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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L217	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	Artibeus gnomus	Artibeus glaucus [synonym of]	N/A	Artibeus gnomus	Artibeus gnomus	Dermanura gnoma	Dermanura gnoma	Dermanura gnoma	Dermanura gnoma	Dermanura gnoma	Dermanura gnomus	Dermanura gnomus	Dermanura gnomus		[MSW3] Subgenus Dermanura. Distinct from cinereus and glaucus; see Handley (1987), Brosset and Charles-Dominique (1990), and Simmons and Voss (1998).; [HMW] Artibeus gnomus Handley, 1987 , “El Manaco (= Km 74), 39 km SE El Dorado, Bolivar , Venezuela , 150 m .” Artibeus gnomus is placed in Dermanura by some authors, but itis reassigned to Artibeus (subgenus Dermanura ) after reinterpretation of molecular results. Before its description, some populations were referred to as A. pumilio , although that name is now associated with A. glaucus . Monotypic.; [batnames2022] Subgenus Dermanura .  Distinct from cinereus and glaucus ; see Handley (1987), Brosset and Charles-Dominique(1990), and Simmons and Voss (1998).; [MDD2022] moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus; [IUCN]  This species was originally described as different from other small Artibeus (sensu Handley 1987), but Koopman (1994) listed it as a synonym of A. cinereus (under A. c. pumilio , along with glaucus ). However, the characters used in the original description clearly differentiated it from similar taxa, and most recent authors (Simmons 2005) listed it as distinct. Molecular analyses (Hoofer et al. 2008, Solari et al. 2009) also validate this distinction.; [batnames2023] Subgenus Dermanura .  Distinct from cinereus and glaucus ; see Handley (1987), Brosset and Charles-Dominique(1990), and Simmons and Voss (1998).; [MDD2023] moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus; [MDD2025_2.0] moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus; [batnames2025_1.7] Subgenus Dermanura. Distinct from cinereus and glaucus; see Handley (1987), Brosset and Charles-Dominique(1990), and Simmons and Voss (1998). The species epithet is a noun in apposition and therefore maintains the original spelling.; [MDD2025_2.2] moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus														gnoma	 This species was originally described as different from other small Artibeus (sensu Handley 1987), but Koopman (1994) listed it as a synonym of A. cinereus (under A. c. pumilio , along with glaucus ). However, the characters used in the original description clearly differentiated it from similar taxa, and most recent authors (Simmons 2005) listed it as distinct. Molecular analyses (Hoofer et al. 2008, Solari et al. 2009) also validate this distinction.			gnoma	gnomus, gnoma			gnomus (Handley, 1987)|gnoma R. D. Owen, 1991 [incorrect subsequent spelling]						N/A							Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Venezuela, Peru, Brazil, Guyana; ref. 4.122																															NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Stenodermatinae	Stenodermatini	Artibeus gnomus	Artibeus	Dermanura	gnomus	Handley		1987		Fieldiana Zool.	39		167		Dwarf Fruit-eating Bat	Venezuela, Bolívar, 59 km SE El Dorado, El Manaco.	Ecuador, Peru, Bolivia, Amazonian Brazil, Venezuela, Guianas.	IUCN 2003 – Not listed (lapsus); IUCN/SSC Action Plan (2001) – Lower Risk (lc).		Subgenus Dermanura. Distinct from cinereus and glaucus; see Handley (1987), Brosset and Charles-Dominique (1990), and Simmons and Voss (1998).	03A687BCFFD1FFD1138AFC7AFE51F910	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	580	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFFD1FFD1138AFC7AFE51F910.xml	Artibeus gnomus	Phyllostomidae	Artibeus	gnomus	Handley	1987	Dermanure naine @fr | Zwergsilberfruchtvampir @de | Artibeoenano @es | Dwarf Little Fruit-eating Bat @en	Artibeus gnomus Handley, 1987 , “El Manaco (= Km 74), 39 km SE El Dorado, Bolivar , Venezuela , 150 m .” Artibeus gnomus is placed in Dermanura by some authors, but itis reassigned to Artibeus (subgenus Dermanura ) after reinterpretation of molecular results. Before its description, some populations were referred to as A. pumilio , although that name is now associated with A. glaucus . Monotypic.	South America, E of Andes, including E Venezuela , the Guianas, E Colombia , E Ecuador , E Peru , Bolivia , and Brazil .	Head—body 44-54 mm (tailless), ear 14-19 mm, hindfoot 9-10 mm, forearm 34-38 mm; weight 8-13 g. The Dwarf Fruit-eating Bat has gray-brown to brown dorsal pelage, and hairs are faintly tricolored (very pale basal band followed by medium to dark gray band and light-brown tip). Ventral fur is paler. Facial stripes are bright white and sharply defined. Noseleaf and horseshoe are gray-brown medially and yellowish to the sides; ears are brownish, edged with yellowish narrow rim, but entire tragusis yellowish. Proximal one-half (dorsal) of forearm is covered with dense and long body fur. Wing membranes are blackish and attach to base of outer toes. Short brown hairs sparsely cover dorsal surface of leg. Uropatagium is sooty brown, with dorsal surface almost naked, and deeply emarginated. Cranially, rostrum is very short, with supraorbital region well inflated. Dental formulais 12/2, C1/1,P 2/2, M 2/2 ( x2 ) = 28. I ! are distinctly bilobated.	Forested areas, pristine humid lowland forests, forest edges, crop fields in recovery (second growth forests), and cerrado formations and savannas at elevations of 100-680 m.	The Dwarf Fruit-eating Bat is considered frugivorous and feeds on fruits in the canopy; however,its specific habits are unknown. Its diet includes figs ( Ficus spp., Moraceae ) and Cecropia sp. ( Urticaceae ).	Pregnant and lactating Dwarf Fruit-eating Bats have been found in January— April in Venezuela , French Guiana , and eastern Brazil (Para); June-July in Venezuela and Brazil ( Mato Grosso ); and October-November in Venezuela and Peru . Based on these data, bimodal polyestry appears to be the reproductive pattern.	In Peru , one Dwarf Fruiteating Bat was located roosting under a cut leaf of Monstera lechleriana ( Araceae ).	Roosting groups of Dwarf Fruiteating Bat have had 2-8 individuals, but no details were given about the make-up of these groups.	Classified as Least Concern on The IUCNRed List (as Dermanura gnoma ). The Dwarf Fruiteating Bat has a wide distribution and occurs in protected areas. Nevertheless, its natural history is poorly understood due to confusion with similar species.	Ascorra et al. (1996) | Bernard & Fenton (2002) | Handley (1987) | Hershkovitz (1949) | Lim, Engstrom, Patton & Bickham (2008) | Marques-Aguiar (2008a) | Solari, Hoofer et al. (2009) | Solari, Munoz-Saba et al. (2013) | Timm (1987)		209. Dwarf Fruit-eating Bat Artibeus gnomus French: Dermanure naine / German: Zwergsilberfruchtvampir / Spanish: Artibeo enano Other common names: Dwarf Little Fruit-eating Bat Taxonomy. Artibeus gnomus Handley, 1987 , “El Manaco (= Km 74), 39 km SE El Dorado, Bolivar , Venezuela , 150 m .” Artibeus gnomus is placed in Dermanura by some authors, but itis reassigned to Artibeus (subgenus Dermanura ) after reinterpretation of molecular results. Before its description, some populations were referred to as A. pumilio , although that name is now associated with A. glaucus . Monotypic. Distribution. South America, E of Andes, including E Venezuela , the Guianas, E Colombia , E Ecuador , E Peru , Bolivia , and Brazil . Descriptive notes. Head—body 44-54 mm (tailless), ear 14-19 mm, hindfoot 9-10 mm, forearm 34-38 mm; weight 8-13 g. The Dwarf Fruit-eating Bat has gray-brown to brown dorsal pelage, and hairs are faintly tricolored (very pale basal band followed by medium to dark gray band and light-brown tip). Ventral fur is paler. Facial stripes are bright white and sharply defined. Noseleaf and horseshoe are gray-brown medially and yellowish to the sides; ears are brownish, edged with yellowish narrow rim, but entire tragusis yellowish. Proximal one-half (dorsal) of forearm is covered with dense and long body fur. Wing membranes are blackish and attach to base of outer toes. Short brown hairs sparsely cover dorsal surface of leg. Uropatagium is sooty brown, with dorsal surface almost naked, and deeply emarginated. Cranially, rostrum is very short, with supraorbital region well inflated. Dental formulais 12/2, C1/1,P 2/2, M 2/2 ( x2 ) = 28. I ! are distinctly bilobated. Habitat. Forested areas, pristine humid lowland forests, forest edges, crop fields in recovery (second growth forests), and cerrado formations and savannas at elevations of 100-680 m. Food and Feeding. The Dwarf Fruit-eating Bat is considered frugivorous and feeds on fruits in the canopy; however,its specific habits are unknown. Its diet includes figs ( Ficus spp., Moraceae ) and Cecropia sp. ( Urticaceae ). Breeding. Pregnant and lactating Dwarf Fruit-eating Bats have been found in January— April in Venezuela , French Guiana , and eastern Brazil (Para); June-July in Venezuela and Brazil ( Mato Grosso ); and October-November in Venezuela and Peru . Based on these data, bimodal polyestry appears to be the reproductive pattern. Activity patterns. In Peru , one Dwarf Fruiteating Bat was located roosting under a cut leaf of Monstera lechleriana ( Araceae ). Movements, Home range and Social organization. Roosting groups of Dwarf Fruiteating Bat have had 2-8 individuals, but no details were given about the make-up of these groups. Status and Conservation. Classified as Least Concern on The IUCNRed List (as Dermanura gnoma ). The Dwarf Fruiteating Bat has a wide distribution and occurs in protected areas. Nevertheless, its natural history is poorly understood due to confusion with similar species. Bibliography. Ascorra et al. (1996), Bernard & Fenton (2002), Handley (1987), Hershkovitz (1949), Lim, Engstrom, Patton & Bickham (2008), Marques-Aguiar (2008a), Solari, Hoofer et al. (2009), Solari, Munoz-Saba et al. (2013), Timm (1987).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Dermanura gnoma	Dermanura		gnoma	Handley	1987	1	Fieldiana Zool.	41:47:00	Dwarf Fruit-eating Bat	None.	Venezuela, Bolívar, 59 km SE El Dorado, El Manaco.	S Colombia, Ecuador, Peru, Bolivia, Brazil, Venezuela, Guianas.	Not listed.	Least Concern	Subgenus Dermanura .  Distinct from cinereus and glaucus ; see Handley (1987), Brosset and Charles-Dominique(1990), and Simmons and Voss (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Dermanura gnoma	23	Dwarf Fruit-eating Bat	Dwarf Little Fruit-eating Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	STENODERMATINAE	STENODERMATINI	Dermanura	NA	gnoma	Handley	1987	1	Artibeus_gnomus	Handley, C. O., Jr. (1987). New Species of Mammals from Northern South America: Fruit-Eating Bats, Genus Artibeus Leach. Fieldiana Zoology, new series 39, 167.	https://www.biodiversitylibrary.org/page/4465287#page/177/mode/1up	USNM 387534		"El Manaco (= Km 74), 59 km SE El Dorado, Bolívar, Venezuela, 150 m."			gnoma (Handley, 1987)	moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus	Hoofer, S. R., Solari, S., Larsen, P. A., Bradley, R. D., & Baker, R. J. (2008). Phylogenetics of the fruit-eating bats (Phyllostomidae: Artibeina) inferred from mitochondrial DNA sequences. Occasional Papers of the Museum of Texas Tech University, 277, 1-15.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.|York, H. A., Rodríguez-Herrera, B., Laval, R. K., Timm, R. M., & Lindsay, K. E. (2019). Field key to the bats of Costa Rica and Nicaragua. Journal of Mammalogy, 100(6), 1726-1749.|Turcios-Casco, M. A., Ávila-Palma, H. D., LaVal, R. К., Stevens, R. D., Ordoñez-Trejo, E. J., Soler-Orellana, J. A., & Ordoñez-Mazier, D. I. (2020). A systematic revision of the bats (Chiroptera) of Honduras: an updated checklist with corroboration of historical specimens and new records. Zoosystematics and Evolution, 96, 411.|	Colombia|Venezuela|Guyana|Suriname|French Guiana|Brazil|Ecuador|Peru|Bolivia	South America	Neotropic	LC	0	0	0	Artibeus_gnomus	0	oldname match	Artibeus_gnomus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	2129	Dermanura gnoma	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Dermanura	gnoma	(Handley, 1987)	 This species was originally described as different from other small Artibeus (sensu Handley 1987), but Koopman (1994) listed it as a synonym of A. cinereus (under A. c. pumilio , along with glaucus ). However, the characters used in the original description clearly differentiated it from similar taxa, and most recent authors (Simmons 2005) listed it as distinct. Molecular analyses (Hoofer et al. 2008, Solari et al. 2009) also validate this distinction.	100000000	Dermanura gnoma	Least Concern		2015	2015-07-20 00:00:00 UTC	3.1	English	This species is listed as Least Concern as it is widespread, is tolerant to a broad range of habitats, and is presumed to be more common than records show.	The ecology of the species is poorly known, but the genus generally feeds on small fruits and insects and roosts under banana-like leaves, and perhaps in tents made from palm leaves (Kunz et al . 1994). The genus is found in mature and disturbed lowland and montane rainforest, plantations and gardens, and cloud forest and deciduous forest (Emmons and Feer 1997). It is one of the smallest species in the genus.	Habitat loss occurs in some parts of its range, although this is not a major threat.	This species is apparently rare but as a canopy frugivore it may be undersampled, and it may be misindentified in the field. As a result it may be more abundant than indicated by current records.	Unknown	This species is found in Venezuela, the Guianas, Brazil, Ecuador and Peru (Handley 1987, Simmons and Voss 1998). It is also in Colombia (Rivas-Pava et al . 1996) and Bolivia (Anderson 1997). Within Brazil, the known distribution has expanded south to Espirito Santo state (Aguiar et al . 1995). It has been found up to 800 m asl in Mato Grosso (Handley 1987).	This species is not used.	Terrestrial	The recommended conservation action is to reduce habitat loss.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Dermanura		gnoma	Handley	1987	1	Fieldiana Zool. N.S.	39(1382): 167	Dwarf Fruit-eating Bat	None.	Venezuela, Bolívar, 59 km SE El Dorado, El Manaco.	S Colombia, Ecuador, Peru, Bolivia, Brazil, Venezuela, Guianas.	Not listed.	Least Concern	Subgenus Dermanura .  Distinct from cinereus and glaucus ; see Handley (1987), Brosset and Charles-Dominique(1990), and Simmons and Voss (1998).	Dermanura gnoma	1005008	23	Dwarf Fruit-eating Bat	Dwarf Little Fruit-eating Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	STENODERMATINAE	STENODERMATINI	Dermanura	NA	gnoma	Handley	1987	1	Artibeus_gnomus	Handley, C. O., Jr. (1987). New Species of Mammals from Northern South America: Fruit-Eating Bats, Genus Artibeus Leach. Fieldiana Zoology, new series 39, 167.	https://www.biodiversitylibrary.org/page/4465287#page/177/mode/1up	USNM 387534		"El Manaco (= Km 74), 59 km SE El Dorado, Bolívar, Venezuela, 150 m."			gnoma (Handley, 1987)	moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus	Hoofer, S. R., Solari, S., Larsen, P. A., Bradley, R. D., & Baker, R. J. (2008). Phylogenetics of the fruit-eating bats (Phyllostomidae: Artibeina) inferred from mitochondrial DNA sequences. Occasional Papers of the Museum of Texas Tech University, 277, 1-15.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.|York, H. A., Rodríguez-Herrera, B., Laval, R. K., Timm, R. M., & Lindsay, K. E. (2019). Field key to the bats of Costa Rica and Nicaragua. Journal of Mammalogy, 100(6), 1726-1749.|Turcios-Casco, M. A., Ávila-Palma, H. D., LaVal, R. К., Stevens, R. D., Ordoñez-Trejo, E. J., Soler-Orellana, J. A., & Ordoñez-Mazier, D. I. (2020). A systematic revision of the bats (Chiroptera) of Honduras: an updated checklist with corroboration of historical specimens and new records. Zoosystematics and Evolution, 96, 411.|				Colombia|Venezuela|Guyana|Suriname|French Guiana|Brazil|Ecuador|Peru|Bolivia	South America	Neotropic	LC	0	0	0	Artibeus_gnomus	0	oldname match	Artibeus_gnomus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Dermanura_gnomus	1005008	23	Dwarf Fruit-eating Bat	Dwarf Little Fruit-eating Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Stenodermatinae	Stenodermatini	Dermanura	NA	gnomus	Handley	1	Artibeus gnomus	Handley, C.O., Jr. 1987-12-31. New species of mammals from northern South America: fruit-eating bats, genus _Artibeus_ Leach. Fieldiana Zoology (n.s.)39:163-172.	https://www.biodiversitylibrary.org/page/4465291	USNM:MAMM:387534	holotype	http://n2t.net/ark:/65665/35427956f-d927-4624-b2cc-d671866c9aed	"El Manaco (= Km 74), 59 km SE El Dorado, Bolívar, Venezuela, 150 m."			moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus	Hoofer, S. R., Solari, S., Larsen, P. A., Bradley, R. D., & Baker, R. J. (2008). Phylogenetics of the fruit-eating bats (Phyllostomidae: Artibeina) inferred from mitochondrial DNA sequences. Occasional Papers of the Museum of Texas Tech University, 277, 1-15.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.|York, H. A., Rodríguez-Herrera, B., Laval, R. K., Timm, R. M., & Lindsay, K. E. (2019). Field key to the bats of Costa Rica and Nicaragua. Journal of Mammalogy, 100(6), 1726-1749.|Turcios-Casco, M. A., Ávila-Palma, H. D., LaVal, R. К., Stevens, R. D., Ordoñez-Trejo, E. J., Soler-Orellana, J. A., & Ordoñez-Mazier, D. I. (2020). A systematic revision of the bats (Chiroptera) of Honduras: an updated checklist with corroboration of historical specimens and new records. Zoosystematics and Evolution, 96, 411.|				Colombia|Venezuela|Guyana|Suriname|French Guiana|Brazil|Ecuador|Peru|Bolivia	South America	Neotropic	LC (as Dermanura gnoma)	0	0	0	Artibeus_gnomus	0	oldname match	Artibeus_gnomus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Dermanura		gnomus	Handley	1987	1	Fieldiana Zool. N.S.	39(1382): 167	Dwarf Fruit-eating Bat	None.	Venezuela, Bolívar, 59 km SE El Dorado, El Manaco.	S Colombia, Ecuador, Peru, Bolivia, Brazil, Venezuela, Guianas.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/2129/97207684/' target='_blank'>Least Concern</a>	Subgenus Dermanura. Distinct from cinereus and glaucus; see Handley (1987), Brosset and Charles-Dominique(1990), and Simmons and Voss (1998). The species epithet is a noun in apposition and therefore maintains the original spelling.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	Artibeina	Artibeus gnomus; Artibeus gnoma; Dermanura gnoma; Dermanura gnomus; gnoma; Dermanure naine; Zwergsilberfruchtvampir; Artibeoenano; Dwarf Little Fruit-eating Bat; Dwarf Fruit-eating Bat; Dwarf Little Fruit-eating Bat; Dwarf Fruit-eating Bat; Dwarf Fruit-eating Bat; Artibeus gnomus gnomus; Artibeus gnoma gnoma; Dermanura gnoma gnoma; Dermanura gnomus gnomus; D. gnoma