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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L211	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Artibeus anderseni	Artibeus anderseni	Artibeus anderseni	Artibeus anderseni	Artibeus anderseni	Artibeus anderseni	Artibeus anderseni	Dermanura anderseni	Dermanura anderseni	Dermanura anderseni	Dermanura anderseni	Dermanura anderseni	Dermanura anderseni	Dermanura anderseni	Dermanura anderseni		[MSW2] Subgenus Dermanura. Previously considered a subspecies of cinereus; but see Koopman (1978b:14).; [MSW3] Subgenus Dermanura. Previously considered a subspecies of cinereus, but see Koopman (1978b) and Handley (1987).; [HMW] Artibeus anderseni Osgood, 1916 , “Porto Velho [ Rondonia ], Brazil .” No other epithets have been associated with A. anderseni , but it was considered a junior synonym or subspecies of A. cinereus . Some authors placed anderseni in Dermanura , but it was reassigned to Artibeus (subgenus Dermanura ) after reinterpretation of molecular results. Monotypic.; [batnames2022] Subgenus Dermanura .  Previously considered a subspecies of cinereus , but see Koopman (1978 b ) and Handley(1987).; [MDD2022] moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus; [IUCN] Subgenus Dermanura (Simmons 2005). This subgenus has been recognized as a separate genus on molecular grounds, but there are no diagnostic morphological differences from Artibeus (Lim et al. 2004).  The previous (2008) assessment for this species was published under the genus Artibeus . This is an amended assessment, under the newly recognised genus Dermanura. ; [batnames2023] Subgenus Dermanura .  Previously considered a subspecies of cinereus , but see Koopman (1978 b ) and Handley(1987).; [MDD2023] moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus; [MDD2025_2.0] moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus; [batnames2025_1.7] Subgenus Dermanura. Previously considered a subspecies of cinereus, but see Koopman (1978b) and Handley(1987).; [MDD2025_2.2] moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus														anderseni	Subgenus Dermanura (Simmons 2005). This subgenus has been recognized as a separate genus on molecular grounds, but there are no diagnostic morphological differences from Artibeus (Lim et al. 2004).  The previous (2008) assessment for this species was published under the genus Artibeus . This is an amended assessment, under the newly recognised genus Dermanura. 			anderseni	anderseni			anderseni (Osgood, 1916)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		E Peru, W Brazil; ref. 4.6	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Artibeus anderseni	Brazil, Rondonia, Porto Velho.	Osgood	1916	Field Mus. Nat. Hist. Publ. Ser. Zool., 10:212.	Distribution: Largely confined to central South America east of the Andes from Ecuador to Bolivia and French Guiana, but also known from northern Colombia.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Bolivia – Peru, W Brazil, French Guiana	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Osgood	1916	Field Mus. Nat. Hist. Publ., Zool. Ser., 10:212.	Subgenus Dermanura. Previously considered a subspecies of cinereus; but see Koopman (1978b:14).	W Brazil, Bolivia, Ecuador, Peru.	Brazil, Rondonia, Porto Velho.		OSGOOD	1916	Size relatively small (forearm length, 34-37 mm; greatest length of skull, 16-19 mm). Preorbital and postorbital processes absent. Last upper and lower molars absent. Transition between braincase and rostrum abrupt. Upper molars relatively narrow. Uropatagium virtually hairless.	Distribution: Largely confined to central South America east of the Andes from Ecuador to Bolivia and French Guiana, but also known from northern Colombia.	No subspecies.		92	species	A. anderseni	OSGOOD	1916	Dermanura	subgenus	Artibeus anderseni				Size relatively small (forearm length, 34-37 mm; greatest length of skull, 16-19 mm). Preorbital and postorbital processes absent. Last upper and lower molars absent. Transition between braincase and rostrum abrupt. Upper molars relatively narrow. Uropatagium virtually hairless.	No subspecies.		15. A. anderseni OSGOOD 1916.	15	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Stenodermatinae	Stenodermatini	Artibeus anderseni	Artibeus	Dermanura	anderseni	Osgood		1916		Field Mus. Nat. Hist. Publ., Zool. Ser.	10		212		Andersen's Fruit-eating Bat	Brazil, Rondonia, Porto Velho.	W Brazil, Bolivia, Ecuador, Peru.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).		Subgenus Dermanura. Previously considered a subspecies of cinereus, but see Koopman (1978b) and Handley (1987).	03A687BCFFEFFFEF1393F29AF923F509	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	578	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFFEFFFEF1393F29AF923F509.xml	Artibeus anderseni	Phyllostomidae	Artibeus	anderseni	Osgood	1916	Dermanure d Andersen @fr | Andersen-Fruchtvampir @de | Artibeo de Andersen @es | Andersen's Little Fruit-eating Bat @en	Artibeus anderseni Osgood, 1916 , “Porto Velho [ Rondonia ], Brazil .” No other epithets have been associated with A. anderseni , but it was considered a junior synonym or subspecies of A. cinereus . Some authors placed anderseni in Dermanura , but it was reassigned to Artibeus (subgenus Dermanura ) after reinterpretation of molecular results. Monotypic.	W Amazon Basin in SE Colombia , NW & W Brazil , and E lowlands of Ecuador , Peru , and Bolivia , with an isolated population in W Colombia ( Antioquia Department ). Records from Venezuela , N Brazil , and Guyana probably represent misidentifications.	Head-body 46-55 mm (tailless), ear 14-18 mm, hindfoot 9-11 mm, forearm 34-38 mm; weight 8-9 g. Dorsal fur of Andersen’s Fruit-eating Bat is pale brown or grayish; fur is medium length, faintly tricolored, and soft; and it extends to base of forearm. Muzzle is short, with two well-marked white facial stripes. Ventralfur is slightly paler than dorsum. Ears are brown and usually have narrow but conspicuous whitish or pale yellow margins. Wing membranes are brownish. Tail membrane is pale, moderate in size, with a Vsshaped groove, and almost completely naked. Dental formula is 12/2, C 1/1,P 2/2, M 2/2 ( x2 ) = 28. Skull is small, with short and broad rostrum, and is characterized by high cranial dome similar to that of the Pygmy Fruiteating Bat (A. phaeotis ).	Usually lowland humid forests, typical of the Amazon Basin, but also cerrado and savanna habitats, forests fragments, and plantations (banana), mostly below elevations of ¢. 1000 m .	Andersen’s Fruit-eating Bat feeds primarily on fruits ( Ficus spp., Moraceae ), mainly from large trees.	Lactating and pregnant Andersen's Fruit-eating Bats have been found in June and October in Brazil and March, June, August, and November in Peru . These could indicate a bimodal polyestrous reproductive pattern.	Andersen’s Fruit-eating Bat is nocturnal. It roosts under banana-like leaves and perhaps in tents made from palm leaves.	Andersen’s Fruit-eating Bat roosts alone or forms small groups (1-3 individuals).	Classified as Least Concern on The IUCN Red List (as Dermanura andersent ). Andersen’s Fruit-eating Bat has a wide distribution.	Albuja (1999) | Ascorra et al. (1996) | Bernard & Fenton (2002) | Davis (1970a) | Graham (1987) | Handley (1987) | Hershkovitz (1949) | Koopman (1978) | Kunz et al. (1994) | Marques-Aguiar (2008a) | Osgood (1916) | Solari et al. (2006)	https://zenodo.org/record/6459003/files/figure.png	204. Andersen’s Fruit-eating Bat Artibeus anderseni French: Dermanure dAndersen / German: Andersen-Fruchtvampir / Spanish: Artibeo de Andersen Other common names: Andersen's Little Fruit-eating Bat Taxonomy. Artibeus anderseni Osgood, 1916 , “Porto Velho [ Rondonia ], Brazil .” No other epithets have been associated with A. anderseni , but it was considered a junior synonym or subspecies of A. cinereus . Some authors placed anderseni in Dermanura , but it was reassigned to Artibeus (subgenus Dermanura ) after reinterpretation of molecular results. Monotypic. Distribution. W Amazon Basin in SE Colombia , NW & W Brazil , and E lowlands of Ecuador , Peru , and Bolivia , with an isolated population in W Colombia ( Antioquia Department ). Records from Venezuela , N Brazil , and Guyana probably represent misidentifications. Descriptive notes. Head-body 46-55 mm (tailless), ear 14-18 mm, hindfoot 9-11 mm, forearm 34-38 mm; weight 8-9 g. Dorsal fur of Andersen’s Fruit-eating Bat is pale brown or grayish; fur is medium length, faintly tricolored, and soft; and it extends to base of forearm. Muzzle is short, with two well-marked white facial stripes. Ventralfur is slightly paler than dorsum. Ears are brown and usually have narrow but conspicuous whitish or pale yellow margins. Wing membranes are brownish. Tail membrane is pale, moderate in size, with a Vsshaped groove, and almost completely naked. Dental formula is 12/2, C 1/1,P 2/2, M 2/2 ( x2 ) = 28. Skull is small, with short and broad rostrum, and is characterized by high cranial dome similar to that of the Pygmy Fruiteating Bat (A. phaeotis ). Habitat. Usually lowland humid forests, typical of the Amazon Basin, but also cerrado and savanna habitats, forests fragments, and plantations (banana), mostly below elevations of ¢. 1000 m . Food and Feeding. Andersen’s Fruit-eating Bat feeds primarily on fruits ( Ficus spp., Moraceae ), mainly from large trees. Breeding. Lactating and pregnant Andersen's Fruit-eating Bats have been found in June and October in Brazil and March, June, August, and November in Peru . These could indicate a bimodal polyestrous reproductive pattern. Activity patterns. Andersen’s Fruit-eating Bat is nocturnal. It roosts under banana-like leaves and perhaps in tents made from palm leaves. Movements, Home range and Social organization. Andersen’s Fruit-eating Bat roosts alone or forms small groups (1-3 individuals). Status and Conservation. Classified as Least Concern on The IUCN Red List (as Dermanura andersent ). Andersen’s Fruit-eating Bat has a wide distribution. Bibliography. Albuja (1999), Ascorra et al. (1996), Bernard & Fenton (2002), Davis (1970a), Graham (1987), Handley (1987), Hershkovitz (1949), Koopman (1978), Kunz et al. (1994), Marques-Aguiar (2008a), Osgood (1916), Solari et al. (2006).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Dermanura anderseni	Dermanura		anderseni	Osgood	1916	1	Field Mus. Nat. Hist. Publ., Zool. Ser.	0.5639	Andersen's Fruit-eating Bat	None.	Brazil, Rondonia, Porto Velho.	W Brazil, Bolivia, Colombia, Ecuador, Peru.	Not listed.	Least Concern	Subgenus Dermanura .  Previously considered a subspecies of cinereus , but see Koopman (1978 b ) and Handley(1987).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Dermanura anderseni	23	Andersen's Fruit-eating Bat	Andersen's Little Fruit-eating Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	STENODERMATINAE	STENODERMATINI	Dermanura	NA	anderseni	Osgood	1916	1	Artibeus_anderseni	Osgood, W. H. (1916). Mammals of the Collins-Day South American expedition. Field Museum of Natural History Zoological Series, 10, 212.	https://www.biodiversitylibrary.org/item/20460#page/18/mode/1up	FMNH 21331		"Porto Velho [Rondônia], Brazil."			anderseni (Osgood, 1916)	moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus	Hoofer, S. R., Solari, S., Larsen, P. A., Bradley, R. D., & Baker, R. J. (2008). Phylogenetics of the fruit-eating bats (Phyllostomidae: Artibeina) inferred from mitochondrial DNA sequences. Occasional Papers of the Museum of Texas Tech University, 277, 1-15.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.|York, H. A., Rodríguez-Herrera, B., Laval, R. K., Timm, R. M., & Lindsay, K. E. (2019). Field key to the bats of Costa Rica and Nicaragua. Journal of Mammalogy, 100(6), 1726-1749.|Turcios-Casco, M. A., Ávila-Palma, H. D., LaVal, R. К., Stevens, R. D., Ordoñez-Trejo, E. J., Soler-Orellana, J. A., & Ordoñez-Mazier, D. I. (2020). A systematic revision of the bats (Chiroptera) of Honduras: an updated checklist with corroboration of historical specimens and new records. Zoosystematics and Evolution, 96, 411.|	Colombia|Brazil|Ecuador|Peru|Bolivia	South America	Neotropic	LC	0	0	0	Artibeus_anderseni	0	oldname match	Artibeus_anderseni	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	2122	Dermanura anderseni	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Dermanura	anderseni	(Osgood, 1916)	Subgenus Dermanura (Simmons 2005). This subgenus has been recognized as a separate genus on molecular grounds, but there are no diagnostic morphological differences from Artibeus (Lim et al. 2004).  The previous (2008) assessment for this species was published under the genus Artibeus . This is an amended assessment, under the newly recognised genus Dermanura. 	20000000	Dermanura anderseni	Least Concern		2016	2016-07-05 00:00:00 UTC	3.1	English	This species is listed as Least Concern as it is widespread, is presumed to tolerate a range of habitats, and is unlikely to be declining rapidly enough to qualify for inclusion in a threat category.	In central Amazonia, Brazil, Bernard and Fenton (2002) found this ;species in forest fragments, primary forests, and savannahs. ;Feeds on fruit, mainly from large trees. Roosts under banana-like leaves, and perhaps in tents made from palm leaves (Kunz ;et al . 1994). ;This species is poorly known, but lowland species in the genus are generally found in mature and disturbed lowland and premontane rainforest, plantations and gardens (Emmons and Feer 1997).	The species is affected by habitat loss although this is not a major threat.	This species is always represented by only a few records at each locality. It is a canopy frugivore and as such is probably undersampled in surveys due to methodology problems, it may well be more common than currently suggested.	Unknown	This bat is found from western Brazil, Bolivia, Ecuador, Peru, and Colombia (Simmons 2005, Gardner 2008). It has been found up to 1,300 m.		Terrestrial	Reduction of habitat loss is needed.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Dermanura		anderseni	Osgood	1916	1	Field Mus. Nat. Hist. Publ., Zool. Ser.	0.563889	Andersen's Fruit-eating Bat	None.	Brazil, Rondonia, Porto Velho.	W Brazil, Bolivia, Colombia, Ecuador, Peru.	Not listed.	Least Concern	Subgenus Dermanura .  Previously considered a subspecies of cinereus , but see Koopman (1978 b ) and Handley(1987).	Dermanura anderseni	1005000	23	Andersen's Fruit-eating Bat	Andersen's Little Fruit-eating Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	STENODERMATINAE	STENODERMATINI	Dermanura	NA	anderseni	Osgood	1916	1	Artibeus_anderseni	Osgood, W. H. (1916). Mammals of the Collins-Day South American expedition. Field Museum of Natural History Zoological Series, 10, 212.	https://www.biodiversitylibrary.org/item/20460#page/18/mode/1up	FMNH 21331		"Porto Velho [Rondônia], Brazil."			anderseni (Osgood, 1916)	moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus	Hoofer, S. R., Solari, S., Larsen, P. A., Bradley, R. D., & Baker, R. J. (2008). Phylogenetics of the fruit-eating bats (Phyllostomidae: Artibeina) inferred from mitochondrial DNA sequences. Occasional Papers of the Museum of Texas Tech University, 277, 1-15.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.|York, H. A., Rodríguez-Herrera, B., Laval, R. K., Timm, R. M., & Lindsay, K. E. (2019). Field key to the bats of Costa Rica and Nicaragua. Journal of Mammalogy, 100(6), 1726-1749.|Turcios-Casco, M. A., Ávila-Palma, H. D., LaVal, R. К., Stevens, R. D., Ordoñez-Trejo, E. J., Soler-Orellana, J. A., & Ordoñez-Mazier, D. I. (2020). A systematic revision of the bats (Chiroptera) of Honduras: an updated checklist with corroboration of historical specimens and new records. Zoosystematics and Evolution, 96, 411.|				Colombia|Brazil|Ecuador|Peru|Bolivia	South America	Neotropic	LC	0	0	0	Artibeus_anderseni	0	oldname match	Artibeus_anderseni	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Dermanura_anderseni	1005000	23	Andersen's Fruit-eating Bat	Andersen's Little Fruit-eating Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Stenodermatinae	Stenodermatini	Dermanura	NA	anderseni	Osgood	1	Artibeus anderseni	Osgood, W.H. 1916-10-31. Mammals of the Collins-Day South American expedition. Field Museum of Natural History, Zoological Series 10(14):199-216.	https://www.biodiversitylibrary.org/page/2651493	FMNH:Mamm:21331	holotype	http://portal.vertnet.org/o/fmnh/mammals?id=5c8cbf89-5472-441c-a227-0a762a9e0865	"Porto Velho [Rondônia], Brazil."			moved to Dermanura from Artibeus, although some recent publications do not agree with this split, there seems to be more publications using this revised taxonomy rather than leaving all species under Artibeus	Hoofer, S. R., Solari, S., Larsen, P. A., Bradley, R. D., & Baker, R. J. (2008). Phylogenetics of the fruit-eating bats (Phyllostomidae: Artibeina) inferred from mitochondrial DNA sequences. Occasional Papers of the Museum of Texas Tech University, 277, 1-15.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.|York, H. A., Rodríguez-Herrera, B., Laval, R. K., Timm, R. M., & Lindsay, K. E. (2019). Field key to the bats of Costa Rica and Nicaragua. Journal of Mammalogy, 100(6), 1726-1749.|Turcios-Casco, M. A., Ávila-Palma, H. D., LaVal, R. К., Stevens, R. D., Ordoñez-Trejo, E. J., Soler-Orellana, J. A., & Ordoñez-Mazier, D. I. (2020). A systematic revision of the bats (Chiroptera) of Honduras: an updated checklist with corroboration of historical specimens and new records. Zoosystematics and Evolution, 96, 411.|				Colombia|Brazil|Ecuador|Peru|Bolivia	South America	Neotropic	LC	0	0	0	Artibeus_anderseni	0	oldname match	Artibeus_anderseni	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Dermanura		anderseni	Osgood	1916	1	Field Mus. Nat. Hist. Publ., Zool. Ser.	0.563889	Andersen's Fruit-eating Bat	None.	Brazil, Rondonia, Porto Velho.	W Brazil, Bolivia, Colombia, Ecuador, Peru.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href=”https://www.iucnredlist.org/species/2122/22000743/” target=”_blank>Least Concern</a>	Subgenus Dermanura. Previously considered a subspecies of cinereus, but see Koopman (1978b) and Handley(1987).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	Artibeina	Artibeus anderseni; Artibeus andersenii; Dermanura anderseni; Dermanura anderseni; Dermanura anderseni; Dermanura anderseni; anderseni; Dermanure d Andersen; Andersen-Fruchtvampir; Artibeo de Andersen; Andersen's Little Fruit-eating Bat; Andersen's Fruit-eating Bat; Andersen's Little Fruit-eating Bat; Andersen's Fruit-eating Bat; Andersen's Fruit-eating Bat; Artibeus anderseni anderseni; D. anderseni