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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L210	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Cyttarops alecto	Cyttarops alecto	Cyttarops alecto	Cyttarops alecto	Cyttarops alecto	Cyttarops alecto	Cyttarops alecto	Cyttarops alecto	Cyttarops alecto	Cyttarops alecto	Cyttarops alecto	Cyttarops alecto	Cyttarops alecto	Cyttarops alecto	Cyttarops alecto		[MSW2] See Starrett (1972, Mammalian Species, 13).; [MSW3] Reviewed by Jones and Hood (1993); also see Starrett (1972). See Emmons (1997) for distribution map.; [HMW] Cyttarops alecto Thomas, 1913 , “ Mucajatuba, near Para ,” Brazil . This species is monotypic.; [batnames2022] Reviewed by Jones and Hood (1993); also see Starrett (1972). See Emmons (1997) for distribution map.; [IUCN] The genus is monotypic.; [batnames2023] Reviewed by Jones and Hood (1993); also see Starrett (1972). See Emmons (1997) for distribution map.; [batnames2025_1.7] Reviewed by Jones and Hood (1993); also see Starrett (1972). See Emmons (1997) for distribution map.														alecto	The genus is monotypic.			alecto	alecto			alecto O. Thomas, 1913		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Costa Rica, Nicaragua, Guyana, Brazil	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Cyttarops alecto	Brazil, Para, Mocajatuba.	Thomas	1913	Ann. Mag. Nat. Hist., ser. 8, 11:135.	Distribution: Same as for genus.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Short-eared bat	Costa Rica, Nicaragua, Guyana, Brazil	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1913	Ann. Mag. Nat. Hist., ser. 8, 11:135.	See Starrett (1972, Mammalian Species, 13).	Nicaragua, Costa Rica, Guyana, Amazonian Brazil.	Brazil, Para, Mocajatuba.		THOMAS	1913	Size relatively small (forearm length, 46-4 8 mm).	Distribution: Same as for genus.	No subspecies.		48	species	C. alecto	THOMAS	1913	Cyttarops	genus	Cyttarops alecto				Size relatively small (forearm length, 46-4 8 mm).	No subspecies.		1. C. alecto THOMAS 1913.	1	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Emballonuridae	Emballonurinae		Cyttarops alecto	Cyttarops		alecto	Thomas		1913		Ann. Mag. Nat. Hist., ser. 8	11		135		Short-eared Bat	Brazil, Pará, Mocajatube.	Nicaragua, Costa Rica, Guyana, French Guiana, Amazonian Brazil.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).		Reviewed by Jones and Hood (1993); also see Starrett (1972). See Emmons (1997) for distribution map.	03D587F2FFDD4C16FF3B3753FCBEF35B	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Emballorunidae.pdf.imd	hash://md5/ffecff8affcf4c04ffa53577fff8ffe9	367	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/D5/87/03D587F2FFDD4C16FF3B3753FCBEF35B.xml	Cyttarops alecto	Emballonuridae	Cyttarops	alecto	Thomas	1913	Short-eared Bat @en | Cytarope furieux @fr | Dunkelgraue Kurzohrfledermaus @de | Embalonuro orejicorto @es	Cyttarops alecto Thomas, 1913 , “ Mucajatuba, near Para ,” Brazil . This species is monotypic.	Central America, locally in Caribbean lowlands from Nicaragua to Panama, and South America in Venezuela (Bolivar State), Guyana, French Guiana, and N Brazil (Para State), also in scattered localities in Amazon Basin (extreme S o ­ lombia, E Peru, W Brazil, and N Bolivia). It might have a much larger distribution in the Amazon drainage of all these countries.	Head-body 47-55 mm, tail 20-25 mm, ear 11-13 mm, hindfoot 10—12 mm, forearm 45-47 mm; weight 6-7 g. Dorsal fur of the Short-eared Bat is long, fluffy, and entirely dark gray; venter is slighdy paler. Ears are broad and rounded, barely rising about crown. Chin is well haired and appears bearded. Flight membranes are black. Hindfeet are small. Thumbs are tiny . Wing sacs are absent. Nostrils are tubelike and slightly divergent. Dental formula is 11/3, C 1/1, P 2/2, M 3/3 (x2) = 32.	Humid lowland forests and along riparian gallery forests from sea level to elevations of500 m. In Brazil, the Short-eared Bat can occur along gallery forests lining small rivers in the savanna-like Cerrado ecoregion. It tolerates second growth disturbed by humans and forages near buildings and in oil palm plantations and gardens.	The Short-eared Bat preys on aerial insects.	A pregnant Short-eared Bat was reported in dry season in Tocantins State, Brazil. In Costa Rica, females with nursing young and subadults are reported in June- August.	Short-eared Bats are nocturnal and roost beneath fronds in palm trees including coconut and oil palms, usually in relatively open areas such as groves or gardens. Roost emergence starts c.45 minutes after sunset. Foraging usually is restricted to immediately around day roosts for 15—30 minutes, after which individuals disperse further from roosts while flying at least 3-4 m aboveground. Echolocation call is unusual among emballonurids: FM call remains above 100 kHz and is short in duration.	Short-eared Bats roost in groups of 1—10 individuals. They hang freely by their feet when roosting, near midribs of fronds. Roosting groups can contain adult males and females, subadults, and dependent young. There is evidence that roosting group composition is not stable even over short periods of time, with different individuals moving into a group.	Classified as Least Concern on The IUCN Red List. The Shorteared Bat has a large distribution and presumably relatively stable overall population. Its rarity in museum collections, known from fewer than 30 global specimens and c.20 localities, is likely due to inefficiency of capture in mist nets. Additional studies are needed on its distribution, habitat, ecology , and threats. Acoustic surveys should produce valuable information to better understand biology and conservation status of Short-eared Bats.	Aguirre , Mammani et al. (2010) | Fonseca et al. (1996) | Koopman (1993) | Lim, B.K. (2009 b) | Lim, B.K. & Engstrom (2005) | Lim, B.K. et al. (1999) | Nunes et al. (2006) | Reid (2009) | Reid & Langtimm (1993) | Rodriguez-Mahecha et al. (1995) | Simmons &Voss (1998) | Starrett (1972) | Starrett & Casebeer (1968) | Tavares, Bobrowiec & Farias (2012) | Velazco et al. (2011)	https://zenodo.org/record/3747970/files/figure.png	37 . Short-eared Bat Cyttarops alecto French: Cytarope furieux / German: Dunkelgraue Kurzohrfledermaus / Spanish: Embalonuro orejicorto Taxonomy . Cyttarops alecto Thomas, 1913 , “ Mucajatuba, near Para ,” Brazil . This species is monotypic. Distribution. Central America, locally in Caribbean lowlands from Nicaragua to Panama, and South America in Venezuela (Bolivar State), Guyana, French Guiana, and N Brazil (Para State), also in scattered localities in Amazon Basin (extreme S o ­ lombia, E Peru, W Brazil, and N Bolivia). It might have a much larger distribution in the Amazon drainage of all these countries. Descriptive notes. Head-body 47-55 mm, tail 20-25 mm, ear 11-13 mm, hindfoot 10—12 mm, forearm 45-47 mm; weight 6-7 g. Dorsal fur of the Short-eared Bat is long, fluffy, and entirely dark gray; venter is slighdy paler. Ears are broad and rounded, barely rising about crown. Chin is well haired and appears bearded. Flight membranes are black. Hindfeet are small. Thumbs are tiny . Wing sacs are absent. Nostrils are tubelike and slightly divergent. Dental formula is 11/3, C 1/1, P 2/2, M 3/3 (x2) = 32. Habitat . Humid lowland forests and along riparian gallery forests from sea level to elevations of500 m. In Brazil, the Short-eared Bat can occur along gallery forests lining small rivers in the savanna-like Cerrado ecoregion. It tolerates second growth disturbed by humans and forages near buildings and in oil palm plantations and gardens. Food and Feeding . The Short-eared Bat preys on aerial insects. Breeding . A pregnant Short-eared Bat was reported in dry season in Tocantins State, Brazil. In Costa Rica, females with nursing young and subadults are reported in June- August. Activity patterns. Short-eared Bats are nocturnal and roost beneath fronds in palm trees including coconut and oil palms, usually in relatively open areas such as groves or gardens. Roost emergence starts c.45 minutes after sunset. Foraging usually is restricted to immediately around day roosts for 15—30 minutes, after which individuals disperse further from roosts while flying at least 3-4 m aboveground. Echolocation call is unusual among emballonurids: FM call remains above 100 kHz and is short in duration. Movements, Home range and Social organization. Short-eared Bats roost in groups of 1—10 individuals. They hang freely by their feet when roosting, near midribs of fronds. Roosting groups can contain adult males and females, subadults, and dependent young. There is evidence that roosting group composition is not stable even over short periods of time, with different individuals moving into a group. Status and Conservation . Classified as Least Concern on The IUCN Red List. The Shorteared Bat has a large distribution and presumably relatively stable overall population. Its rarity in museum collections, known from fewer than 30 global specimens and c.20 localities, is likely due to inefficiency of capture in mist nets. Additional studies are needed on its distribution, habitat, ecology , and threats. Acoustic surveys should produce valuable information to better understand biology and conservation status of Short-eared Bats. Bibliography. Aguirre , Mammani et al. (2010), Fonseca et al. (1996), Koopman (1993), Lim, B.K. (2009 b ), Lim, B.K. & Engstrom (2005), Lim, B.K. et al. (1999), Nunes et al. (2006), Reid (2009), Reid & Langtimm (1993), Rodriguez-Mahecha et al. (1995), Simmons &Voss (1998), Starrett (1972), Starrett & Casebeer (1968), Tavares, Bobrowiec & Farias (2012), Velazco et al. (2011).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Emballonuridae	Cyttarops alecto	Cyttarops		alecto	Thomas	1913	0	Ann. Mag. Nat. Hist.	ser. 8, 11: 135	Short-eared Bat	None.	Brazil, Par&aacute;, Mocajatube.	Nicaragua, Costa Rica, Guyana, French Guiana, Amazonian Brazil, Bolivia.	Not listed.	Least Concern	Reviewed by Jones and Hood (1993); also see Starrett (1972). See Emmons (1997) for distribution map.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Cyttarops alecto	23	Short-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	EMBALLONURIDAE	EMBALLONURINAE	DICLIDURINI	Cyttarops	NA	alecto	O. Thomas	1913	0	Cyttarops_alecto	Thomas, O. (1913). On some rare Amazonian mammals from the collection of the Para Museum. Annals and Magazine of Natural History, ser.8, 11, 135.	https://www.biodiversitylibrary.org/item/61788#page/147/mode/1up	BM 1912.11.4.5		"Mucajatuba, near Para," Brazil.			alecto O. Thomas, 1913	NA	NA	Nicaragua|Costa Rica|Panama|Venezuela|Guyana|Suriname?|French Guiana|Brazil|Colombia|Peru|Bolivia	North America|South America	Neotropic	LC	0	0	0	Cyttarops_alecto	0	sciname match	Cyttarops_alecto	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	6206	Cyttarops alecto	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	EMBALLONURIDAE	Cyttarops	alecto	Thomas, 1913	The genus is monotypic.	20000000	Cyttarops alecto	Least Concern		2016	2016-07-01 00:00:00 UTC	3.1	English	This species is listed as Least Concern because, although it is widely distributed, and is probably not as rare as indicated by museum specimens due to difficulties in capture (mist nets) and is unlikely to be declining fast enough to qualify for inclusion in the threat categories.	Roosts in groups of 1 to 10 at the top of palm trees on fronds, usually in relatively open areas such as groves or gardens. It hangs freely by the feet when roosting, near the midrib of a frond. Activity starts about 45 min after sunset but is usually restricted to immediately around the roost for 15 to 30 min, after which time, in complete darkeness, individuals disperse, flying at least 3 to 4 m above ground (Starrett 1972, Reid and Langtimm 1993). Adaptable to human disturbed areas. Aerial insectivore.	Threats for this species are unknown.	This is one of the rarest Neotropical bats, known from fewer than 30 individuals taken from less than 20 localities in humid lowland areas (Reid 2009, Velazco et al . 2011, Tavares et al . 2012).	Unknown	This species is known from a few localities in the Caribbean lowlands of Nicaragua (Koopman 1993), Costa Rica (Starrett and Casebeer 1968, Reid 2009), ;Panama (Reid 2009), ;Colombia (Rodriguez et al.  ;1995), Guyana (Lim and Engstrom 2005), Suriname (Lim 2009), French Guiana (Simmons and Voss 1998), Amazonian Brazil (Fonseca et al . 1996), Peru (Velazco et al . 2011) and Bolivia (Aguirre et al . 2010). All localities are at or below 500 m elevation (Tavares et al . 2012).		Terrestrial	Further studies are needed into the distribution, habitat, ecology, and threats to this species.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Emballonuridae	Cyttarops		alecto	Thomas	1913	0	Ann. Mag. Nat. Hist.	ser. 8, 11: 135	Short-eared Bat	None.	Brazil, Par&aacute;, Mocajatube.	Nicaragua, Costa Rica, Guyana, French Guiana, Amazonian Brazil, Bolivia.	Not listed.	Least Concern	Reviewed by Jones and Hood (1993); also see Starrett (1972). See Emmons (1997) for distribution map.	Cyttarops alecto	1004782	23	Short-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	EMBALLONUROIDEA	Emballonuridae	EMBALLONURINAE	DICLIDURINI	Cyttarops	NA	alecto	O. Thomas	1913	0	Cyttarops_alecto	Thomas, O. (1913). On some rare Amazonian mammals from the collection of the Para Museum. Annals and Magazine of Natural History, ser.8, 11, 135.	https://www.biodiversitylibrary.org/item/61788#page/147/mode/1up	BM 1912.11.4.5		"Mucajatuba, near Para," Brazil.			alecto O. Thomas, 1913	NA	NA				Nicaragua|Costa Rica|Panama|Venezuela|Guyana|Suriname?|French Guiana|Brazil|Colombia|Peru|Bolivia	North America|South America	Neotropic	LC	0	0	0	Cyttarops_alecto	0	sciname match	Cyttarops_alecto	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Cyttarops_alecto	1004782	23	Short-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Emballonuroidea	Emballonuridae	Emballonurinae	Diclidurini	Cyttarops	NA	alecto	O. Thomas	0	Cyttarops alecto	Thomas, O. 1913-01-01. On some rare Amazonian mammals from the collection of the Para Museum. Annals and Magazine of Natural History (8)11(61):130-136.	https://www.biodiversitylibrary.org/page/18635696	BMNH:Mamm:1912.11.4.5	holotype	https://data.nhm.ac.uk/object/8a11975b-37b9-4c2b-991e-2af7cc4239d8	"Mucajatuba, near Para," Brazil.			NA	NA				Nicaragua|Costa Rica|Panama|Venezuela|Guyana|Suriname?|French Guiana|Brazil|Colombia|Peru|Bolivia	North America|South America	Neotropic	LC	0	0	0	Cyttarops_alecto	0	sciname match	Cyttarops_alecto	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Emballonuridae	Cyttarops		alecto	Thomas	1913	0	Ann. Mag. Nat. Hist.	ser. 8, 11: 135	Short-eared Bat	None.	Brazil, Par&aacute;, Mocajatube.	Nicaragua, Costa Rica, Guyana, French Guiana, Amazonian Brazil, Bolivia.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/6206/22022820/' target='_blank'>Least Concern</a>	Reviewed by Jones and Hood (1993); also see Starrett (1972). See Emmons (1997) for distribution map.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Cyttarops alecto; Cyttarops alecto; Cyttarops alecto; Cyttarops alecto; Cyttarops alecto; Cyttarops alecto; alecto; Short-eared Bat; Cytarope furieux; Dunkelgraue Kurzohrfledermaus; Embalonuro orejicorto; Short-eared Bat; Short-eared Bat; Short-eared Bat; C. alecto