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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L203	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Cynopterus horsfieldi	Cynopterus horsfieldi	Cynopterus horsfieldii	Cynopterus horsfieldi	Cynopterus horsfieldi	Cynopterus horsfieldii	Cynopterus horsfieldii	Cynopterus horsfieldii	Cynopterus horsfieldii	Cynopterus horsfieldii	Cynopterus horsfieldii	Cynopterus horsfieldii	Cynopterus horsfieldii	Cynopterus horsfieldii	Cynopterus horsfieldii		[MSW2] Includes harpax (see Hill, 1961fl).; [MSW3] Includes harpax; see Hill (1961a). This name is sometimes spelled horsefieldi.; [HMW] Cynopterus horsfieldii J. E. Gray, 1843 , Java , Indonesia . Four subspecies are recognized.; [batnames2022] Includes harpax ; see Hill (1961 a ). While this name is sometimes spelled horsefieldi , the correct original spelling is <horsfieldii following ICZN artcile 33.4.; [batnames2023] Includes harpax ; see Hill (1961 a ). While this name is sometimes spelled horsefieldi , the correct original spelling is <horsfieldii following ICZN artcile 33.4.; [batnames2025_1.7] Includes harpax; see Hill (1961). See also Campbell and Kunz (2006). While this name is sometimes spelled horsefieldi, the correct original spelling is horsfieldii following ICZN article 33.4.				harpax		harpax, lyoni; minor Lyon; persimilis, princeps.	horsfieldi, persimilis, harpax, princeps	horsfieldii, harpax, persimilis, princeps	harpax - lyoni, minor	horsfieldii, harpax, persimilis, princeps		horsfieldii, harpax, lyoni, persimilis, princeps	lyoni - minor	horsfieldii, princeps, harpax, minor, lyoni, persimilis		horsfieldii, harpax, persimilis, princeps	harpax - lyoni, minor	horsfieldii, princeps, harpax, minor, lyoni, persimilis	horsfieldii, princeps, minor, harpax, lyoni, persimilis, horsfieldi	harpax, horsfieldii, persimilis, princeps	harpax - lyoni, minor 	horsfieldii J. E. Gray, 1843|princeps G. S. Miller, 1906|minor (Lyon, 1908) [preoccupied]|harpax O. Thomas & Wroughton, 1909|lyoni Andersen, 1912 [nomen novum]|persimilis Andersen, 1912|horsfieldi D. D. Davis, 1962 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Horsfield's fruit bat	S Thailand – Java, Borneo	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Cynopterus horsfieldi	Indonesia, Java	Gray	1843	List Mamm. Br. Mus., p.38.	Distribution: From Thai land to Java and Borneo.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Horsfield's fruit bat	S Thailand – Java, Borneo	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Gray	1843	List Specimens Mamm. Coll. Brit. Mus., p. 38.	Includes harpax (see Hill, 1961fl).	Thailand, W Malaysia, Borneo, Java, Sumatra, Lesser Sunda Isis, and adjacent small islands.	Indonesia, Java.		GRAY	1843	Molars and pre molars relatively broad and subrectangular in outline. Surface cusp on last lower premolar and first lower molar always well developed. Size me dium to large (forearm length, 64-90 mm; ear length, 17-18 mm).	Distribution: From Thai land to Java and Borneo.	Four currently rec ognized subspecies:	C. h. horsfieldi (Java), C. h. persimilis (Borneo), C. h. harpax (= minor, lyoni) (Sumatra, Malaya, Thailand), C. h. princeps (Nias island off western Sumatra).	33	species	C. horsfieldi	GRAY	1843	Cynopterus	genus	Cynopterus horsfieldi				Molars and pre molars relatively broad and subrectangular in outline. Surface cusp on last lower premolar and first lower molar always well developed. Size me dium to large (forearm length, 64-90 mm; ear length, 17-18 mm).	Four currently rec ognized subspecies:		5. C. horsfieldi GRAY 1843.	5	_C. h. harpax_ Thomas & Wroughton, 1909 (synonyms: _lyoni_ Andersen, 1912, _minor_ (Lyon, 1908)); _C. h. horsfieldii_ Gray, 1843; _C. h. persimilis_ Andersen, 1912; _C. h. princeps_ Miller, 1906			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Pteropodidae			Cynopterus horsfieldii	Cynopterus		horsfieldii	Gray		1843		List Specimens Mamm. Coll. Brit. Mus.			38		Horsfield’s Fruit Bat	Indonesia, Java.	Thailand, Cambodia, W Malaysia, Borneo, Java, Sumatra, Lesser Sunda Isls, and adjacent small islands.	IUCN/SSC Action Plan (1992) – Not Threatened. IUCN 2003 – Lower Risk (lc).	harpax Thomas and Wroughton, 1909; lyoni K. Andersen, 1912; minor Lyon, 1908 [not Trouessart, 1878]; persimilis K. Andersen, 1912; princeps Miller, 1906.	Includes harpax; see Hill (1961a). This name is sometimes spelled horsefieldi.	03AD87FAFFCCF62289B03D43F684F552	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Pteropodidae_16.pdf.imf	hash://md5/ff94ff82ffc4f62a891e341cffa5ff9b	66	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/AD/87/03AD87FAFFCCF62289B03D43F684F552.xml	Cynopterus horsfieldii	Pteropodidae	Cynopterus	horsfieldii	J. E. Gray	1843	Cynoptére de Horsfield @fr | Horsfield-Kurznasenflughund @de | Cynéptero de Horsfield @es | Codot Horsfield @en | Horsfield's Fruit Bat @en	Cynopterus horsfieldii J. E. Gray, 1843 , Java , Indonesia . Four subspecies are recognized.	C.h.horsfieldiiJ.E.Gray,1843~Java,Bali,Lombok,andFloresIs;itmaybealsopresentonSumbawaI. C.h.harpaxThomas&Wroughton,1909—scatteredpopulationsinW&CThailand,MalayPeninsula,includingTiomanI,andSumatra. C.h.persimilisK.Andersen,1912—Borneo. C. h. princeps G. S. Miller, 1906 — Nias I. Population on Simelue I may also represent this subspecies.	Head—body 85-120 mm,tail 12-19 mm, ear 19-22 mm, hindfoot 14-16 mm, forearm 74-3 mm (64-5— 71 mm in nominate horsfieldii , 87-89-5 mm in princeps); weight 43-63 g. Horsfield’s Short-nosed Fruit Bat is medium-sized, with rimmed ears and surface cusps on lower cheekteeth. Muzzle is short and deep; nostrils are shortly tubular and divergent, and their rims are thickened; and muzzle skin is brown. Eyes are large; iris is warm brown or cocoa-brown. Ears are long and oval, with slightly attenuated tips and white rims. Head is wide, and pelage is brown or rust-brown, longer on nape and lighter on dorsum. Tail is longer than broad uropatagium rim, and calcar is short. Pelage on throat is sparse. Conspicuous reddish brown to orange ruff extends to sides of neck and chest;it is stiffer, brighter, and richer on males. Lower chest and belly are light gray-brown; flanks have yellowish tinge. Genitals are dark brown. Wing membranes are dark grayish brown, originate on sides of body, and attach to first toe; index claw is present; and metacarpals and phalanges are dorsally white. Skull lacks basicranial deflection. Rostrum is short, forehead gently slopes, orbitis large, braincase is slightly flattened anteriorly and more rounded posteriorly, zygomatic root is above alveolar line, and zygoma is much arched and wide open. Dorsally, rostrum is wide and short; paranasal recesses are inflated and surpass posteriorly supraorbital foramina; postorbital processes are relatively thick and pointed posterolaterally; postorbital constriction is small; sagittal crest is low; braincase is oval; and nuchal crest present. Palate is flat, post-dental palate is long and convergent; palatine spine is joined to midsphenoidal ridge. Ectotympanic is small and wide anteriorly; entotympanic long and thin. Mandible has moderately thick body; coronoid is tall and rather steep, with wide tip; condyle is above lower alveolar line; and angle is round off, with demarked rim. Upper incisors are small and long; C' is long and strongly decurved, with marked cingula and strong secondary cusp on inner edge; P' is minute; and posterior cheekteeth are tall and strong, with marked cusps, from almost square to rectangular outline posteriorly. Lower incisors are small; C,is strong but short; P| is well formed, with cusp; next premolar (P,) is large and astall as C,, following teeth decreasing in height; P, and M have conspicuous additional surface cusp (especially in princeps); and M, is small, with normal cusps distinguishable. Chromosomal complement has 2n = 34 and FN = 58, with eleven metacentric or submetacentric, two subacrocentric, and three acrocentric autosomes. X-chromosome is subacrocentric, and Y-chromosome is small acrocentric.	Lowland to montane forests, mangrove forests, and transition from forest to open/disturbed forest or cultivated land from sea level to elevations of ¢. 1460 m . Horsfield’s Short-nosed Fruit Bat uses understory and subcanopy strata and prefers to roost in forest edges and secondary forests.	Horsfield’s Short-nosed Fruit Bat is mainly frugivorous. It is transient, with aseasonal changes in abundance, and uses mainly large-crop, big-bang fruiting plants. Single fruit are taken from a tree and consumed in nearby day or feeding roost where seeds and dry pellets are discarded. Feeding roosts are located in foliage c. 3 m aboveground. Plants consumed include mainly Ficus (FE fistulosa and FE variegata ) and Artocarpus (both Moraceae ); Elaeocarpus (Elacocarpaceae); Payena ( Sapotaceae ); Plernandra ( Melastomataceae ); and introduced Piper aduncum ( Piperaceae ). Fruits from gardens and orchards are often eaten. Flower products are important during dry season when fruits are scarce, with up to eleven pollen types eaten (e.g. Parkia ; Fabaceae ) and unidentified leaves.	Mating system of the Horsfield’s Short-nosed Fruit Bat is polygynous. Females are polyestrous, with postpartum estrus. Reproduction is largely aseasonal, with pregnant females found throughout the year. Litter size is one, with up to two litters per year.	Activity pattern is bimodal. Horsfield’s Short-nosed Fruit Bats are most active 2-4 hours after sunset and again c.3 hours before sunrise, with resting period around midnight.	Horsfield’s Short-nosed Fruit Bats mainly move between day roosts and feeding areas,traveling 400-1600 m. Mean home range is 8 ha for adult males and 6 ha for adult females. Home ranges widely overlap among males and females and with individuals of other species of Cynopterus (e.g. Lesser Short-nosed Fruit Bat, C. brachyotis ). Social organization is harem-based, with a single male associated with 2-5 reproductive females and their young. Harem group roosts in a tent built by the male or in unmodified roosts. In Peninsular Malaysia , banana ( Musa , Musaceae ) leaves are preferred roosts; ¢.50% of them is modified as tents by chewing mid-rib midway between base and tip. Other roosts include withering roots of epiphytic ferns ( Asplenium , Aspleniaceae ) and palm fronds ( Arenga , Cocos , Corypha , and Nypa ; Arecaceae ). Roosts are spatially clumped in forests. Males and females have low roost fidelity, and harem system is labile. Males change roosts every 1-7 days, and females change roosts every 3-14 days. Alternative roost sites include cave entrances and limestone solution cavities, where Horsfield’s Short-nosed Fruit Bats are more gregarious.	Classified as Least Concern on The IUCN Red List. Horsfield’s Short-nosed Fruit Bat is abundant and uses modified forests as roosting and feeding habitat. Population trend is unknown.	Andersen (1912b) | Bates, Francis, Gumal & Bumrungsri (2008) | Campbell (2008) | Campbell & Kunz (2006) | Campbell, Reid et al. (2006) | Campbell, Schneider et al. (2006) | Fletcher et al. (2012) | Francis (1994) | Funakoshi & Zubaid (1997) | Hodgkison (2001) | Jones, Bielby et al. (2009) | Kingston et al. (2006) | Kitchener, Gunnell & Maharadatunkamsi (1990) | Tingga et al. (2012) | Yong et al. (1973)		7. Horsfield’s Short-nosed Fruit Bat Cynopterus horsfieldii French: Cynoptére de Horsfield / German: Horsfield-Kurznasenflughund / Spanish: Cynéptero de Horsfield Other common names: Codot Horsfield , Horsfield's Fruit Bat Taxonomy. Cynopterus horsfieldii J. E. Gray, 1843 , Java , Indonesia . Four subspecies are recognized. Subspecies and Distribution. C.h.horsfieldiiJ.E.Gray,1843~Java,Bali,Lombok,andFloresIs;itmaybealsopresentonSumbawaI. C.h.harpaxThomas&Wroughton,1909—scatteredpopulationsinW&CThailand,MalayPeninsula,includingTiomanI,andSumatra. C.h.persimilisK.Andersen,1912—Borneo. C. h. princeps G. S. Miller, 1906 — Nias I. Population on Simelue I may also represent this subspecies. Descriptive notes. Head—body 85-120 mm,tail 12-19 mm, ear 19-22 mm, hindfoot 14-16 mm, forearm 74-3 mm (64-5— 71 mm in nominate horsfieldii , 87-89-5 mm in princeps); weight 43-63 g. Horsfield’s Short-nosed Fruit Bat is medium-sized, with rimmed ears and surface cusps on lower cheekteeth. Muzzle is short and deep; nostrils are shortly tubular and divergent, and their rims are thickened; and muzzle skin is brown. Eyes are large; iris is warm brown or cocoa-brown. Ears are long and oval, with slightly attenuated tips and white rims. Head is wide, and pelage is brown or rust-brown, longer on nape and lighter on dorsum. Tail is longer than broad uropatagium rim, and calcar is short. Pelage on throat is sparse. Conspicuous reddish brown to orange ruff extends to sides of neck and chest;it is stiffer, brighter, and richer on males. Lower chest and belly are light gray-brown; flanks have yellowish tinge. Genitals are dark brown. Wing membranes are dark grayish brown, originate on sides of body, and attach to first toe; index claw is present; and metacarpals and phalanges are dorsally white. Skull lacks basicranial deflection. Rostrum is short, forehead gently slopes, orbitis large, braincase is slightly flattened anteriorly and more rounded posteriorly, zygomatic root is above alveolar line, and zygoma is much arched and wide open. Dorsally, rostrum is wide and short; paranasal recesses are inflated and surpass posteriorly supraorbital foramina; postorbital processes are relatively thick and pointed posterolaterally; postorbital constriction is small; sagittal crest is low; braincase is oval; and nuchal crest present. Palate is flat, post-dental palate is long and convergent; palatine spine is joined to midsphenoidal ridge. Ectotympanic is small and wide anteriorly; entotympanic long and thin. Mandible has moderately thick body; coronoid is tall and rather steep, with wide tip; condyle is above lower alveolar line; and angle is round off, with demarked rim. Upper incisors are small and long; C' is long and strongly decurved, with marked cingula and strong secondary cusp on inner edge; P' is minute; and posterior cheekteeth are tall and strong, with marked cusps, from almost square to rectangular outline posteriorly. Lower incisors are small; C,is strong but short; P| is well formed, with cusp; next premolar (P,) is large and astall as C,, following teeth decreasing in height; P, and M have conspicuous additional surface cusp (especially in princeps); and M, is small, with normal cusps distinguishable. Chromosomal complement has 2n = 34 and FN = 58, with eleven metacentric or submetacentric, two subacrocentric, and three acrocentric autosomes. X-chromosome is subacrocentric, and Y-chromosome is small acrocentric. Habitat. Lowland to montane forests, mangrove forests, and transition from forest to open/disturbed forest or cultivated land from sea level to elevations of ¢. 1460 m . Horsfield’s Short-nosed Fruit Bat uses understory and subcanopy strata and prefers to roost in forest edges and secondary forests. Food and Feeding. Horsfield’s Short-nosed Fruit Bat is mainly frugivorous. It is transient, with aseasonal changes in abundance, and uses mainly large-crop, big-bang fruiting plants. Single fruit are taken from a tree and consumed in nearby day or feeding roost where seeds and dry pellets are discarded. Feeding roosts are located in foliage c. 3 m aboveground. Plants consumed include mainly Ficus (FE fistulosa and FE variegata ) and Artocarpus (both Moraceae ); Elaeocarpus (Elacocarpaceae); Payena ( Sapotaceae ); Plernandra ( Melastomataceae ); and introduced Piper aduncum ( Piperaceae ). Fruits from gardens and orchards are often eaten. Flower products are important during dry season when fruits are scarce, with up to eleven pollen types eaten (e.g. Parkia ; Fabaceae ) and unidentified leaves. Breeding. Mating system of the Horsfield’s Short-nosed Fruit Bat is polygynous. Females are polyestrous, with postpartum estrus. Reproduction is largely aseasonal, with pregnant females found throughout the year. Litter size is one, with up to two litters per year. Activity patterns. Activity pattern is bimodal. Horsfield’s Short-nosed Fruit Bats are most active 2-4 hours after sunset and again c.3 hours before sunrise, with resting period around midnight. Movements, Home range and Social organization. Horsfield’s Short-nosed Fruit Bats mainly move between day roosts and feeding areas,traveling 400-1600 m. Mean home range is 8 ha for adult males and 6 ha for adult females. Home ranges widely overlap among males and females and with individuals of other species of Cynopterus (e.g. Lesser Short-nosed Fruit Bat, C. brachyotis ). Social organization is harem-based, with a single male associated with 2-5 reproductive females and their young. Harem group roosts in a tent built by the male or in unmodified roosts. In Peninsular Malaysia , banana ( Musa , Musaceae ) leaves are preferred roosts; ¢.50% of them is modified as tents by chewing mid-rib midway between base and tip. Other roosts include withering roots of epiphytic ferns ( Asplenium , Aspleniaceae ) and palm fronds ( Arenga , Cocos , Corypha , and Nypa ; Arecaceae ). Roosts are spatially clumped in forests. Males and females have low roost fidelity, and harem system is labile. Males change roosts every 1-7 days, and females change roosts every 3-14 days. Alternative roost sites include cave entrances and limestone solution cavities, where Horsfield’s Short-nosed Fruit Bats are more gregarious. Status and Conservation. Classified as Least Concern on The IUCN Red List. Horsfield’s Short-nosed Fruit Bat is abundant and uses modified forests as roosting and feeding habitat. Population trend is unknown. Bibliography. Andersen (1912b), Bates, Francis, Gumal & Bumrungsri (2008), Campbell (2008), Campbell & Kunz (2006), Campbell, Reid et al. (2006), Campbell, Schneider et al. (2006), Fletcher et al. (2012), Francis (1994), Funakoshi & Zubaid (1997), Hodgkison (2001), Jones, Bielby et al. (2009), Kingston et al. (2006), Kitchener, Gunnell & Maharadatunkamsi (1990), Tingga et al. (2012), Yong et al. (1973).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Pteropodidae	Cynopterus horsfieldii	Cynopterus		horsfieldii	Gray	1843	0	List Specimens Mamm. Coll. Brit. Mus.	p. 38	Horsfield&apos;s Fruit Bat	<b> harpax </b>Thomas and Wroughton, 1909; <b></b> lyoni K. Andersen, 1912; minor Lyon, 1908 [not Trouessart, 1878]; <b> persimilis K.</b> Andersen, 1912; <b> princeps </b> Miller, 1906.	Indonesia, Java.	Thailand, Cambodia, W Malaysia, Borneo, Java, Sumatra, Lesser Sunda Isls, and adjacent small islands.	Not listed.	Least Concern	Includes harpax ; see Hill (1961 a ). While this name is sometimes spelled horsefieldi , the correct original spelling is <horsfieldii following ICZN artcile 33.4.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Cynopterus horsfieldii	23	Horsfield's Short-nosed Fruit Bat	Codot Horsfield|Horsfield's Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	PTEROPODIDAE	CYNOPTERINAE	CYNOPTERINI	Cynopterus	NA	horsfieldii	J. E. Gray	1843	0						Java, Indonesia.			horsfieldii J. E. Gray, 1843|princeps G. S. Miller, 1906|harpax O. Thomas & Wroughton, 1909|minor (Lyon, 1908) [preoccupied by minor Trouessart, 1878]|lyoni K. Andersen, 1912|persimilis K. Andersen, 1912	NA	NA	Thailand|Malaysia|Singapore|Indonesia|Brunei	Asia	Indomalaya	LC	0	0	0	Cynopterus_horsfieldii	0	sciname match	Cynopterus_horsfieldii	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	6104	Cynopterus horsfieldii	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PTEROPODIDAE	Cynopterus	horsfieldii	Gray, 1843		20000000	Cynopterus horsfieldii	Least Concern		2019	2019-07-06 00:00:00 UTC	3.1	English	Listed as Least Concern as it is a common and widespread species, it is tolerant of habitat modification, and because it is unlikely to be declining at nearly the rate required to qualify for listing in a more threatened category.	It is found in many habitats in Indonesia but only pristine forests in Thailand. In Malaysia it has been found in agricultural areas, suburban parks, fruit orchards and secondary forest as well as good forest (Campbell and Kunst 2006). They have a harem social structure, and are highly gregarious when roosting in caves, but are found in smaller groups when roosting in forests. It makes tent roosts.	There are no major threats to this species throughout its range. Deforestation is a threat in Thailand, but elsewhere this species adapts to disturbed habitats.	This species is common in Indonesia (I. Maryanto pers. comm.), but locally rare in northern and western Thailand (S. Bumrungsri pers. comm.).	Unknown	This species is found in Thailand, Cambodia, Malaysia, Borneo, Java, Sumatra, Lesser Sunda Islands, and adjacent small islands in Indonesia. It has also been recorded from Bali, Lombok, Simelue and Nias (I. Maryanto pers. comm.).		Terrestrial	Isolated populations from Thailand need to be examined taxonomically (S. Bumrungsri pers. comm.). It is found in a number of protected areas throughout its range.	Australasian|Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Pteropodidae	Cynopterus		horsfieldii	Gray	1843	0	List Specimens Mamm. Coll. Brit. Mus.	p. 38	Horsfield&apos;s Fruit Bat	<b> harpax </b>Thomas and Wroughton, 1909; <b></b> lyoni K. Andersen, 1912; minor Lyon, 1908 [not Trouessart, 1878]; <b> persimilis K.</b> Andersen, 1912; <b> princeps </b> Miller, 1906.	Indonesia, Java.	Thailand, Cambodia, W Malaysia, Borneo, Java, Sumatra, Lesser Sunda Isls, and adjacent small islands.	Not listed.	Least Concern	Includes harpax ; see Hill (1961 a ). While this name is sometimes spelled horsefieldi , the correct original spelling is <horsfieldii following ICZN artcile 33.4.	Cynopterus horsfieldii	1004375	23	Horsfield's Short-nosed Fruit Bat	Codot Horsfield|Horsfield's Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	PTEROPODOIDEA	Pteropodidae	CYNOPTERINAE	CYNOPTERINI	Cynopterus	NA	horsfieldii	J. E. Gray	1843	0						Java, Indonesia.			horsfieldii J. E. Gray, 1843|princeps G. S. Miller, 1906|harpax O. Thomas & Wroughton, 1909|minor (Lyon, 1908) [preoccupied by minor Trouessart, 1878]|lyoni K. Andersen, 1912|persimilis K. Andersen, 1912	NA	NA				Thailand|Malaysia|Singapore|Indonesia|Brunei	Asia	Indomalaya	LC	0	0	0	Cynopterus_horsfieldii	0	sciname match	Cynopterus_horsfieldii	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Cynopterus_horsfieldii	1004375	23	Horsfield's Short-nosed Fruit Bat	Codot Horsfield|Horsfield's Fruit Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Pteropodoidea	Pteropodidae	Cynopterinae	Cynopterini	Cynopterus	NA	horsfieldii	J. E. Gray	0	Cynopterus Horsfieldii	Gray, J.E. 1843-05-13. List of the Specimens of Mammalia in the Collection of the British Museum. British Museum, London, 216 pp.	https://www.biodiversitylibrary.org/page/53729633	BMNH:Mamm:40a, BMNH:Mamm:40d	syntypes	https://data.nhm.ac.uk/object/308ba208-1f88-4a78-84ec-bc755a7c8e6c | https://data.nhm.ac.uk/object/af290465-d8a1-4ad8-83dc-616ee757b2e9 | https://data.nhm.ac.uk/object/e896ede7-5ab5-4887-8e87-fb8f693c194b	Java, Indonesia.			NA	NA				Thailand|Malaysia|Singapore|Indonesia|Brunei	Asia	Indomalaya	LC	0	0	0	Cynopterus_horsfieldii	0	sciname match	Cynopterus_horsfieldii	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Pteropodidae	Cynopterus		horsfieldii	Gray	1843	0	List Specimens Mamm. Coll. Brit. Mus.	p. 38	Horsfield's Short-nosed Fruit Bat	Yes.	Indonesia, Java	Thailand, Cambodia, W Malaysia, Borneo, Java, Sumatra, Lesser Sunda Isls, and adjacent small islands	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/6104/22113239/' target='_blank'>Least Concern</a>	Includes harpax; see Hill (1961). See also Campbell and Kunz (2006). While this name is sometimes spelled horsefieldi, the correct original spelling is horsfieldii following ICZN article 33.4.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Cynopterus horsfieldii; Cynopterus horsfieldii; Cynopterus horsfieldii; Cynopterus horsfieldii; Cynopterus horsfieldii; Cynopterus horsfieldii; horsfieldii; harpax; persimilis; princeps; harpax - lyoni; minor; horsfieldii; harpax; persimilis; princeps; harpax; lyoni; persimilis; princeps; lyoni - minor; horsfieldii; princeps; harpax; minor; lyoni; persimilis; Cynoptére de Horsfield; Horsfield-Kurznasenflughund; Cynéptero de Horsfield; Codot Horsfield; Horsfield's Fruit Bat; Horsfield's Short-nosed Fruit Bat; Codot Horsfield; Horsfield's Fruit Bat; Horsfield’s Fruit Bat; Horsfield&apos;s Fruit Bat; C. horsfieldii