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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L225	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Desmalopex microleucopterus	Desmalopex microleucopterus	Desmalopex microleucopterus	Desmalopex microleucopterus	Desmalopex microleucopterus	Desmalopex microleucoptera	Desmalopex microleucoptera	Desmalopex microleucoptera	Desmalopex microleucoptera		[HMW] Desmalopex microleucopterus Esselstyn et al, 2008 , “Mount Siburan, Batong Buhay, Sablayan, Occidental Mindoro , Mindoro Island, Philippines , approximately 100 m above sea level ( 12.83488 N . 120.93028 E ).” This species is monotypic.; [batnames2022]  Pteropus sp. A of Heaney et al. (1998); [MDD2022] recently described; [batnames2023]  Pteropus sp. A of Heaney et al. (1998); [MDD2023] recently described; usually spelt 'microleucopterus', but the name is an adjective and Desmalopex is feminine, so the name should be spelt 'microleucoptera'; [MDD2025_2.0] recently described; usually spelt 'microleucopterus', but the name is an adjective and Desmalopex is feminine, so the name should be spelt 'microleucoptera'; [batnames2025_1.7] Pteropus sp. A of Heaney et al. (1998). We have changed the spelling of the species epithet, originally microleucopterus, to agree with the feminine genus name, Desmalopex; see Zijlstra (2023).; [MDD2025_2.2] recently described; usually spelt 'microleucopterus', but the name is an adjective and Desmalopex is feminine, so the name should be spelt 'microleucoptera'														microleucopterus				microleucoptera	microleucoptera			microleucoptera Esselstyn, Garcia, Saulog, & Heaney, 2008						N/A																																								NA																											03AD87FAFF80F66E89B43DCBF78DF9B1	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	126	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFF80F66E89B43DCBF78DF9B1.xml	Desmalopex microleucopterus	Pteropodidae	Desmalopex	microleucopterus	Esselstyn et al.	2008	Roussette microptére @fr | Mindoro-Wei Rfllgelflughund @de | Desmalopex de Mindoro @es	Desmalopex microleucopterus Esselstyn et al, 2008 , “Mount Siburan, Batong Buhay, Sablayan, Occidental Mindoro , Mindoro Island, Philippines , approximately 100 m above sea level ( 12.83488 N . 120.93028 E ).” This species is monotypic.	N Philippines (Mindoro).	Head-body 133- 155 mm (tailless), hindfoot 29-34 mm , forearm 97-103 mm ; weight 129-156 g . Muzzle of the Mindoro Pallid Flying Fox is long, almost hairless, and whitish brown to pinkish; nostrils are large and divergent; and philtrum is undivided. Eyes are large, with orange irises. Ears are moderately long, pale brown, and narrowly rounded at tips. Body pelage is pale brown, soft, relatively long, and woollier in mantle. Uropatagium is narrow in center, tibia is sparsely furred, calcar is small, and index claw is present. Wing membranes are brown, with white blotches rather uniformly distributed on entire wing and slightly more intensely mottled in white at wingtips. Skull is typical pteropine, with pronounced basicranial deflection. Laterally, rostrum is long and tapering; nasal process of premaxilla is wide and procumbent; orbit is large, and rim is complete, with thin postorbital process joined to spine of thin arched zygoma; and zygomatic rootisjust above upper alveolar line. Dorsally, rostrum is long and rectangular, nasals are salient, paranasal recesses are moderately inflated reaching small postorbital foramina, postorbital constriction is moderately marked, sagittal crest is absent, braincase is oval, and nuchal crest is barely marked. Ventrally, palate is very long and flat; tooth rows are nearly parallel; post-dental palate is short, ending in deep concavity; ectopterygoids are comparatively large; and ectotympanic is annular, very small, and internally sided by ribbon-like entotympanic. Mandible is delicate, with long flat symphysis, thickening caudally; coronoid is relatively weak, long, and sloping, with square tip; condyle is slightly above lower alveolar line; and angle is gently rounded off. Upper incisors are moderately large, with subtle lingual basal shelf; I* is slightly larger; C! is strongly decurved; P' is tiny; posterior cheekteeth are relatively short and rectangular in occlusal outline, with posterobasal ledge and decreasing in height; molars are quadritubercular in appearance; and M* is small, with reduced cusps. Lower dentition has very large, triangular I, with basal lingual ledge, and very small, spatulated 1; C, is strongly decurved, slanted outward, and moderately small; P, is relatively very large, with tall labial cusp; posterior cheekteeth decrease in height and are rectangular in occlusal outline, with posterobasal ledge; and M, is peg-like.	Secondary lowland forests and disturbed agricultural areas from sea level up to elevations of ¢. 750 m .	The Mindoro Pallid Flying Fox is frugivorous. Seeds of Ficus spp. ( Moraceae ), Musa balbisiana ( Musaceae ), and one undetermined Melastomataceae were found in feces.	No information.	Mindoro Pallid Flying Foxes are nocturnal.	No information.	Not assessed on The IUCN Red List. The Mindoro Pallid Flying Fox is a narrow endemic of lowland Mindoro known from just two localities. It has been commonly found in secondary forests and highly disturbed agricultural areas with patchy forest remnants. This suggests adaptability to modified environments, which might be key for its survival in highly deforested Mindoro. It is likely best categorized as Data Deficient and in need of studies to determine actual status. It was captured in immediate vicinity of Mounts Iglit-Baco National Park, Mindoro.	Esselstyn et al. (2008) | Heaney et al. (2010)		117. Mindoro Pallid Flying Fox Desmalopex microleucopterus French: Roussette microptére / German: Mindoro-WeiRfllgelflughund / Spanish: Desmalopex de Mindoro Taxonomy. Desmalopex microleucopterus Esselstyn et al, 2008 , “Mount Siburan, Batong Buhay, Sablayan, Occidental Mindoro , Mindoro Island, Philippines , approximately 100 m above sea level ( 12.83488 N . 120.93028 E ).” This species is monotypic. Distribution. N Philippines (Mindoro). Descriptive notes. Head-body 133- 155 mm (tailless), hindfoot 29-34 mm , forearm 97-103 mm ; weight 129-156 g . Muzzle of the Mindoro Pallid Flying Fox is long, almost hairless, and whitish brown to pinkish; nostrils are large and divergent; and philtrum is undivided. Eyes are large, with orange irises. Ears are moderately long, pale brown, and narrowly rounded at tips. Body pelage is pale brown, soft, relatively long, and woollier in mantle. Uropatagium is narrow in center, tibia is sparsely furred, calcar is small, and index claw is present. Wing membranes are brown, with white blotches rather uniformly distributed on entire wing and slightly more intensely mottled in white at wingtips. Skull is typical pteropine, with pronounced basicranial deflection. Laterally, rostrum is long and tapering; nasal process of premaxilla is wide and procumbent; orbit is large, and rim is complete, with thin postorbital process joined to spine of thin arched zygoma; and zygomatic rootisjust above upper alveolar line. Dorsally, rostrum is long and rectangular, nasals are salient, paranasal recesses are moderately inflated reaching small postorbital foramina, postorbital constriction is moderately marked, sagittal crest is absent, braincase is oval, and nuchal crest is barely marked. Ventrally, palate is very long and flat; tooth rows are nearly parallel; post-dental palate is short, ending in deep concavity; ectopterygoids are comparatively large; and ectotympanic is annular, very small, and internally sided by ribbon-like entotympanic. Mandible is delicate, with long flat symphysis, thickening caudally; coronoid is relatively weak, long, and sloping, with square tip; condyle is slightly above lower alveolar line; and angle is gently rounded off. Upper incisors are moderately large, with subtle lingual basal shelf; I* is slightly larger; C! is strongly decurved; P' is tiny; posterior cheekteeth are relatively short and rectangular in occlusal outline, with posterobasal ledge and decreasing in height; molars are quadritubercular in appearance; and M* is small, with reduced cusps. Lower dentition has very large, triangular I, with basal lingual ledge, and very small, spatulated 1; C, is strongly decurved, slanted outward, and moderately small; P, is relatively very large, with tall labial cusp; posterior cheekteeth decrease in height and are rectangular in occlusal outline, with posterobasal ledge; and M, is peg-like. Habitat. Secondary lowland forests and disturbed agricultural areas from sea level up to elevations of ¢. 750 m . Food and Feeding. The Mindoro Pallid Flying Fox is frugivorous. Seeds of Ficus spp. ( Moraceae ), Musa balbisiana ( Musaceae ), and one undetermined Melastomataceae were found in feces. Breeding. No information. Activity patterns. Mindoro Pallid Flying Foxes are nocturnal. Movements, Home range and Social organization. No information. Status and Conservation. Not assessed on The IUCN Red List. The Mindoro Pallid Flying Fox is a narrow endemic of lowland Mindoro known from just two localities. It has been commonly found in secondary forests and highly disturbed agricultural areas with patchy forest remnants. This suggests adaptability to modified environments, which might be key for its survival in highly deforested Mindoro. It is likely best categorized as Data Deficient and in need of studies to determine actual status. It was captured in immediate vicinity of Mounts Iglit-Baco National Park, Mindoro. Bibliography. Esselstyn et al. (2008), Heaney et al. (2010).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Desmalopex microleucopterus	Desmalopex		microleucopterus	Esselstyn, Garcia, Saulog & Heaney	2008	0	J. Mammal.	89(4): 817	Mindoro Pallid Flying Fox	None.	Phillippines, Mindoro Isl., Occidental Mindoro, Sablayan, Batong Buhay, Mount Siburan	Mindoro Isl. Phillippines	Not listed.	Endangered	 Pteropus sp. A of Heaney et al. (1998)	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Desmalopex microleucopterus	23	Mindoro Pallid Flying Fox		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	PTEROPODINAE	PTERALOPINI	Desmalopex	NA	microleucopterus	Esselstyn, H. J. D. Garcia, Saulog, & Heaney	2008	0						"Mount Siburan, Batong Buhay, Sablayan, Occidental Mindoro, Mindoro Island, Philippines, approximately 100 m above sea level (12.83488 N, 120.93028 E)."	12.835	120.93	microleucopterus Esselstyn, H. J. D. Garcia, Saulog, & Heaney, 2008	recently described	Esselstyn, J. A., Garcia, H. J., Saulog, M. G., & Heaney, L. R. (2008). A new species of Desmalopex (Pteropodidae) from the Philippines, with a phylogenetic analysis of the Pteropodini. Journal of Mammalogy, 89(4), 815-825.	Philippines	Asia	Indomalaya	EN	0	0	0	Desmalopex_microleucopterus	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Desmalopex microleucopterus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Desmalopex	microleucopterus	Esselstyn, Garcia, Saulog &; Heaney, 2008		80000000	Desmalopex microleucopterus	Endangered	B1ab(iii,v)	2019	2018-08-31 00:00:00 UTC	3.1	English	This species is listed as Endangered because it is a Mindoro Island endemic with an extent of occurrence (EOO) of 1,589 kmÂ², it is only known from four localities, and the species is threatened from hunting by local communities, faces habitat loss and degradation, and is disturbed from recreational mountaineering.	There is limited information on the ecology and habitat requirements for D. microleucopterus . Since the species was described from Mt. Siburan, is has been captured on Mt. Halcon in secondary lowland and secondary lower montane forests. In Mts. Iglit-Baco National Park, Calintaan, Occidental Mindoro, individuals were captured in a disturbed lowland forest adjacent to grassland, agroforest and kaingin area. And in the Mt. Tallulah Peak area, Sablayan, Occidental Mindoro (140-180 m asl), individuals were captured in lowland forest with semi-evergreen structure. Additional research is required to expand and refine the understanding of the species ecology.	This species is only known from four areas on Mindoro Island and is threatened by local subsistence hunting for food, habitat loss and degradation, and disturbance brought about by tourism (mountaineering activity in Mt. Halcon). At this stage the precise threats to the species are not clear.	Very little information is known about the population status of the species as limited numbers of individuals only have been captured from four localities on Mindoro Island. It is inferred that the number of mature individuals is declining given the ongoing hunting of the species and decline in the extent and quality of its habitat.	Decreasing	Desmalopex microleucopterus is a Mindoro Island endemic (EOO is 1,589 kmÂ²) and has only been documented from four localities, including Mt. Halcon, Bayanan, Oriental Mindoro (450 and 1,480 m asl), Mts. Iglit-Baco National Park, Calintaan, Occidental Mindoro (120-250 m asl), Mt. Tallulah Peak, Sablayan, Occidental Mindoro (140-180 m asl), and Mt. Siburan (100 m asl, Type Locality). Additional information is needed to refine the understanding of the species distribution across Mindoro.	The species is hunted for local consumption.	Terrestrial	While most of the current known distribution for the species is outside of any known protected area, it has been found in the Mounts Iglit-Baco National Park which is also recognized as an ASEAN Heritage Park. It is unknown what percent of the species is found within the protected area. While there is no species-level conservation plan in place, a conservation management plan has been drafted for the Mt. Halcon area and is awaiting approval by local communities. There are no active monitoring activities in place. Further work is needed to determine the distribution, abundance, ecological requirements, threats and conservation needs of this species.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Desmalopex		microleucopterus	Esselstyn, Garcia, Saulog & Heaney	2008	0	J. Mammal.	89(4): 817	Mindoro Pallid Flying Fox	None.	Phillippines, Mindoro Isl., Occidental Mindoro, Sablayan, Batong Buhay, Mount Siburan	Mindoro Isl. Phillippines	Not listed.	Endangered	 Pteropus sp. A of Heaney et al. (1998)	Desmalopex microleucoptera	1004435	23	Mindoro Pallid Flying Fox		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	PTEROPODINAE	PTERALOPINI	Desmalopex	NA	microleucoptera	Esselstyn, H. J. D. Garcia, Saulog, & Heaney	2008	0						"Mount Siburan, Batong Buhay, Sablayan, Occidental Mindoro, Mindoro Island, Philippines, approximately 100 m above sea level (12.83488 N, 120.93028 E)."	12.83488	120.9303	microleucoptera Esselstyn, H. J. D. Garcia, Saulog, & Heaney, 2008	recently described; usually spelt 'microleucopterus', but the name is an adjective and Desmalopex is feminine, so the name should be spelt 'microleucoptera'	Esselstyn, J. A., Garcia, H. J., Saulog, M. G., & Heaney, L. R. (2008). A new species of Desmalopex (Pteropodidae) from the Philippines, with a phylogenetic analysis of the Pteropodini. Journal of Mammalogy, 89(4), 815-825.				Philippines	Asia	Indomalaya	EN	0	0	0	Desmalopex_microleucopterus	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Desmalopex_microleucoptera	1004435	23	Mindoro Pallid Flying Fox		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Pteropodinae	Pteralopini	Desmalopex	NA	microleucoptera	Esselstyn, Garcia, Saulog, & Heaney	0	Desmalopex microleucopterus	Esselstyn, J.A., Garcia, H.J.D., Saulog, M.G. and Heaney, L.R. 2008-08-15. A new species of _Desmalopex_ (Pteropodidae) from the Philippines, with a phylogenetic analysis of the Pteropodini. Journal of Mammalogy 89(4):815-825.	https://doi.org/10.1644/07-MAMM-A-285.1	NMPh 5202	holotype		"Mount Siburan, Batong Buhay, Sablayan, Occidental Mindoro, Mindoro Island, Philippines, approximately 100 m above sea level (12.83488 N, 120.93028 E)."	12.83488	120.9303	recently described; usually spelt 'microleucopterus', but the name is an adjective and Desmalopex is feminine, so the name should be spelt 'microleucoptera'	Esselstyn, J. A., Garcia, H. J., Saulog, M. G., & Heaney, L. R. (2008). A new species of Desmalopex (Pteropodidae) from the Philippines, with a phylogenetic analysis of the Pteropodini. Journal of Mammalogy, 89(4), 815-825.				Philippines	Asia	Indomalaya	EN (as Desmalopex microleucopterus)	0	0	0	Desmalopex_microleucopterus	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Desmalopex		microleucoptera	Esselstyn, Garcia, Saulog & Heaney	2008	0	J. Mammal.	89(4): 817	Mindoro Pallid Flying Fox	None.	Phillippines, Mindoro Isl., Occidental Mindoro, Sablayan, Batong Buhay, Mount Siburan	Mindoro Isl. Phillippines	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/84457227/84457393/' target='_blank'>Endangered as Desmalopex microleucopterus</a>	Pteropus sp. A of Heaney et al. (1998). We have changed the spelling of the species epithet, originally microleucopterus, to agree with the feminine genus name, Desmalopex; see Zijlstra (2023).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Desmalopex microleucopterus; Desmalopex microleucopterus; Desmalopex microleucopterus; Desmalopex microleucopterus; Desmalopex microleucopterus; microleucopterus; Roussette microptére; Mindoro-Wei Rfllgelflughund; Desmalopex de Mindoro; Mindoro Pallid Flying Fox; Mindoro Pallid Flying Fox; D. microleucoptera
