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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L218	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Artibeus phaeotis	Artibeus phaeotis	Artibeus phaeotis	Artibeus phaeotis	Artibeus phaeotis	Artibeus phaeotis	Artibeus phaeotis	Dermanura phaeotis	Dermanura phaeotis	Dermanura phaeotis	Dermanura phaeotis	Dermanura phaeotis	Dermanura phaeotis	Dermanura phaeotis	Dermanura phaeotis		[MSW2] Subgenus Dermanura. Includes nanus and turpis; see Jones and Lawlor (1965:412) and Davis (1970). For including ravus and other synomyms see Timm (1985, Mammalian Species, 235).; [MSW3] Subgenus Dermanura. Includes nanus and turpis; see Jones and Lawlor (1965) and Davis (1970). For including ravus and other synomyms see Timm (1985) and Handley (1987).; [HMW] Dermanura phaeotis G. S. Miller, 1902 , “Chichen Itza [= Chichén-Itzd], Yucatan ,” Mexico . Artibeus turpis from Tabasco (southern Mexico ), described by RK. Andersen in 1906, 1s a junior synonym of A. p. phaeotis . Considered in the genus Dermanura by some authors but reassigned to Artibeus (subgenus Dermanura ) after reinterpretation of molecular results. Some authors recognized A. ravus as a synonym of A. phaeotis , but according to molecular evidence, they are considered different species. Three subspecies recognized.; [batnames2022] Subgenus Dermanura . Includes nanus and turpis ; see Jones and Lawlor (1965) and Davis (1970). Does not include ravus (see Hoofer et al. 2008 and Solari et al. 2009). For including other synomyms see Timm (1985) and Handley (1987).; [MDD2022] previously included A. ravus; moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus; [IUCN] The definition of this species has changed through the years and distinct researchers, Handley (1987) included A. ravus as a synonym, without further discussion, but previously Davis (1970) also synonymized other names under A. phaeotis . This view was kept by subsequent authors, like Simmons (2005). However, the molecular analyses by Solari et al. (2009) plus the side-by-side comparison of the type specimens (S. Solari pers. comm.) showed these two names represent distinct entities, and therefore valid species.; [batnames2023] Subgenus Dermanura . Includes nanus and turpis ; see Jones and Lawlor (1965) and Davis (1970). Does not include ravus (see Hoofer et al. 2008 and Solari et al. 2009). For including other synomyms see Timm (1985) and Handley (1987).; [MDD2023] previously included A. ravus; moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus; [MDD2025_2.0] previously included A. ravus; moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus; [batnames2025_1.7] Subgenus Dermanura. Includes nanus and turpis; see Jones and Lawlor (1965) and Davis (1970). Does not include ravus (see Hoofer et al. 2008 and Solari et al. 2009). For including other synomyms see Timm (1985) and Handley (1987).; [MDD2025_2.2] previously included A. ravus; moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus				nanus, turpis, ravus		nanus, palatinus, ravus, turpis.	nanus, palatinus, phaeotis, ravus	phaeotis, nanus, palatinus, ravus	turpis	phaeotis, nanus, palatinus	ravus	phaeotis, nanus, palatinus	phaeotis - turpis	phaeotis, nanus, turpis, palatinus	The definition of this species has changed through the years and distinct researchers, Handley (1987) included A. ravus as a synonym, without further discussion, but previously Davis (1970) also synonymized other names under A. phaeotis . This view was kept by subsequent authors, like Simmons (2005). However, the molecular analyses by Solari et al. (2009) plus the side-by-side comparison of the type specimens (S. Solari pers. comm.) showed these two names represent distinct entities, and therefore valid species.	phaeotis, nanus, palatinus	phaeotis - turpis	phaeotis, nanus, turpis, palatinus	phaeotis, nanus, turpis, palatina, nana 	nanus, palatinus, phaeotis	phaeotis - turpis	phaeotis G. S. Miller, 1902|nanus (Andersen, 1906)|turpis (Andersen, 1906)|palatina (W. B. Davis, 1970)|nana R. D. Owen, 1987 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		W Mexico – E Peru	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Artibeus phaeotis	Mexico, Yucatan, Chichen-Itza.	Miller	1902	Proc. Acad. Nat. Sci. Phila., 54:405.	Distribution: Ranging from tropical Mexico to northwestern Ecuador, southeastern Colombia, and Guyana. A Peruvian record probably refers to A. anderseni.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Dwarf fruit-eating bat (Pygmy fruit-eating bat)	W Mexico – E Peru; Guyana	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Miller	1902	Proc. Acad. Nat. Sci. Philadelphia, 54:405.	Subgenus Dermanura. Includes nanus and turpis; see Jones and Lawlor (1965:412) and Davis (1970). For including ravus and other synomyms see Timm (1985, Mammalian Species, 235).	Veracruz and Sinaloa (Mexico) to Ecuador and Guyana.	Mexico, Yucatan, Chichen-Itza.		MILLER	1902	Size relatively small (forearm length, 34-42 mm; greatest length of skull, 17-21 mm). Preorbital and postorbital processes absent. Last upper and lower molars absent. Forehead concave but transition to rostrum varying from gradual to abrupt. Upper molars relatively narrow. Uropatagium virtually hairless.	Distribution: Ranging from tropical Mexico to northwestern Ecuador, southeastern Colombia, and Guyana. A Peruvian record probably refers to A. anderseni.	Four subspecies are here recognized:	A. p. nanus (western Mexico from Sinaloa to Guerrero), A. p. palatinus (Pacific side of Middle America from Oaxaca to Costa Rica), A. p. phaeotis (Atlantic side of Middle America from Veracruz to Costa Rica), A. p. ravus (Panama and South American range).	92	species	A. phaeotis	MILLER	1902	Dermanura	subgenus	Artibeus phaeotis				Size relatively small (forearm length, 34-42 mm; greatest length of skull, 17-21 mm). Preorbital and postorbital processes absent. Last upper and lower molars absent. Forehead concave but transition to rostrum varying from gradual to abrupt. Upper molars relatively narrow. Uropatagium virtually hairless.	Four subspecies are here recognized:		14. A.phaeotis (MILLER 1902).	14	_D. p. nanus_ (Andersen, 1906); _D. p. palatina_ (Davis, 1970); _D. p. phaeotis_ Miller, 1902 (synonyms: _turpis_ (Andersen, 1906))			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Stenodermatinae	Stenodermatini	Artibeus phaeotis	Artibeus	Dermanura	phaeotis	Miller	y	1902		Proc. Acad. Nat. Sci. Phil.	54		405		Pygmy Fruit-eating Bat	Mexico, Yucatán, Chichén-Itzá.	Veracruz, Sinaloa, and Michoacan (Mexico) south to Ecuador and Guyana.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	turpis K. Andersen, 1906; nanus K. Andersen, 1906; palatinus Davis, 1970; ravus Miller, 1902.	Subgenus Dermanura. Includes nanus and turpis; see Jones and Lawlor (1965) and Davis (1970). For including ravus and other synomyms see Timm (1985) and Handley (1987).	03A687BCFFECFFEF1643F9B0FBA4F0BF	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	577	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFFECFFEF1643F9B0FBA4F0BF.xml	Artibeus phaeotis	Phyllostomidae	Artibeus	phaeotis		1902	Dermanure pygmée @fr | Zwergfruchtvampir @de | Artibeopigmeo @es	Dermanura phaeotis G. S. Miller, 1902 , “Chichen Itza [= Chichén-Itzd], Yucatan ,” Mexico . Artibeus turpis from Tabasco (southern Mexico ), described by RK. Andersen in 1906, 1s a junior synonym of A. p. phaeotis . Considered in the genus Dermanura by some authors but reassigned to Artibeus (subgenus Dermanura ) after reinterpretation of molecular results. Some authors recognized A. ravus as a synonym of A. phaeotis , but according to molecular evidence, they are considered different species. Three subspecies recognized.	A.p.phaeotisG.S.Miller,1902—E&SEMexico(fromVeracruztoYucatanPeninsula),NGuatemala,Belize,Honduras,ENicaragua,ECostaRica,PanamaandWColombia. A.p.nanusK.Andersen,1906—W&SWMexico(fromSinaloatoOaxaca). A. p. palatinus W. B. Davis, 1970 — S Mexico (SW Chiapas ), S Guatemala , W El Salvador , W Nicaragua , and W Costa Rica . Records from W Ecuador and N Peru are unverified; they could represent the Little Fruit-eating Bat (A. ravus), which was thought to be part of this taxon. Those from Venezuela , N Brazil , and Guyana probably represent misidentifications.	Head—body 51-60 mm (tailless), ear 14-18 mm, hindfoot 8-12 mm, forearm 35-2-41-8 mm; weight 8-15-6 g. Skull of the Pygmy Fruit-eating Batis small, with short and broad rostrum, and is characterized by high cranial dome. Dorsal fur is light pale brown or grayish, short (4-6 mm), faintly tricolored, and soft; it extends to base of forearm. Light brown ocular ring is present, with two well-marked white facial stripes. Ventralfur is slightly paler than dorsum. Noseleaf is thick and pointed compared with other species of Stenodermatinae. Ears are rounded, moderated in size, and brown and usually have conspicuous whitish or pale yellow tips. Tragus is pointed, with many lobes of different sizes on the outside, giving a serrated appearance. Fur covers more than one-half of each forearm. Wing membranes are black, except for second interdigital membrane that lacks pigmentation. Tail membrane is pale, moderated in size (9-11 mm), U-shaped, and almost completely naked. Dental formulais12/2,C1/1,P 2/2, M 2/2 ( x2 ) = 28. Upper and lower molars are relatively small compared with other species of similarly sized Artibeus .	Seemingly associated with tropical deciduous forest, tropical rainforest, tropical dry forests, shrublands, and second growth forests and plantations (banana). The Pygmy Fruit-eating Bat might occur in dryareas (e.g. Yucatan Peninsula), using tropical forest islands or “petenes” of coastal wetlands.	The Pygmy Fruit-eating Bat is mainly frugivorous and occasionally eats insects and pollen. In dry forests of Costa Rica , eleven plants species are used for food and ten plant genera in Panama , including Ficus spp. ( Moraceae ), Cecropia spp. ( Urticaceae ), and Spondias spp. ( Anacardiaceae ).	Breeding pattern of the PygmyFruit-eating Bat has been characterized as bimodal polyestry. In Costa Rica , there is information of births at the end of dry seasons (April) and middle of rainy seasons (August-September). Females have one young per pregnancy. In the Yucatan Peninsula, breeding season occurs in rainy season.	The Pygmy Fruit-eating Bat is nocturnal. It uses caves or tents made from leaves of bananas, Heliconia ( Heliconiaceae ), Philodendron ( Araceae ), and palms. Long leaves are chewed along edge of midrib to form an “upturned boat” shape; fanshaped leaves are cut in a polygonal shape around their bases.	Pygmy Fruit-eating Bats roost alone or in small groups.	Classified as Least Concern on The IUCN Red List (as Dermanura phaeotis ).	Andersen (1906b) | Davis (1970a) | LaVal & Rodriguez-Herrera (2002) | Le6én & Montiel (2006) | Miller et al. (2015c) | Reid (2009) | Rodriguez-Herrera et al. (2007) | Timm (1985, 1987)	https://zenodo.org/record/6458999/files/figure.png	202. Pygmy Fruit-eating Bat Artibeus phaeotis French: Dermanure pygmée / German: Zwergfruchtvampir / Spanish: Artibeo pigmeo Taxonomy. Dermanura phaeotis G. S. Miller, 1902 , “Chichen Itza [= Chichén-Itzd], Yucatan ,” Mexico . Artibeus turpis from Tabasco (southern Mexico ), described by RK. Andersen in 1906, 1s a junior synonym of A. p. phaeotis . Considered in the genus Dermanura by some authors but reassigned to Artibeus (subgenus Dermanura ) after reinterpretation of molecular results. Some authors recognized A. ravus as a synonym of A. phaeotis , but according to molecular evidence, they are considered different species. Three subspecies recognized. Subspecies and Distribution. A.p.phaeotisG.S.Miller,1902—E&SEMexico(fromVeracruztoYucatanPeninsula),NGuatemala,Belize,Honduras,ENicaragua,ECostaRica,PanamaandWColombia. A.p.nanusK.Andersen,1906—W&SWMexico(fromSinaloatoOaxaca). A. p. palatinus W. B. Davis, 1970 — S Mexico (SW Chiapas ), S Guatemala , W El Salvador , W Nicaragua , and W Costa Rica . Records from W Ecuador and N Peru are unverified; they could represent the Little Fruit-eating Bat (A. ravus), which was thought to be part of this taxon. Those from Venezuela , N Brazil , and Guyana probably represent misidentifications. Descriptive notes. Head—body 51-60 mm (tailless), ear 14-18 mm, hindfoot 8-12 mm, forearm 35-2-41-8 mm; weight 8-15-6 g. Skull of the Pygmy Fruit-eating Batis small, with short and broad rostrum, and is characterized by high cranial dome. Dorsal fur is light pale brown or grayish, short (4-6 mm), faintly tricolored, and soft; it extends to base of forearm. Light brown ocular ring is present, with two well-marked white facial stripes. Ventralfur is slightly paler than dorsum. Noseleaf is thick and pointed compared with other species of Stenodermatinae. Ears are rounded, moderated in size, and brown and usually have conspicuous whitish or pale yellow tips. Tragus is pointed, with many lobes of different sizes on the outside, giving a serrated appearance. Fur covers more than one-half of each forearm. Wing membranes are black, except for second interdigital membrane that lacks pigmentation. Tail membrane is pale, moderated in size (9-11 mm), U-shaped, and almost completely naked. Dental formulais12/2,C1/1,P 2/2, M 2/2 ( x2 ) = 28. Upper and lower molars are relatively small compared with other species of similarly sized Artibeus . Habitat. Seemingly associated with tropical deciduous forest, tropical rainforest, tropical dry forests, shrublands, and second growth forests and plantations (banana). The Pygmy Fruit-eating Bat might occur in dryareas (e.g. Yucatan Peninsula), using tropical forest islands or “petenes” of coastal wetlands. Food and Feeding. The Pygmy Fruit-eating Bat is mainly frugivorous and occasionally eats insects and pollen. In dry forests of Costa Rica , eleven plants species are used for food and ten plant genera in Panama , including Ficus spp. ( Moraceae ), Cecropia spp. ( Urticaceae ), and Spondias spp. ( Anacardiaceae ). Breeding. Breeding pattern of the PygmyFruit-eating Bat has been characterized as bimodal polyestry. In Costa Rica , there is information of births at the end of dry seasons (April) and middle of rainy seasons (August-September). Females have one young per pregnancy. In the Yucatan Peninsula, breeding season occurs in rainy season. Activity patterns. The Pygmy Fruit-eating Bat is nocturnal. It uses caves or tents made from leaves of bananas, Heliconia ( Heliconiaceae ), Philodendron ( Araceae ), and palms. Long leaves are chewed along edge of midrib to form an “upturned boat” shape; fanshaped leaves are cut in a polygonal shape around their bases. Movements, Home range and Social organization. Pygmy Fruit-eating Bats roost alone or in small groups. Status and Conservation. Classified as Least Concern on The IUCN Red List (as Dermanura phaeotis ). Bibliography. Andersen (1906b), Davis (1970a), LaVal & Rodriguez-Herrera (2002), Le6én & Montiel (2006), Miller et al. (2015c), Reid (2009), Rodriguez-Herrera et al. (2007), Timm (1985, 1987).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Dermanura phaeotis	Dermanura		phaeotis	Miller	1902	0	Proc. Acad. Nat. Sci. Phil.	60:45:00	Pygmy Fruit-eating Bat	 turpis K. Andersen, 1906; <b> nanus </b> K. Andersen, 1906; <b>palatinus</b> Davis, 1970.	Mexico, Yucat&aacute;n, ChichÃ©n-Itz&aacute;.	Veracruz, Sinaloa, and Michoacan (Mexico) south to Ecuador and Guyana.	Not listed.	Least Concern	Subgenus Dermanura . Includes nanus and turpis ; see Jones and Lawlor (1965) and Davis (1970). Does not include ravus (see Hoofer et al. 2008 and Solari et al. 2009). For including other synomyms see Timm (1985) and Handley (1987).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Dermanura phaeotis	23	Pygmy Fruit-eating Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	STENODERMATINAE	STENODERMATINI	Dermanura	NA	phaeotis	G. S. Miller	1902	0	Dermanura_phaeotis	Miller, G. S., Jr. (1902). Twenty New American Bats. Proceedings of the Academy of Natural Sciences of Philadelphia, 54, 405.	https://www.biodiversitylibrary.org/page/10309520#page/439/mode/1up	USNM 108176		"Chichen Itza [= ChichÃ©n-ItzÃ¡], Yucatan," Mexico.			phaeotis G. S. Miller, 1902|nanus (K. Andersen, 1906)|turpis (K. Andersen, 1906)|palatinus (W. B. Davis, 1970)	previously included A. ravus; moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus	Solari, S., Hoofer, S. R., Larsen, P. A., Brown, A. D., Bull, R. J., Guerrero, J. A., ... & Baker, R. J. (2009). Operational criteria for genetically defined species: analysis of the diversification of the small fruit-eating bats, Dermanura (Phyllostomidae: Stenodermatinae). Acta Chiropterologica, 11(2), 279-288.|Hoofer, S. R., Solari, S., Larsen, P. A., Bradley, R. D., & Baker, R. J. (2008). Phylogenetics of the fruit-eating bats (Phyllostomidae: Artibeina) inferred from mitochondrial DNA sequences. Occasional Papers of the Museum of Texas Tech University, 277, 1-15.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.|York, H. A., RodrÃ­guez-Herrera, B., Laval, R. K., Timm, R. M., & Lindsay, K. E. (2019). Field key to the bats of Costa Rica and Nicaragua. Journal of Mammalogy, 100(6), 1726-1749.|Turcios-Casco, M. A., Ãvila-Palma, H. D., LaVal, R. Ðš., Stevens, R. D., OrdoÃ±ez-Trejo, E. J., Soler-Orellana, J. A., & OrdoÃ±ez-Mazier, D. I. (2020). A systematic revision of the bats (Chiroptera) of Honduras: an updated checklist with corroboration of historical specimens and new records. Zoosystematics and Evolution, 96, 411.|	Mexico|Belize|Guatemala|Honduras|El Salvador|Nicaragua|Costa Rica|Panama|Colombia|Ecuador?|Peru?|Venezuela?|Guyana?|Brazil?	North America|South America	Nearctic|Neotropic	LC	0	0	0	Artibeus_phaeotis	0	oldname match	Artibeus_phaeotis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Dermanura phaeotis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Dermanura	phaeotis	Miller, 1902	The definition of this species has changed through the years and distinct researchers, Handley (1987) included A. ravus as a synonym, without further discussion, but previously Davis (1970) also synonymized other names under A. phaeotis . This view was kept by subsequent authors, like Simmons (2005). However, the molecular analyses by Solari et al. (2009) plus the side-by-side comparison of the type specimens (S. Solari pers. comm.) showed these two names represent distinct entities, and therefore valid species.	20000000	Dermanura phaeotis	Least Concern		2015	2015-07-20 00:00:00 UTC	3.1	English	This species is listed as Least Concern because of its wide distribution, presumed large population, occurrence in a number of protected areas, tolerance to some degree of habitat modification, and because it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	This species is difficult to distinguish in the field from Dermanura watsoni (Reid 1997). It is found in lowland evergreen and deciduous forests up to 1,200 m asl. It is found in secondary forests and disturbed habitats. ;It feeds in highly cluttered space as a gleaning frugivore (Kalko et al. 1996) but it occasionally consumes pollen and insects. It may modify leaves to form tent roosts. Reproduction is seasonally polyestrus, with birth peaks around April and September.	There are no major threats.	It is locally common.	Stable	It occurs in Veracruz, Sinaloa and Michoacan (Mexico) south to Ecuador and Guyana (Simmons 2005).	This species is not used.	Terrestrial	It is found in protected areas.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Dermanura		phaeotis	Miller	1902	0	Proc. Acad. Nat. Sci. Phil.	60:45:00	Pygmy Fruit-eating Bat	 turpis K. Andersen, 1906; <b> nanus </b> K. Andersen, 1906; <b>palatinus</b> Davis, 1970.	Mexico, Yucat&aacute;n, ChichÃ©n-Itz&aacute;.	Veracruz, Sinaloa, and Michoacan (Mexico) south to Ecuador and Guyana.	Not listed.	Least Concern	Subgenus Dermanura . Includes nanus and turpis ; see Jones and Lawlor (1965) and Davis (1970). Does not include ravus (see Hoofer et al. 2008 and Solari et al. 2009). For including other synomyms see Timm (1985) and Handley (1987).	Dermanura phaeotis	1005014	23	Pygmy Fruit-eating Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	STENODERMATINAE	STENODERMATINI	Dermanura	NA	phaeotis	G. S. Miller	1902	0	Dermanura_phaeotis	Miller, G. S., Jr. (1902). Twenty New American Bats. Proceedings of the Academy of Natural Sciences of Philadelphia, 54, 405.	https://www.biodiversitylibrary.org/page/10309520#page/439/mode/1up	USNM 108176		"Chichen Itza [= ChichÃ©n-ItzÃ¡], Yucatan," Mexico.			phaeotis G. S. Miller, 1902|nanus (K. Andersen, 1906)|turpis (K. Andersen, 1906)|palatinus (W. B. Davis, 1970)	previously included A. ravus; moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus	Solari, S., Hoofer, S. R., Larsen, P. A., Brown, A. D., Bull, R. J., Guerrero, J. A., ... & Baker, R. J. (2009). Operational criteria for genetically defined species: analysis of the diversification of the small fruit-eating bats, Dermanura (Phyllostomidae: Stenodermatinae). Acta Chiropterologica, 11(2), 279-288.|Hoofer, S. R., Solari, S., Larsen, P. A., Bradley, R. D., & Baker, R. J. (2008). Phylogenetics of the fruit-eating bats (Phyllostomidae: Artibeina) inferred from mitochondrial DNA sequences. Occasional Papers of the Museum of Texas Tech University, 277, 1-15.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.|York, H. A., RodrÃ­guez-Herrera, B., Laval, R. K., Timm, R. M., & Lindsay, K. E. (2019). Field key to the bats of Costa Rica and Nicaragua. Journal of Mammalogy, 100(6), 1726-1749.|Turcios-Casco, M. A., Ãvila-Palma, H. D., LaVal, R. Ðš., Stevens, R. D., OrdoÃ±ez-Trejo, E. J., Soler-Orellana, J. A., & OrdoÃ±ez-Mazier, D. I. (2020). A systematic revision of the bats (Chiroptera) of Honduras: an updated checklist with corroboration of historical specimens and new records. Zoosystematics and Evolution, 96, 411.|				Mexico|Belize|Guatemala|Honduras|El Salvador|Nicaragua|Costa Rica|Panama|Colombia|Ecuador?|Peru?|Venezuela?|Guyana?|Brazil?	North America|South America	Nearctic|Neotropic	LC	0	0	0	Artibeus_phaeotis	0	oldname match	Artibeus_phaeotis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Dermanura_phaeotis	1005014	23	Pygmy Fruit-eating Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Stenodermatinae	Stenodermatini	Dermanura	NA	phaeotis	G. S. Miller	0	Dermanura phaeotis	Miller, G.S., Jr. 1902-09-12. Twenty new American bats. Proceedings of the Academy of Natural Sciences of Philadelphia 54(2):389-412.	https://www.biodiversitylibrary.org/page/10309536	USNM:MAMM:108176	holotype	http://n2t.net/ark:/65665/32ea1c147-1db7-4703-9290-be0a629ad219	"Chichen Itza [= ChichÃ©n-ItzÃ¡], Yucatan," Mexico.			previously included A. ravus; moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus	Solari, S., Hoofer, S. R., Larsen, P. A., Brown, A. D., Bull, R. J., Guerrero, J. A., ... & Baker, R. J. (2009). Operational criteria for genetically defined species: analysis of the diversification of the small fruit-eating bats, Dermanura (Phyllostomidae: Stenodermatinae). Acta Chiropterologica, 11(2), 279-288.|Hoofer, S. R., Solari, S., Larsen, P. A., Bradley, R. D., & Baker, R. J. (2008). Phylogenetics of the fruit-eating bats (Phyllostomidae: Artibeina) inferred from mitochondrial DNA sequences. Occasional Papers of the Museum of Texas Tech University, 277, 1-15.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.|York, H. A., RodrÃ­guez-Herrera, B., Laval, R. K., Timm, R. M., & Lindsay, K. E. (2019). Field key to the bats of Costa Rica and Nicaragua. Journal of Mammalogy, 100(6), 1726-1749.|Turcios-Casco, M. A., Ãvila-Palma, H. D., LaVal, R. Ðš., Stevens, R. D., OrdoÃ±ez-Trejo, E. J., Soler-Orellana, J. A., & OrdoÃ±ez-Mazier, D. I. (2020). A systematic revision of the bats (Chiroptera) of Honduras: an updated checklist with corroboration of historical specimens and new records. Zoosystematics and Evolution, 96, 411.|				Mexico|Belize|Guatemala|Honduras|El Salvador|Nicaragua|Costa Rica|Panama|Colombia|Ecuador?|Peru?|Venezuela?|Guyana?|Brazil?	North America|South America	Nearctic|Neotropic	LC	0	0	0	Artibeus_phaeotis	0	oldname match	Artibeus_phaeotis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Dermanura		phaeotis	Miller	1902	0	Proc. Acad. Nat. Sci. Phil.	60:45:00	Pygmy Fruit-eating Bat	turpis K. Andersen, 1906; nanus K. Andersen, 1906; palatinus Davis, 1970.	Mexico, Yucat&aacute;n, ChichÃ©n-Itz&aacute;.	Veracruz, Sinaloa, and Michoacan (Mexico) south to Ecuador and Guyana.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/83683287/21997769/' target='_blank'>Least Concern</a>	Subgenus Dermanura. Includes nanus and turpis; see Jones and Lawlor (1965) and Davis (1970). Does not include ravus (see Hoofer et al. 2008 and Solari et al. 2009). For including other synomyms see Timm (1985) and Handley (1987).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	Artibeina	Artibeus phaeotis; Artibeus phaeotis; Dermanura phaeotis; Dermanura phaeotis; Dermanura phaeotis; Dermanura phaeotis; phaeotis; nanus; palatinus; ravus; turpis; phaeotis; nanus; palatinus; ravus; nanus; palatinus; turpis; phaeotis; nanus; turpis; palatinus; Dermanure pygmée; Zwergfruchtvampir; Artibeopigmeo; Pygmy Fruit-eating Bat; Pygmy Fruit-eating Bat; Pygmy Fruit-eating Bat; Artibeus phaeotis phaeotis; Artibeus phaeotis nanus; Artibeus phaeotis palatinus; Artibeus phaeotis ravus; Artibeus phaeotis turpis; Dermanura phaeotis phaeotis; Dermanura phaeotis nanus; Dermanura phaeotis palatinus; Dermanura phaeotis ravus; Dermanura phaeotis turpis; Artibeus phaeotis; Artibeus nanus; Artibeus palatinus; Artibeus ravus; Artibeus nanus; Artibeus palatinus; Artibeus turpis; Dermanura phaeotis; Dermanura nanus; Dermanura palatinus; Dermanura ravus; Dermanura nanus; Dermanura palatinus; Dermanura turpis; Dermanura phaeotis phaeotis; D. phaeotis
