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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L224	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Pteropus leucopterus	Pteropus leucopterus	Pteropus leucopterus	Pteropus leucopterus	Pteropus leucopterus	Pteropus leucopterus	Desmalopex leucopterus	Desmalopex leucopterus	Desmalopex leucopterus	Desmalopex leucopterus	Desmalopex leucopterus	Desmalopex leucoptera	Desmalopex leucoptera	Desmalopex leucoptera	Desmalopex leucoptera		[MSW3] pselaphon species group.; [HMW] Pteropus leucopterus Temminck, 1853 , “les Philippines .” Desmalopex has been under Pteropus as a junior synonym since K. Andersen in 1912. Recent phylogenies demonstrated that it is a valid genus more closely related to monkeyfaced bats than typical pteropodines such as Pteropus . Desmalopex leucopterus is present in its typical form in the Luzon faunal region (Luzon and Catanduanes islands); other specimens referred might represent an undescribed species in the Mindanao faunal region (on Dinagat Island). Monotypic. On following pages: 117. Mindoro Pallid Flying Fox ( Desmalopex microleucopterus ); 118. Fijian Monkey-faced Fruit Bat ( Mirimiri acrodonta ); 119. Bougainville Monkey-faced Fruit Bat ( Pteralopex anceps ); 120. Guadalcanal Monkey-faced Fruit Bat ( Pteralopex atrata ); 121. Montane Monkey-faced Fruit Bat ( Pteralopex pulchra ); 122. New Georgia Monkey-faced Fruit Bat ( Pteralopex taki ); 123. Greater Monkey-faced Fruit Bat ( Pteralopex flanneryi ), 124. Black-bellied Blossom Bat ( Melonycteris melanops ); 125. Fardoulis's Blossom Bat ( Nesonycteris fardoulisi ); 126. Woodford's Blossom Bat ( Nesonycteris woodford)).; [batnames2022] Formerly included in Pteropus ; see Giannini et al. (2008).; [MDD2022] moved from Pteropus to Desmalopex; [IUCN] Desmalopex was originally described by Miller (1907) and included leucopterus based on distinctive cranial features. Leucopterus was subsequently placed in Pteropus . Recent DNA sequence data demonstrates that leucopterus is more closely related to Pteralopex and Acerodon than to Pteropus , supporting the recognition of Desmalopex as a valid genus (Giannini et al. ;2008, Almeida et al. ;2014).; [batnames2023] Formerly included in Pteropus ; see Giannini et al. (2008).; [MDD2023] usually spelt 'leucopterus', but the name is an adjective and Desmalopex is feminine, so the name should be spelt 'leucoptera'; moved from Pteropus to Desmalopex; [MDD2025_2.0] usually spelt 'leucopterus', but the name is an adjective and Desmalopex is feminine, so the name should be spelt 'leucoptera'; moved from Pteropus to Desmalopex; [batnames2025_1.7] Formerly included in Pteropus; see Giannini et al. (2008). Although the species epithet has been spelled leucopterus since Miller (1907) transferred the species from Pteropus to Desmalopex, Desmalopex is feminine, and we have changed the epithet spelling to leucoptera accordingly; see Zijlstra (2023).; [MDD2025_2.2] usually spelt 'leucopterus', but the name is an adjective and Desmalopex is feminine, so the name should be spelt 'leucoptera'; moved from Pteropus to Desmalopex						chinensis.			chinensis					leucopterus, chinensis	Desmalopex was originally described by Miller (1907) and included leucopterus based on distinctive cranial features. Leucopterus was subsequently placed in Pteropus . Recent DNA sequence data demonstrates that leucopterus is more closely related to Pteralopex and Acerodon than to Pteropus , supporting the recognition of Desmalopex as a valid genus (Giannini et al. ;2008, Almeida et al. ;2014).			leucoptera, chinensis	leucoptera, chinensis			leucoptera (Temminck, 1853)|chinensis (J. E. Gray, 1871)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Luzon, Philippines	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Pteropus leucopterus	Philippines.	Temminck	1853	Esquisses Zool. sur la Cote de Guine, p. 60.	Distribution: Confined to the Philippines.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Philippines	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Temminck	1853	Esquisses Zool. sur la Cote de Guine, p. 60.		Philippines (known only from Luzon and Dinagat).	Philippines.		TEMMINCK	1853	Tibia hairy dorsally. Upper molariform teeth shortened and subquadrate. Postorbital processes forming a complete postorbital bar. Size large for group (forearm length, 136-143 mm).	Distribution: Confined to the Philippines.	No subspecies.		25	species	P. leucopterus	TEMMINCK	1853	Pteropus	genus	Pteropus leucopterus				Tibia hairy dorsally. Upper molariform teeth shortened and subquadrate. Postorbital processes forming a complete postorbital bar. Size large for group (forearm length, 136-143 mm).	No subspecies.		41. P. leucopterus TEMMINCK 1853 [pselaphon group].	41	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Pteropus leucopterus	Pteropus		leucopterus	Temminck		1853		Esquisses Zool. sur la Côte de Guine			60		White-winged Flying Fox	Philippines.	Luzon, Catanduanes, and Dinagat Isls (Philippines).	CITES – Appendix II. IUCN/SSC Action Plan (1992) – Vulnerable. IUCN 2003 – Endangered.	chinensis Gray, 1871.	pselaphon species group.	03AD87FAFF87F66E8C673F9BFE54F614	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	125	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFF87F66E8C673F9BFE54F614.xml	Desmalopex leucopterus	Pteropodidae	Desmalopex	leucopterus		1853	Roussette leucoptere @fr | \Weilkfligelflughund @de | Desmalopex de alas blancas @es | Mottle-winged Flying Fox @en	Pteropus leucopterus Temminck, 1853 , “les Philippines .” Desmalopex has been under Pteropus as a junior synonym since K. Andersen in 1912. Recent phylogenies demonstrated that it is a valid genus more closely related to monkeyfaced bats than typical pteropodines such as Pteropus . Desmalopex leucopterus is present in its typical form in the Luzon faunal region (Luzon and Catanduanes islands); other specimens referred might represent an undescribed species in the Mindanao faunal region (on Dinagat Island). Monotypic. On following pages: 117. Mindoro Pallid Flying Fox ( Desmalopex microleucopterus ); 118. Fijian Monkey-faced Fruit Bat ( Mirimiri acrodonta ); 119. Bougainville Monkey-faced Fruit Bat ( Pteralopex anceps ); 120. Guadalcanal Monkey-faced Fruit Bat ( Pteralopex atrata ); 121. Montane Monkey-faced Fruit Bat ( Pteralopex pulchra ); 122. New Georgia Monkey-faced Fruit Bat ( Pteralopex taki ); 123. Greater Monkey-faced Fruit Bat ( Pteralopex flanneryi ), 124. Black-bellied Blossom Bat ( Melonycteris melanops ); 125. Fardoulis's Blossom Bat ( Nesonycteris fardoulisi ); 126. Woodford's Blossom Bat ( Nesonycteris woodford)).	N Philippines (Luzon and Catanduanes ).	Head-body 185- 240 mm (tailless), ear 26-28 mm , hindfoot 44-46 mm , forearm 135-145 mm ; weight 250-375 g . Head of the Whitewinged Flying Fox is rounded, with long fox-like muzzle; tip is almost hairless and whitish brown to pinkish; nostrils are large and divergent; and philtrum is undivided. Eyes are moderately large, with warm brown to orange-brown irises. Ears are short, pale brown, and attenuated at tips. Body pelage is generally pale brown; soft; of medium length; longer in rather indistinct mantle; and paler turning yellowish in crown, nape, and dorsum in some specimens; forearm is densely furred on dorsal proximal one-half. Uropatagium is narrow in center,tibia is thickly furred to ankle, and calcar is small. Index claw is present. Wing membranes are brown, with white blotches, more intensely mottled in white on long thumb, wing leading edge, and wingtip. Skull is typical pteropine, with noticeable basicranial deflection; laterally rostrum is long and tapering; nasal process of premaxilla is wide; orbit is large, slightly tilted upward; zygomatic root is just above upper alveolar line; and zygomais strong and arched. Dorsally, rostrum is long; nasals are salient; paranasal recesses are inflated, reaching small postorbital foramina; postorbital processes are long, arched, and annectant to zygoma in most adult specimens; postorbital constriction is very obvious; temporal lines joined in low sagittal crest; braincase is oval; and nuchal crest is very obvious. Ventrally, palate is very long and flat; tooth rows are nearly parallel; post-dental palate is short, ending in deep concavity; and ectotympanic is annular, very small, and internally sided by incomplete entotympanic. Mandible has long, flat symphysis, thickening caudally; coronoid is long and sloping, with square tip; condyle is slightly above lower alveolar line; and angle is gently rounded off. Dental formula for all species of Desmalopexis12/2,C 1/1, P 3/3, M 2/3 (x2) = 34. Dentition is generally heavy. Upper incisors are large, with noticeable lingual basal shelf; I* is slightly larger; C' is strong and decurved; P' is small, with round and flat crown; posterior cheekteeth are relatively short and squarish in occlusal outline, with posterobasal ledge, decreasing in height; molars are quadritubercular in appearance; and M* is small, with reduced cusps. Lower dentition has very large, tricuspidate I, with wide basal lingual ledge, and much smaller, spatulate I; C,is strongly decurved and moderately small; P, is relatively very large, with marked labial cusp; posterior cheekteeth decrease in height and are rectangular in occlusal outline, with posterobasal ledge; and M,is peg-like. Diploid number is 2n = 38, with one pair submetacentric and 18 pairs acrocentric or subtelocentric autosomes.	Primary lowland and montane forests from sea level up to elevations of ¢. 1200 m .	The White-winged Flying Fox is frugivorous.	In captivity, White-winged Flying Foxes have one young/year.	White-winged Flying Foxes are nocturnal.	No information.	CITES Appendix II (as Pteropus leucopterus ) since 1990. Classified as Least Concern on The IUCN Red Lust (as Pteropus leucopterus ). The White-winged Flying Fox is tolerant to some habitat modification and relatively widely distributed. A captive breeding program exists at Silliman University, Philippines .	Andersen (1912b) | Esselstyn et al. (2008) | Giannini et al. (2008) | Heaney & Rabor (1982) | Heaney, Balete et al. (1998) | Heaney, Dolar et al. (2010) | Miller (1907) | Ong, Rosell-Ambal, Tabaranza, Balete et al. (2008) | Rickart et al. (1999)		116. White-winged Flying Fox Desmalopex leucopterus French: Roussette leucoptere / German: \Weilkfligelflughund / Spanish: Desmalopex de alas blancas Other common names: Mottle-winged Flying Fox Taxonomy. Pteropus leucopterus Temminck, 1853 , “les Philippines .” Desmalopex has been under Pteropus as a junior synonym since K. Andersen in 1912. Recent phylogenies demonstrated that it is a valid genus more closely related to monkeyfaced bats than typical pteropodines such as Pteropus . Desmalopex leucopterus is present in its typical form in the Luzon faunal region (Luzon and Catanduanes islands); other specimens referred might represent an undescribed species in the Mindanao faunal region (on Dinagat Island). Monotypic. On following pages: 117. Mindoro Pallid Flying Fox ( Desmalopex microleucopterus ); 118. Fijian Monkey-faced Fruit Bat ( Mirimiri acrodonta ); 119. Bougainville Monkey-faced Fruit Bat ( Pteralopex anceps ); 120. Guadalcanal Monkey-faced Fruit Bat ( Pteralopex atrata ); 121. Montane Monkey-faced Fruit Bat ( Pteralopex pulchra ); 122. New Georgia Monkey-faced Fruit Bat ( Pteralopex taki ); 123. Greater Monkey-faced Fruit Bat ( Pteralopex flanneryi ), 124. Black-bellied Blossom Bat ( Melonycteris melanops ); 125. Fardoulis's Blossom Bat ( Nesonycteris fardoulisi ); 126. Woodford's Blossom Bat ( Nesonycteris woodford)). Distribution. N Philippines (Luzon and Catanduanes ). Descriptive notes. Head-body 185- 240 mm (tailless), ear 26-28 mm , hindfoot 44-46 mm , forearm 135-145 mm ; weight 250-375 g . Head of the Whitewinged Flying Fox is rounded, with long fox-like muzzle; tip is almost hairless and whitish brown to pinkish; nostrils are large and divergent; and philtrum is undivided. Eyes are moderately large, with warm brown to orange-brown irises. Ears are short, pale brown, and attenuated at tips. Body pelage is generally pale brown; soft; of medium length; longer in rather indistinct mantle; and paler turning yellowish in crown, nape, and dorsum in some specimens; forearm is densely furred on dorsal proximal one-half. Uropatagium is narrow in center,tibia is thickly furred to ankle, and calcar is small. Index claw is present. Wing membranes are brown, with white blotches, more intensely mottled in white on long thumb, wing leading edge, and wingtip. Skull is typical pteropine, with noticeable basicranial deflection; laterally rostrum is long and tapering; nasal process of premaxilla is wide; orbit is large, slightly tilted upward; zygomatic root is just above upper alveolar line; and zygomais strong and arched. Dorsally, rostrum is long; nasals are salient; paranasal recesses are inflated, reaching small postorbital foramina; postorbital processes are long, arched, and annectant to zygoma in most adult specimens; postorbital constriction is very obvious; temporal lines joined in low sagittal crest; braincase is oval; and nuchal crest is very obvious. Ventrally, palate is very long and flat; tooth rows are nearly parallel; post-dental palate is short, ending in deep concavity; and ectotympanic is annular, very small, and internally sided by incomplete entotympanic. Mandible has long, flat symphysis, thickening caudally; coronoid is long and sloping, with square tip; condyle is slightly above lower alveolar line; and angle is gently rounded off. Dental formula for all species of Desmalopexis12/2,C 1/1, P 3/3, M 2/3 (x2) = 34. Dentition is generally heavy. Upper incisors are large, with noticeable lingual basal shelf; I* is slightly larger; C' is strong and decurved; P' is small, with round and flat crown; posterior cheekteeth are relatively short and squarish in occlusal outline, with posterobasal ledge, decreasing in height; molars are quadritubercular in appearance; and M* is small, with reduced cusps. Lower dentition has very large, tricuspidate I, with wide basal lingual ledge, and much smaller, spatulate I; C,is strongly decurved and moderately small; P, is relatively very large, with marked labial cusp; posterior cheekteeth decrease in height and are rectangular in occlusal outline, with posterobasal ledge; and M,is peg-like. Diploid number is 2n = 38, with one pair submetacentric and 18 pairs acrocentric or subtelocentric autosomes. Habitat. Primary lowland and montane forests from sea level up to elevations of ¢. 1200 m . Food and Feeding. The White-winged Flying Fox is frugivorous. Breeding. In captivity, White-winged Flying Foxes have one young/year. Activity patterns. White-winged Flying Foxes are nocturnal. Movements, Home range and Social organization. No information. Status and Conservation. CITES Appendix II (as Pteropus leucopterus ) since 1990. Classified as Least Concern on The IUCN Red Lust (as Pteropus leucopterus ). The White-winged Flying Fox is tolerant to some habitat modification and relatively widely distributed. A captive breeding program exists at Silliman University, Philippines . Bibliography. Andersen (1912b), Esselstyn et al. (2008), Giannini et al. (2008), Heaney & Rabor (1982), Heaney, Balete et al. (1998), Heaney, Dolar et al. (2010), Miller (1907), Ong, Rosell-Ambal, Tabaranza, Balete et al. (2008), Rickart et al. (1999).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Desmalopex leucopterus	Desmalopex		leucopterus	Temminck	1853	1	Esquisses Zool. sur la C&ocirc;te de Guine	p. 60	White-winged Flying Fox	None.	Phillipines	Luzon, Catanduanes, and Dinagat Isls (Philippines).	Not listed.	Vulnerable	Formerly included in Pteropus ; see Giannini et al. (2008).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Desmalopex leucopterus	23	White-winged Flying Fox	Mottle-winged Flying Fox	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	PTEROPODINAE	PTERALOPINI	Desmalopex	NA	leucopterus	Temminck	1853	1						"îles Philippines."			leucopterus (Temminck, 1853)|chinensis (J. E. Gray, 1871)	moved from Pteropus to Desmalopex	Giannini, N. P., Almeida, F. C., Simmons, N. B., & Helgen, K. M. (2008). The systematic position of Pteropus leucopterus and its bearing on the monophyly and relationships of Pteropus (Chiroptera: Pteropodidae). Acta Chiropterologica, 10(1), 11-20.	Philippines	Asia	Indomalaya	VU	0	0	0	Desmalopex_leucopterus	0	oldname match	Pteropus_leucopterus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	18731	Desmalopex leucopterus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Desmalopex	leucopterus	Temminck, 1853	Desmalopex was originally described by Miller (1907) and included leucopterus based on distinctive cranial features. Leucopterus was subsequently placed in Pteropus . Recent DNA sequence data demonstrates that leucopterus is more closely related to Pteralopex and Acerodon than to Pteropus , supporting the recognition of Desmalopex as a valid genus (Giannini et al. ;2008, Almeida et al. ;2014).	20000000	Desmalopex leucopterus	Vulnerable	A2cd	2020	2020-07-19 00:00:00 UTC	3.1	English	Desmalopex leucopterus is assessed as Vulnerable under criterion A2cd as the species is suspected to have had a significant 30-35% decline over the last 24.3 years (three generations, GL = 8.1 years, Pacifici et al. 2013) due to extensive hunting (Scheffers et al. ; 2012), roost disturbance, and widespread loss and degradation of lowland forest habitats (Gonzalez et al. 2018) resulting in a decline in the species EOO and AOO. The species is also assessed as Vulnerable under the Philippines Red List (Gonzalez et al. 2018) as critical lowland forest have been reduced to less than 8% of its original extent (ca 2% old growth, 6% secondary) (Walpole 2010).	Although Desmalopex leucopterus has been reported from the limited primary montane forest on Catanduanes island, it is predominantly known from lowland primary dipterocarp forests (where they remain), secondary forests, forest fragments, plantation forests (Heaney et al. 1991, Mickleburgh et al. 1992, Utzurrum 1992, van Weerd et al. , 2003). It has also been documented in grasslands in Benguet and San Mariano (L. Heaney pers. comm. 2006) and in limestone forest in Bulacan (Duya pers. comm. 2020). The species was also reported to be frequently captured in ultramafic forest (Duya 2019). Lowland forest have been reduced to less than 8% of its original extent (ca. 2% old growth, 6% secondary) (Walpole 2010) and it is not known if the species is able to persist in heavily degraded habitats (L. Heaney pers. comm. 2007). The species has been documented from sea level to 1,200 m. This species is not known to form large conspicuous roosts in trees or to co-roost with other species of flying foxes, but probably roosts singly or in small groups (van Weerd et al. ;2003). It appears to have a seasonal presence in some areas (Balbalasan) which suggests that it might be migratory (Heaney and Utzurrum 1991, L. Heaney pers. comm. 2006). The species breeds once per year in captivity (A. Cariño pers. comm. 2006) and generally gives birth to one pup and occasionally two pups (E. Alcala pers. comm. 2006). An individual carrying a pup was also observed in June in a limestone forest fragment in Bulacan (Duya pers comm. 2020).<p></p>	The primary threats to the species include hunting as bushmeat (Scheffers et al. ;2012), roost disturbance, and loss and degradation of lowland forests (Utzurrum 1992, Heaney et al. 1998, Gonzalez et al. ;2018) due to agriculture, timber production, road construction, and expanding human settlements. Lowland forests have been reduced to less than 8% of its original extent (ca 2% old growth, 6% secondary) (Walpole 2010). The species is hunted, including in forest reserves, where the demand from local communities provide an important market to hunters (Scheffers et al. ;2012), flying foxes are the most sought after bushmeat and sell for ca. 150 PhP per flying fox - about $3 USD (Scheffers et al. ;2012).	The population trend of Desmalopex leucopterus is suspected to have a significant 30-35% decline over the last 24.3 years (three generations, GL = 8.1 years, Pacifici et al. 2013) due to extensive hunting (Scheffers et al. , 2012), roost disturbance, and widespread loss and degradation of lowland forest habitats (Utzurrum 1992, Heaney et al. 1998, Gonzalez et al. 2018) resulting in a decline in its extent of occurrence (EOO) and area of occupancy (AOO). It is a high-flying species that is difficult to capture in surveys (L. Heaney pers. comm. 2007). There are seasonal fluctuations in relative number of captures which might indicate that it is migratory (Heaney and Utzurrum 1991).	Decreasing	Desmalopex leucopterus , the White-winged Flying Fox, is endemic to the Philippines, where it is found in the Luzon faunal region and Dinagat. It has been recorded on Luzon (provinces including: Abra, Aurora, Benguet, Bicol, Bulacan, Cagayan, Isabela, Laguna, Nueva Ecija, Nueva Viscaya, Quezon, Sorsogon, Tarlac, and Zambales (D. Balete pers. comm. 2006, L. Heaney pers. comm. 2006, M.R. Duya pers. com. 2020)), Catanduanes and Dinagat Islands (Heaney et al. 1998, Heaney et al. ;2010), occurring at an elevation range from 0–1,200 m asl (Heaney et al. 2010, M.R. Duya pers. com. 2020).	The species is thought to be hunted extensively for local markets and consumption (Scheffers et al. ;2012).	Terrestrial	Desmalopex leucopterus is known to occur in some protected areas (e.g., Pantabangan-Carranglan Protected Landscape and Seascape, Northern Sierra Madre Natural Park) and has been included in CITES Appendix II since 1990. Given hunting of the species is predominantly for local consumption, local education and awareness programs are warranted to reduce hunting pressure (Scheffers et al. ;2012). Enforcement of local ordinances and fines may provide effective deterrents to hunters (Scheffers et al. ;2012) and alternative livelihoods may prove effective to reduce hunting pressures. Forest protection and restoration are needed to expand roosting and foraging areas. Additional research is needed on the species population status and trends, ecology, distribution, habitat preferences, and threats.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Desmalopex		leucopterus	Temminck	1853	1	Esquisses Zool. sur la C&ocirc;te de Guine	p. 60	White-winged Flying Fox	None.	Phillipines	Luzon, Catanduanes, and Dinagat Isls (Philippines).	Not listed.	Vulnerable	Formerly included in Pteropus ; see Giannini et al. (2008).	Desmalopex leucoptera	1004434	23	White-winged Flying Fox	Mottle-winged Flying Fox	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	PTEROPODINAE	PTERALOPINI	Desmalopex	NA	leucoptera	Temminck	1853	1						"îles Philippines."			leucoptera (Temminck, 1853)|chinensis (J. E. Gray, 1871)	usually spelt 'leucopterus', but the name is an adjective and Desmalopex is feminine, so the name should be spelt 'leucoptera'; moved from Pteropus to Desmalopex	Giannini, N. P., Almeida, F. C., Simmons, N. B., & Helgen, K. M. (2008). The systematic position of Pteropus leucopterus and its bearing on the monophyly and relationships of Pteropus (Chiroptera: Pteropodidae). Acta Chiropterologica, 10(1), 11-20.				Philippines	Asia	Indomalaya	VU	0	0	0	Desmalopex_leucopterus	0	oldname match	Pteropus_leucopterus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Desmalopex_leucoptera	1004434	23	White-winged Flying Fox	Mottle-winged Flying Fox	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Pteropodinae	Pteralopini	Desmalopex	NA	leucoptera	Temminck	1	Pteropus leucopterus	Temminck, C.J. 1853. Esquisses zoologiques sur la côte de Guiné. E. J. Brill, Leiden, 256 pp.	https://www.biodiversitylibrary.org/page/14835807	RMNH.MAM.37855	holotype	https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.37855.a | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.37855.b	"îles Philippines."			usually spelt 'leucopterus', but the name is an adjective and Desmalopex is feminine, so the name should be spelt 'leucoptera'; moved from Pteropus to Desmalopex	Giannini, N. P., Almeida, F. C., Simmons, N. B., & Helgen, K. M. (2008). The systematic position of Pteropus leucopterus and its bearing on the monophyly and relationships of Pteropus (Chiroptera: Pteropodidae). Acta Chiropterologica, 10(1), 11-20.				Philippines	Asia	Indomalaya	VU (as Desmalopex leucopterus)	0	0	0	Desmalopex_leucopterus	0	oldname match	Pteropus_leucopterus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Desmalopex		leucoptera	Temminck	1853	1	Esquisses Zool. sur la C&ocirc;te de Guine	p. 60	White-winged Flying Fox	None.	Phillipines	Luzon, Catanduanes, and Dinagat Isls (Philippines).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/18731/22081331/' target='_blank'>Vulnerable as Desmalopex leucopterus</a>	Formerly included in Pteropus; see Giannini et al. (2008). Although the species epithet has been spelled leucopterus since Miller (1907) transferred the species from Pteropus to Desmalopex, Desmalopex is feminine, and we have changed the epithet spelling to leucoptera accordingly; see Zijlstra (2023).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Pteropus leucopterus; Desmalopex leucopterus; Desmalopex leucopterus; Desmalopex leucopterus; Desmalopex leucopterus; Desmalopex leucopterus; chinensis; leucopterus; chinensis; Roussette leucoptere; \Weilkfligelflughund; Desmalopex de alas blancas; Mottle-winged Flying Fox; White-winged Flying Fox; Mottle-winged Flying Fox; White-winged Flying Fox; White-winged Flying Fox; D. leucoptera