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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L219	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	Artibeus phaeotis [synonym of]	N/A	Artibeus phaeotis [synonym of]	Artibeus phaeotis ravus	Artibeus phaeotis ravus	Artibeus ravus	Dermanura rava	Dermanura rava	Dermanura rava	Dermanura rava	Dermanura rava	Dermanura rava	Dermanura rava	Dermanura rava		[HMW] Dermanura rava G. S. Miller, 1902 , “San Javier [ Esmeraldas ], northern Ecuador .” Artibeus ravus , along with smaller species of Artibeus , 1s placed in Dermanura ; however, after reinterpretation of molecular analyses, this clade was reassigned to Artibeus (subgenus Dermanura ). Several authors listed A. ravus as a synonym of A. phaeotis , but according to molecular and morphological analyses,it is a distinct species. Monotypic.; [batnames2022] Subgenus Dermanura . Previously considered a subspecies of phaeotis , but see Hoofer et al. (2008) and Solari et al. (2009).; [MDD2022] split from A. phaeotis; moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus; [IUCN] <span>Previously this was assessed as the subspecies D. phaeotis rava . It has now been raised to species level (Solari et al. 2009).; [batnames2023] Subgenus Dermanura . Previously considered a subspecies of phaeotis , but see Hoofer et al. (2008) and Solari et al. (2009).; [MDD2023] split from A. phaeotis; moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus; [MDD2025_2.0] split from A. phaeotis; moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus; [batnames2025_1.7] Subgenus Dermanura. Previously considered a subspecies of phaeotis, but see Hoofer et al. (2008) and Solari et al. (2009).; [MDD2025_2.2] split from A. phaeotis; moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus														rava	<span>Previously this was assessed as the subspecies D. phaeotis rava . It has now been raised to species level (Solari et al. 2009).			rava	rava			rava G. S. Miller, 1902						N/A																																								NA																											03A687BCFFEFFFEE1696F82BFB84F056	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	578	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFFEFFFEE1696F82BFB84F056.xml	Artibeus ravus	Phyllostomidae	Artibeus	ravus		1902	Dermanure jaunatre @fr | Kleiner Fruchtvampir @de | Artibeoamarillento @es | Amber Fruit-eating Bat @en | Yellowish Little Fruit-eating Bat @en	Dermanura rava G. S. Miller, 1902 , “San Javier [ Esmeraldas ], northern Ecuador .” Artibeus ravus , along with smaller species of Artibeus , 1s placed in Dermanura ; however, after reinterpretation of molecular analyses, this clade was reassigned to Artibeus (subgenus Dermanura ). Several authors listed A. ravus as a synonym of A. phaeotis , but according to molecular and morphological analyses,it is a distinct species. Monotypic.	W Colombia ( Choco Department and middle and lower Cauca and Magdalena valleys) S through W slope of Andes to SW Ecuador and extreme NW Peru ( Tumbes Department).	Head-body 44-62 mm (tailless), ear 11-17 mm, hindfoot 9-14 mm, forearm 37-42 mm; weight 10-17 g. The Little Fruit-eating Bat is small, with facial stripes and short and broad rostrum. Dorsal fur is light yellowish brown, more pale than dark; fur is short and faintly tricolored (paler bases). Whitish facial stripes are almost well defined but never too contrasting. Ventral fur is slightly paler than dorsum. Ears and noseleaf are pale brown, and bases and margins of ears are usually narrowly rimmed in white or cream. Fur covers more than one-half of each forearm. Wing membranes are brownish and attach to metatarsus. Tail membrane is pale brown, moderated in size, U-shaped, and sparsely haired. Dental formulais12/2,C1/1,P 2/2, M 2/2 ( x2 ) = 28. Skull is elongated, without inflated frontal and supraorbital areas. M' has long hypocone (talon), and M,is always absent.	Evergreen lowlands and pre-montane forests, second growth, tropical dry forests, fruit groves, and inter-Andean dry valleys from lowlands up to elevations of¢. 2000 m (usually below 600 m ). The Little Fruit-eating Bat has broad variation in habitat preferences through its distribution.	Based on resemblance to congeners and its habitats in western Ecuador , the Little Fruit-eating Bat is assumed to be mainly frugivorous.	Pregnant and lactating Little Fruit-eating Bats have been found in June-July in Colombia .	The Little Fruit-eating Bat is nocturnal. It is usually more active 2-3 hours after sunset.	No information.	Not assessed on The IUCN Red List. Additional information on population status and ecology of the Little Fruit-eating Bat is needed for an accurate conservation assessment.	Andersen (1908c) | Carrera et al. (2010) | Davis (1969) | Handley (1987) | Marques-Aguiar (2008a) | Miller (1902a) | Solari, Hoofer et al. (2009) | Solari, Munoz-Saba et al. (2013) | Timm (1985)	https://zenodo.org/record/6459005/files/figure.png	205. Little Fruit-eating Bat Artibeus ravus French: Dermanure jaunatre / German: Kleiner Fruchtvampir / Spanish: Artibeo amarillento Other common names: Amber Fruit-eating Bat , Yellowish Little Fruit-eating Bat Taxonomy. Dermanura rava G. S. Miller, 1902 , “San Javier [ Esmeraldas ], northern Ecuador .” Artibeus ravus , along with smaller species of Artibeus , 1s placed in Dermanura ; however, after reinterpretation of molecular analyses, this clade was reassigned to Artibeus (subgenus Dermanura ). Several authors listed A. ravus as a synonym of A. phaeotis , but according to molecular and morphological analyses,it is a distinct species. Monotypic. Distribution. W Colombia ( Choco Department and middle and lower Cauca and Magdalena valleys) S through W slope of Andes to SW Ecuador and extreme NW Peru ( Tumbes Department). Descriptive notes. Head-body 44-62 mm (tailless), ear 11-17 mm, hindfoot 9-14 mm, forearm 37-42 mm; weight 10-17 g. The Little Fruit-eating Bat is small, with facial stripes and short and broad rostrum. Dorsal fur is light yellowish brown, more pale than dark; fur is short and faintly tricolored (paler bases). Whitish facial stripes are almost well defined but never too contrasting. Ventral fur is slightly paler than dorsum. Ears and noseleaf are pale brown, and bases and margins of ears are usually narrowly rimmed in white or cream. Fur covers more than one-half of each forearm. Wing membranes are brownish and attach to metatarsus. Tail membrane is pale brown, moderated in size, U-shaped, and sparsely haired. Dental formulais12/2,C1/1,P 2/2, M 2/2 ( x2 ) = 28. Skull is elongated, without inflated frontal and supraorbital areas. M' has long hypocone (talon), and M,is always absent. Habitat. Evergreen lowlands and pre-montane forests, second growth, tropical dry forests, fruit groves, and inter-Andean dry valleys from lowlands up to elevations of¢. 2000 m (usually below 600 m ). The Little Fruit-eating Bat has broad variation in habitat preferences through its distribution. Food and Feeding. Based on resemblance to congeners and its habitats in western Ecuador , the Little Fruit-eating Bat is assumed to be mainly frugivorous. Breeding. Pregnant and lactating Little Fruit-eating Bats have been found in June-July in Colombia . Activity patterns. The Little Fruit-eating Bat is nocturnal. It is usually more active 2-3 hours after sunset. Movements, Home range and Social organization. No information. Status and Conservation. Not assessed on The IUCN Red List. Additional information on population status and ecology of the Little Fruit-eating Bat is needed for an accurate conservation assessment. Bibliography. Andersen (1908c), Carrera et al. (2010), Davis (1969), Handley (1987), Marques-Aguiar (2008a), Miller (1902a), Solari, Hoofer et al. (2009), Solari, Munoz-Saba et al. (2013), Timm (1985).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Dermanura rava	Dermanura		rava	Miller	1902	0	Proc. Acad. Nat. Sci. Phil.	60:44:00	Amber Fruit-eating Bat	None.	Ecuador, Esmeraldas, San Javier.	W Colombia	Not listed.	Least Concern	Subgenus Dermanura . Previously considered a subspecies of phaeotis , but see Hoofer et al. (2008) and Solari et al. (2009).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Dermanura rava	23	Little Fruit-eating Bat	Amber Fruit-eating Bat|Yellowish Little Fruit-eating Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	STENODERMATINAE	STENODERMATINI	Dermanura	NA	rava	G. S. Miller	1902	0	Dermanura_rava	Miller, G. S., Jr. (1902). Twenty New American Bats. Proceedings of the Academy of Natural Sciences of Philadelphia, 54, 404.	https://www.biodiversitylibrary.org/page/10309520#page/438/mode/1up	USNM 113338		"San Javier [Esmeraldas], northern Ecuador."			rava G. S. Miller, 1902	split from A. phaeotis; moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus	Solari, S., Hoofer, S. R., Larsen, P. A., Brown, A. D., Bull, R. J., Guerrero, J. A., ... & Baker, R. J. (2009). Operational criteria for genetically defined species: analysis of the diversification of the small fruit-eating bats, Dermanura (Phyllostomidae: Stenodermatinae). Acta Chiropterologica, 11(2), 279-288.|Hoofer, S. R., Solari, S., Larsen, P. A., Bradley, R. D., & Baker, R. J. (2008). Phylogenetics of the fruit-eating bats (Phyllostomidae: Artibeina) inferred from mitochondrial DNA sequences. Occasional Papers of the Museum of Texas Tech University, 277, 1-15.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.|York, H. A., Rodríguez-Herrera, B., Laval, R. K., Timm, R. M., & Lindsay, K. E. (2019). Field key to the bats of Costa Rica and Nicaragua. Journal of Mammalogy, 100(6), 1726-1749.|Turcios-Casco, M. A., Ávila-Palma, H. D., LaVal, R. К., Stevens, R. D., Ordoñez-Trejo, E. J., Soler-Orellana, J. A., & Ordoñez-Mazier, D. I. (2020). A systematic revision of the bats (Chiroptera) of Honduras: an updated checklist with corroboration of historical specimens and new records. Zoosystematics and Evolution, 96, 411.|	Colombia|Ecuador|Peru	South America	Neotropic	LC	0	0	0	Artibeus_ravus	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Dermanura rava	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Dermanura	rava	Miller, 1902	<span>Previously this was assessed as the subspecies D. phaeotis rava . It has now been raised to species level (Solari et al. 2009).	80000000	Dermanura rava	Least Concern		2019	2016-07-12 00:00:00 UTC	3.1	English	Listed as Least Concern because of ;its wide geographic distribution, including large sections of continuous native forests, and ;natural abundance at local scale. There is no major threats to these habitats or the species itself.	Dermanura rava inhabits dry and wet forests of western Ecuador and northwestern Peru, although it seems to prefer dry forests (Pacheco et al. 2009, Carrera et al. 2010). In Colombia, it is present from the Pacific lowland forests of the Chocó to the dry valleys of the Cauca and Magdalena rivers (Solari et al. 2013). This small frugivore may be found in open areas too, being able to move long distances every night to get to its key resources, typically fruits (such as Ficus sp.).	The threats to this species are unknown.	In western Ecuador, this species inhabits dry and wet forests, being more commonly collected in dry forests (Carrera et al. 2010). This is a common species along the interandean valleys of western Colombia (Solari, pers. obs.).	Stable	This species is found from southeastern Panama through the Pacific versant of Colombia and Ecuador, western and central Colombia, and northwestern Peru (Pacheco et al. 2009, Solari et al. 2009, Carrera et al. 2010, Solari et al. 2013).		Terrestrial	A couple of national reserves exist in northwestern Peru, where the southern limit of the species is reached. Also, small natural reserves protect the Tropical dry forests of western Ecuador (Carrera et al. 2010). A few protected areas occur in western Colombia, including ;natural reserves and ;national parks.	Nearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Dermanura		rava	Miller	1902	0	Proc. Acad. Nat. Sci. Phil.	60:44:00	Amber Fruit-eating Bat	None.	Ecuador, Esmeraldas, San Javier.	W Colombia	Not listed.	Least Concern	Subgenus Dermanura . Previously considered a subspecies of phaeotis , but see Hoofer et al. (2008) and Solari et al. (2009).	Dermanura rava	1005016	23	Little Fruit-eating Bat	Amber Fruit-eating Bat|Yellowish Little Fruit-eating Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	STENODERMATINAE	STENODERMATINI	Dermanura	NA	rava	G. S. Miller	1902	0	Dermanura_rava	Miller, G. S., Jr. (1902). Twenty New American Bats. Proceedings of the Academy of Natural Sciences of Philadelphia, 54, 404.	https://www.biodiversitylibrary.org/page/10309520#page/438/mode/1up	USNM 113338		"San Javier [Esmeraldas], northern Ecuador."			rava G. S. Miller, 1902	split from A. phaeotis; moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus	Solari, S., Hoofer, S. R., Larsen, P. A., Brown, A. D., Bull, R. J., Guerrero, J. A., ... & Baker, R. J. (2009). Operational criteria for genetically defined species: analysis of the diversification of the small fruit-eating bats, Dermanura (Phyllostomidae: Stenodermatinae). Acta Chiropterologica, 11(2), 279-288.|Hoofer, S. R., Solari, S., Larsen, P. A., Bradley, R. D., & Baker, R. J. (2008). Phylogenetics of the fruit-eating bats (Phyllostomidae: Artibeina) inferred from mitochondrial DNA sequences. Occasional Papers of the Museum of Texas Tech University, 277, 1-15.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.|York, H. A., Rodríguez-Herrera, B., Laval, R. K., Timm, R. M., & Lindsay, K. E. (2019). Field key to the bats of Costa Rica and Nicaragua. Journal of Mammalogy, 100(6), 1726-1749.|Turcios-Casco, M. A., Ávila-Palma, H. D., LaVal, R. К., Stevens, R. D., Ordoñez-Trejo, E. J., Soler-Orellana, J. A., & Ordoñez-Mazier, D. I. (2020). A systematic revision of the bats (Chiroptera) of Honduras: an updated checklist with corroboration of historical specimens and new records. Zoosystematics and Evolution, 96, 411.|				Colombia|Ecuador|Peru	South America	Neotropic	LC	0	0	0	Artibeus_ravus	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Dermanura_rava	1005016	23	Little Fruit-eating Bat	Amber Fruit-eating Bat|Yellowish Little Fruit-eating Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Stenodermatinae	Stenodermatini	Dermanura	NA	rava	G. S. Miller	0	Dermanura rava	Miller, G.S., Jr. 1902-09-12. Twenty new American bats. Proceedings of the Academy of Natural Sciences of Philadelphia 54(2):389-412.	https://www.biodiversitylibrary.org/page/10309535	USNM:MAMM:113338	holotype	http://n2t.net/ark:/65665/373e4b913-d937-4367-8702-ceaa68acc042	"San Javier [Esmeraldas], northern Ecuador."			split from A. phaeotis; moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus	Solari, S., Hoofer, S. R., Larsen, P. A., Brown, A. D., Bull, R. J., Guerrero, J. A., ... & Baker, R. J. (2009). Operational criteria for genetically defined species: analysis of the diversification of the small fruit-eating bats, Dermanura (Phyllostomidae: Stenodermatinae). Acta Chiropterologica, 11(2), 279-288.|Hoofer, S. R., Solari, S., Larsen, P. A., Bradley, R. D., & Baker, R. J. (2008). Phylogenetics of the fruit-eating bats (Phyllostomidae: Artibeina) inferred from mitochondrial DNA sequences. Occasional Papers of the Museum of Texas Tech University, 277, 1-15.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.|York, H. A., Rodríguez-Herrera, B., Laval, R. K., Timm, R. M., & Lindsay, K. E. (2019). Field key to the bats of Costa Rica and Nicaragua. Journal of Mammalogy, 100(6), 1726-1749.|Turcios-Casco, M. A., Ávila-Palma, H. D., LaVal, R. К., Stevens, R. D., Ordoñez-Trejo, E. J., Soler-Orellana, J. A., & Ordoñez-Mazier, D. I. (2020). A systematic revision of the bats (Chiroptera) of Honduras: an updated checklist with corroboration of historical specimens and new records. Zoosystematics and Evolution, 96, 411.|				Colombia|Ecuador|Peru	South America	Neotropic	LC	0	0	0	Artibeus_ravus	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Dermanura		rava	Miller	1902	0	Proc. Acad. Nat. Sci. Phil.	60:44:00	Amber Fruit-eating Bat	None.	Ecuador, Esmeraldas, San Javier.	W Colombia	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/83683265/83683270/' target='_blank'>Least Concern</a>	Subgenus Dermanura. Previously considered a subspecies of phaeotis, but see Hoofer et al. (2008) and Solari et al. (2009).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	Artibeina	Artibeus ravus; Dermanura rava; Dermanura rava; Dermanura rava; Dermanura rava; rava; Dermanure jaunatre; Kleiner Fruchtvampir; Artibeoamarillento; Amber Fruit-eating Bat; Yellowish Little Fruit-eating Bat; Little Fruit-eating Bat; Amber Fruit-eating Bat; Yellowish Little Fruit-eating Bat; Amber Fruit-eating Bat; D. rava