenberg (1989), Emmons & eer (1997), Hood & Gardner (2008), Lim, B.K. et al. (1999), Mantilla-Meluk, Jiménez-Ortega, Palacios & Baker (2009), Ojasti & Linares (1971).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Emballonuridae	Diclidurus ingens	Diclidurus	Diclidurus	ingens	Hernandez-Camacho	1955	0	Caldasia	0.3521	Greater Ghost Bat	None.	Colombia, Caqueta, R&iacute;o Putumayo, Puerto Leguizamo.	Venezuela, SE Colombia, Guyana, NW Brazil.	Not listed.	Data Deficient	Subgenus Diclidurus .	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Diclidurus ingens	23	Greater Ghost Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	EMBALLONURIDAE	EMBALLONURINAE	DICLIDURINI	Diclidurus	Diclidurus	ingens	Hernández-Camacho	1955	0	Diclidurus_ingens	Hernández-Camacho, J. (1955). Una nueva especie Colombiana del genero Diclidurus (Mammalia: Chiroptera): Diclidurus ingens. Caldasia, 7(31), 87.	https://www.jstor.org/stable/44240739?refreqid=excelsior%3A6f7ac4c472fcecd4f04757c797055762&seq=1#metadata_info_tab_contents	ICN 546		"Puerto Leguízamo, Intendencia del Caquetá [= Putumayo]," Colombia.			ingens Hernández-Camacho, 1955	NA	NA	Colombia|Venezuela|Guyana|Brazil	South America	Neotropic	DD	0	0	0	Diclidurus_ingens	0	sciname match	Diclidurus_ingens	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	6562	Diclidurus ingens	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	EMBALLONURIDAE	Diclidurus	ingens	Hernandez-Camacho, 1955		20000000	Diclidurus ingens	Data Deficient		2016	2016-06-27 00:00:00 UTC	3.1	English	Listed as Data Deficient because, although it is known from very few specimens and few locations - there is not enough information on the distribution, status or threats to this species to make an assessment.	Ecology is little known. These bats fly high in open space, such as above rivers, streams, and lagoons, and above the forest canopy. They are attracted to concentrations of insects around high floodlights and are among the many species swooping around such lights, even in towns (Emmons and Feer 1997). It is associated with multistratal evergreen forest and prefers moist areas (Eisenberg 1989). Their roosts are not well known; perhaps they are found in abandoned towers, or between the leaves of palm trees (Emmons and Feer 1997). Maybe well adapted to human habitation.	This is one of the least known species in the genus, however its distribution has been recently updated with new geographic records. However, it is still known from very few specimens, probably suggesting its restricted distribution is associated to specific habitat and roost requirements.	They are known from only a few specimens, and this is the least known species in the genus. The high roosting and flying behaviour of these bats makes detecting them extremely difficult, and it is not possible to say whether they are common or rare or to estimate population size.	Unknown	The species ranges from southeastern Colombia and adjacent Venezuela to Guyana, extending south to northwestern Brazil (Hood and Gardner 2008). Recently, its distribution has extended to Mato Grosso, central Brazil (Dalponte and Aguiar 2009) and the Choco of Colombia (Mantilla-Meluk et al. 2009). Many specimens have been taken below 200 m elevation (Eisenberg 1989).		Terrestrial	This specis is listed as the least known species of the genus, thus research actions should be taken.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Emballonuridae	Diclidurus	Diclidurus	ingens	Hernandez-Camacho	1955	0	Caldasia	0.352083	Greater Ghost Bat	None.	Colombia, Caqueta, R&iacute;o Putumayo, Puerto Leguizamo.	Venezuela, SE Colombia, Guyana, NW Brazil.	Not listed.	Data Deficient	Subgenus Diclidurus .	Diclidurus ingens	1004784	23	Greater Ghost Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	Emballonuridae	EMBALLONURINAE	DICLIDURINI	Diclidurus	Diclidurus	ingens	Hernández-Camacho	1955	0	Diclidurus_ingens	Hernández-Camacho, J. (1955). Una nueva especie Colombiana del genero Diclidurus (Mammalia: Chiroptera): Diclidurus ingens. Caldasia, 7(31), 87.	https://www.jstor.org/stable/44240739?refreqid=excelsior%3A6f7ac4c472fcecd4f04757c797055762&seq=1#metadata_info_tab_contents	ICN 546		"Puerto Leguízamo, Intendencia del Caquetá [= Putumayo]," Colombia.			ingens Hernández-Camacho, 1955	NA	NA				Colombia|Venezuela|Guyana|Brazil	South America	Neotropic	DD	0	0	0	Diclidurus_ingens	0	sciname match	Diclidurus_ingens	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Diclidurus_ingens	1004784	23	Greater Ghost Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Emballonuroidea	Emballonuridae	Emballonurinae	Diclidurini	Diclidurus	NA	ingens	Hernández-Camacho	0	Diclidurus ingens	Hernández-Camacho, J.I. 1955. Una nueva especie colombiana del genero _Diclidurus_ (Mammalia: Chiroptera): _Diclidurus ingens_. Caldasia 7(31):87-98.	https://www.jstor.org/stable/44240739	ICN 546	holotype		"Puerto Leguízamo, Intendencia del Caquetá [= Putumayo]," Colombia.			NA	NA				Colombia|Venezuela|Guyana|Brazil	South America	Neotropic	DD	0	0	0	Diclidurus_ingens	0	sciname match	Diclidurus_ingens	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Emballonuridae	Diclidurus	Diclidurus	ingens	Hernandez-Camacho	1955	0	Caldasia	0.352083	Greater Ghost Bat	None.	Colombia, Caqueta, R&iacute;o Putumayo, Puerto Leguizamo.	Venezuela, SE Colombia, Guyana, NW Brazil.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/6562/21986793/' target='_blank'>Data Deficient</a>	Subgenus Diclidurus.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Diclidurus ingens; Diclidurus ingens; Diclidurus ingens; Diclidurus ingens; Diclidurus ingens; Diclidurus ingens; ingens; Greater Ghost Bat; Grande Diclidure; Grosse Gespenstfledermaus; Dicliduro grande; Greater Ghost Bat; Greater Ghost Bat; Greater Ghost Bat; D. ingens